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Alouatta Coibensis Coibensis
The Coiba Island howler (''Alouatta coibensis'') is a type of howler monkey, a type of New World monkey, endemic to Panama. Although the Coiba Island howler has been recognized as a separate species by a number of authorities since a 1987 study of its fingerprints, mitochondrial DNA testing found it does not differ from mantled howler populations in any significant way. A reason given for treating it as a separate species is that the dermal ridges of its hands and feet differ from those of the mantled howler. ''A. c. coibensis'' is smaller than other Central American howler monkeys and has duller pelage than the Azuero howler, ''Alouatta coibensis trabeata''. Subspecies Two subspecies of this howler have been recognized by those who consider it a separate species: *''Alouatta coibensis coibensis'' Thomas, 1902, found on Coiba Island and Jicaron, off the Pacific coast of Panama *Azuero howler, ''A. c. trabeata'' Lawrence, 1933, endemic to the Azuero Peninsula Azuero Peninsul ...
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Oldfield Thomas
Michael Rogers Oldfield Thomas (21 February 1858 – 16 June 1929) was a British zoologist. Career Thomas worked at the Natural History Museum on mammals, describing about 2,000 new species and subspecies for the first time. He was appointed to the museum secretary's office in 1876, transferring to the zoological department in 1878. In 1891, Thomas married Mary Kane, daughter of Sir Andrew Clark, heiress to a small fortune, which gave him the finances to hire mammal collectors and present their specimens to the museum. He also did field work himself in Western Europe and South America. His wife shared his interest in natural history, and accompanied him on collecting trips. In 1896, when William Henry Flower took control of the department, he hired Richard Lydekker to rearrange the exhibitions, allowing Thomas to concentrate on these new specimens. Thomas viewed his taxonomy efforts from the scope of British imperialism. "You and I in our scientific lives have seen th ...
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Mammals Described In 1902
Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or hair, and three middle ear bones. These characteristics distinguish them from reptiles (including birds) from which they diverged in the Carboniferous, over 300 million years ago. Around 6,400 extant species of mammals have been described divided into 29 orders. The largest orders, in terms of number of species, are the rodents, bats, and Eulipotyphla (hedgehogs, moles, shrews, and others). The next three are the Primates (including humans, apes, monkeys, and others), the Artiodactyla (cetaceans and even-toed ungulates), and the Carnivora (cats, dogs, seals, and others). In terms of cladistics, which reflects evolutionary history, mammals are the only living members of the Synapsida (synapsids); this clade, together with Saurops ...
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Primates Of Central America
Primates are a diverse order of mammals. They are divided into the strepsirrhines, which include the lemurs, galagos, and lorisids, and the haplorhines, which include the tarsiers and the simians (monkeys and apes, the latter including humans). Primates arose 85–55 million years ago first from small terrestrial mammals, which adapted to living in the trees of tropical forests: many primate characteristics represent adaptations to life in this challenging environment, including large brains, visual acuity, color vision, a shoulder girdle allowing a large degree of movement in the shoulder joint, and dextrous hands. Primates range in size from Madame Berthe's mouse lemur, which weighs , to the eastern gorilla, weighing over . There are 376–524 species of living primates, depending on which classification is used. New primate species continue to be discovered: over 25 species were described in the 2000s, 36 in the 2010s, and three in the 2020s. Primates have larg ...
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Howler Monkeys
Howler monkeys (genus ''Alouatta'', monotypic in subfamily Alouattinae) are the most widespread primate genus in the Neotropics and are among the largest of the platyrrhines along with the muriquis (''Brachyteles''), the spider monkeys (''Ateles'') and woolly monkeys (''Lagotrix''). These monkeys are native to South and Central American forests. They are famous for their loud howls, which can travel more than a mile through dense rain forest. Fifteen species are recognized. Previously classified in the family Cebidae, they are now placed in the family Atelidae. They are primarily folivores but also significant frugivores, acting as seed dispersal agents through their digestive system and their locomotion. Threats include human predation, habitat destruction, and capture for pets or zoo animals. Classification * ''A. palliata'' group ** Coiba Island howler, ''Alouatta coibensis'' *** ''Alouatta coibensis coibensis'' *** Azuero howler, ''Alouatta coibensis trabeata'' ** Ma ...
