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Ambedus
''Ambedus'' is an extinct genus of possible diadectid reptiliomorph. Fossils have been found from the Early Permian Dunkard Group of Monroe County, Ohio, Monroe County, Ohio. The type species ''A. pusillus'' was named in 2004. The genus name comes from the Latin word ''ambedo'' meaning "to nibble", in reference to its herbivorous diet. The specific name ''pusillus'' means "tiny" in Latin. Description ''Ambedus'' is a small tetrapod, possible Diadectid, known only from maxillae and dentary bones. It is usually considered the most primitive diadectid because unlike other forms it had a small, shallow lower jaw and many simple, conical teeth. It was also smaller than the later, rather bulky diadectids. Later diadectids have deep jaws with few teeth and forward-projecting incisor, incisiforms at the tips of the jaws for consuming plant material, and a corresponding greater girth. David Berman's 2013 paper argued against the inclusion of Ambedus in Diadectidae, for several reasons. He ...
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Diadectids
Diadectidae is an extinct family (biology), family of early tetrapods that lived in what is now North America and Europe during the Late Carboniferous and Early Permian in Asia during the Late Permian. They were the first herbivore, herbivorous tetrapods, and also the first fully terrestrial animals to attain large sizes. Footprints indicate that diadectids walked with an erect posture. They were the first to exploit plant material in terrestrial food chains, making their appearance an important stage in both vertebrate evolution and the development of terrestrial ecosystems. The best known and largest representative of the family is ''Diadectes'', a heavily built animal that attained a maximum length of several metres. Several other genera and various fragmentary fossil remains are also known. Although well known genera like ''Diadectes'' first appear in the Late Pennsylvanian, fragmentary remains of possible diadectids are known from much earlier deposits, including a piece of l ...
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Diadectid
Diadectidae is an extinct family of early tetrapods that lived in what is now North America and Europe during the Late Carboniferous and Early Permian in Asia during the Late Permian. They were the first herbivorous tetrapods, and also the first fully terrestrial animals to attain large sizes. Footprints indicate that diadectids walked with an erect posture. They were the first to exploit plant material in terrestrial food chains, making their appearance an important stage in both vertebrate evolution and the development of terrestrial ecosystems. The best known and largest representative of the family is ''Diadectes'', a heavily built animal that attained a maximum length of several metres. Several other genera and various fragmentary fossil remains are also known. Although well known genera like ''Diadectes'' first appear in the Late Pennsylvanian, fragmentary remains of possible diadectids are known from much earlier deposits, including a piece of lower jaw found in Missis ...
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Early Permian
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Lungfish
Lungfish are freshwater vertebrates belonging to the order Dipnoi. Lungfish are best known for retaining ancestral characteristics within the Osteichthyes, including the ability to breathe air, and ancestral structures within Sarcopterygii, including the presence of lobed fins with a well-developed internal skeleton. Lungfish represent the closest living relatives of the tetrapods. Today there are only six known species of lungfish, living in Africa, South America, and Australia. The fossil record shows that lungfish were abundant since the Triassic. While vicariance would suggest this represents an ancient distribution limited to the Mesozoic supercontinent Gondwana, the fossil record suggests advanced lungfish had a widespread freshwater distribution and the current distribution of modern lungfish species reflects extinction of many lineages subsequent to the breakup of Pangaea, Gondwana and Laurasia. Lungfish have historically been referred to as salamanderfish, but this t ...
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Paleontology In Ohio
Paleontology in Ohio refers to paleontological research occurring within or conducted by people from the U.S. state of Ohio. Ohio is well known for having a great quantity and diversity of fossils preserved in its Rock (geology), rocks. The state's fossil record begins early in the Paleozoic era, during the Cambrian period. Ohio was generally covered by seawater from that time on through the rest of the early Paleozoic. Local invertebrates included brachiopods, cephalopods, coral, graptolites, and trilobites. Vertebrates included bony fishes and sharks. The first land plants in the state grew during the Devonian. During the Carboniferous, Ohio became a more Ecoregion#Terrestrial, terrestrial environment with an increased diversity of plants that formed expansive swampy River delta, deltas. Amphibians and reptiles began to inhabit the state at this time, and remained present into the ensuing Permian. A gap in the local rock record spans from this point until the start of the Pleistoce ...
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Permian Ohio
The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleozoic Era; the following Triassic Period belongs to the Mesozoic Era. The concept of the Permian was introduced in 1841 by geologist Sir Roderick Murchison, who named it after the region of Perm in Russia. The Permian witnessed the diversification of the two groups of amniotes, the synapsids and the sauropsids (reptiles). The world at the time was dominated by the supercontinent Pangaea, which had formed due to the collision of Euramerica and Gondwana during the Carboniferous. Pangaea was surrounded by the superocean Panthalassa. The Carboniferous rainforest collapse left behind vast regions of desert within the continental interior. Amniotes, which could better cope with these drier conditions, rose to dominance in place of their amphibian ...
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Cisuralian Tetrapods Of North America
The Cisuralian is the first series/epoch of the Permian. The Cisuralian was preceded by the Pennsylvanian and followed by the Guadalupian. The Cisuralian Epoch is named after the western slopes of the Ural Mountains in Russia and Kazakhstan and dates between 298.9 ± 0.15 – 272.3 ± 0.5 Mya. The Cisuralian is often synonymous with the informal terms early Permian or lower Permian. It corresponds approximately with the Wolfcampian in southwestern North America. The series saw the appearance of beetles and flies and was a relatively stable warming period of about 21 million years. Name and background The Cisuralian is the first series or epoch of the Permian. The Cisuralian was preceded by the last Pennsylvanian epoch (Gzhelian) and is followed by the Permian Guadalupian Epoch. The name "Cisuralian" was proposed in 1982, and approved by the International Subcommission on Permian Stratigraphy in 1996. The Cisuralian Epoch is named after the western slopes of the Ural Mountain ...
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Kungurian Genera
In the geologic timescale, the Kungurian is an age (geology), age or stage (stratigraphy), stage of the Permian. It is the latest or upper of four subdivisions of the Cisuralian Epoch (geology), Epoch or series (stratigraphy), Series. The Kungurian lasted between and megaannum, million years ago (Ma). It was preceded by the Artinskian and followed by the Roadian. It corresponds roughly to the Leonardian Stage, covering the span from 280 to 270.6 ± 0.7 Ma in the North American system. Stratigraphy The Kungurian is named after the Russian city of Kungur in Perm Krai. The stage was introduced into scientific literature by Russian geologist Alexandr Antonovich Stukenberg (Alexander Stuckenberg) in 1890.; 2002: ''Progress report on the base of the Artinskian and base of the Kungurian by the Cisuralian Working Group'', Permophiles 41: pp 13–16. The base of the Kungurian Stage is defined as the place in the stratigraphic record where fossils of conodont species ''Neostrept ...
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Artinskian Genera
In the geologic timescale, the Artinskian is an age (geology), age or stage (stratigraphy), stage of the Permian. It is a subdivision of the Cisuralian Epoch (geology), Epoch or series (stratigraphy), Series. The Artinskian likely lasted between and megaannum, million years ago (Ma) according to the most recent revision of the International Commission on Stratigraphy (ICS) in 2022. It was preceded by the Sakmarian and followed by the Kungurian. Stratigraphy The Artinskian is named after the small Russian city of Arti, Russia, Arti (formerly ''Artinsk''), situated in the southern Ural mountains, about 200 km southwest of Yekaterinburg. The stage was introduced into scientific literature by Alexander Karpinsky in 1874. Base of the Artinskian The base of the Artinskian Stage is defined as the first appearance datum (FAD) of the conodont species ''Sweetognathus whitei'' and ''Mesogondolella bisselli''. In order to constrain this age, the ICS subcommission on Permian stra ...
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Ctenospondylus
''Ctenospondylus'' ("comb vertebra") is an extinct genus of sphenacodontid synapsid Species were about three meters (ten feet) long. The genus is known only from the 'Seymouran' Land Vertebrate Faunachron, which is equivalent to the upper part of the Artinskian stage and the lowermost Kungurian stage of the Early Permian. Its fossils were found in the U.S. states of Ohio and Texas. It was a carnivore and preyed upon animals close to its size. ''Ctenospondylus'' had a long tail, short back spines, and a very deep yet narrow skull with massive jaws that had sharp teeth. Because of its large size, it was probably the apex predator in its environment, and might have competed with other predators like ''Dimetrodon'' for food. A sphenacodontid, it was a close relative of ''Dimetrodon''. In 1936, Alfred Sherwood Romer named the type species ''Ctenospondylus casei''. The generic name, from Greek κτείς, ''kteis'', "comb", and σφόνδυλος, ''sphondylos'', "vertebra", refe ...
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Synapsid
Synapsids + (, 'arch') > () "having a fused arch"; synonymous with ''theropsids'' (Greek, "beast-face") are one of the two major groups of animals that evolved from basal amniotes, the other being the sauropsids, the group that includes reptiles and birds. The group includes mammals and every animal more closely related to mammals than to sauropsids. Unlike other amniotes, synapsids have a single temporal fenestra, an opening low in the skull roof behind each eye orbit, leaving a bony arch beneath each; this accounts for their name. The distinctive temporal fenestra developed about 318 million years ago during the Late Carboniferous period, when synapsids and sauropsids diverged, but was subsequently merged with the orbit in early mammals. Traditionally, non-mammalian synapsids were believed to have evolved from reptiles, and therefore described as mammal-like reptiles in classical systematics, and primitive synapsids were also referred to as pelycosaurs, or pelycosaur-gr ...
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Embolomeri
Embolomeri is an order of tetrapods or stem-tetrapods, possibly members of Reptiliomorpha. Embolomeres first evolved in the Early Carboniferous ( Mississippian) Period and were the largest and most successful predatory tetrapods of the Late Carboniferous ( Pennsylvanian) Period. They were specialized semiaquatic predators with long bodies for eel-like undulatory swimming. Embolomeres are characterized by their vertebral centra, which are formed by two cylindrical segments, the pleurocentrum at the rear and intercentrum at the front. These segments are equal in size. Most other tetrapods have pleurocentra and intercentra which are drastically different in size and shape. Embolomeres were among the earliest large carnivorous tetrapods, with members such as the crocodilian-like ''Proterogyrinus'' appearing in the Visean stage of the Carboniferous. They declined in diversity during the Permian period, though at least one representative ('' Archeria)'' was common in the Early Permian ...
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