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Azuero Peninsula
Azuero Peninsula ( es, Península de Azuero) is a large peninsula in southern Panama. It is surrounded by the Pacific Ocean in the south; the Pacific and Gulf of Montijo to the west, and by the Gulf of Panama in the east. The peninsula is effectively divided into two regions; the Western Azuero and the Eastern Azuero, as no serviceable roads join the two peninsula regions past the Pan-American Highway. The Eastern Azuero Peninsula is known for baseball and is also a center of activity during the annual '' carnaval'' ( carnival), with Las Tablas being the hub. Pedasi is a small fishing town with sport fishing. The Western Azuero Peninsula is known for its cattle ranching, farming, fishing, sunsets and beaches. Tourism has begun to increase in the area both for the aforementioned sport fishing, surfing and for the local charm of cities like Chitré, Las Tablas and Pedasi. Due to a rise in tourism, real estate development has begun. The area enjoys some of the best weather in Pa ...
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Endemism
Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsewhere. For example, the Cape sugarbird is found exclusively in southwestern South Africa and is therefore said to be ''endemic'' to that particular part of the world. An endemic species can be also be referred to as an ''endemism'' or in scientific literature as an ''endemite''. For example '' Cytisus aeolicus'' is an endemite of the Italian flora. '' Adzharia renschi'' was once believed to be an endemite of the Caucasus, but it was later discovered to be a non-indigenous species from South America belonging to a different genus. The extreme opposite of an endemic species is one with a cosmopolitan distribution, having a global or widespread range. A rare alternative term for a species that is endemic is "precinctive", which applies t ...
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Pacific
The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the continents of Asia and Oceania in the west and the Americas in the east. At in area (as defined with a southern Antarctic border), this largest division of the World Ocean—and, in turn, the hydrosphere—covers about 46% of Earth's water surface and about 32% of its total surface area, larger than Earth's entire land area combined .Pacific Ocean
. '' Britannica Concise.'' 2008: Encyclopædia Britannica, Inc.
The centers of both the



Coiba Island
Coiba is the largest island in Central America, with an area of , off the Pacific coast of the Panamanian province of Chiriquí. It is part of the Tolé District of that province. History Coiba separated from continental Panama about 12,000 to 18,000 years ago when sea levels rose. Plants and animals on the new island became isolated from mainland populations and over the millennia most animals have diverged in appearance and behaviour from their mainland counterparts. The island is home to many endemic subspecies, including the Coiba Island howler monkey, the Coiba agouti and the Coiba spinetail. Coiba was home to the Coiba Cacique Indians until about 1560, when they were conquered by the Spanish and forced into slavery. In 1919 a penal colony was built on the island and during the years that Panama was under the dictatorships of Omar Torrijos and Manuel Noriega, the prison on Coiba was a feared place with a reputation for brutal conditions, extreme tortures, executions ...
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Howler Monkey
Howler monkeys (genus ''Alouatta'', monotypic in subfamily Alouattinae) are the most widespread primate genus in the Neotropics and are among the largest of the platyrrhines along with the muriquis (''Brachyteles''), the spider monkeys (''Ateles'') and woolly monkeys (''Lagotrix''). These monkeys are native to South and Central American forests. They are famous for their loud howls, which can travel more than a mile through dense rain forest. Fifteen species are recognized. Previously classified in the family Cebidae, they are now placed in the family Atelidae. They are primarily folivores but also significant frugivores, acting as seed dispersal agents through their digestive system and their locomotion. Threats include human predation, habitat destruction, and capture for pets or zoo animals. Classification * ''A. palliata'' group ** Coiba Island howler, ''Alouatta coibensis'' *** ''Alouatta coibensis coibensis'' *** Azuero howler, ''Alouatta coibensis trabeata'' ** Mant ...
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Azuero Howler
The Azuero howler (''Alouatta coibensis trabeata'') a type of monkey that is a subspecies of the Coiba Island howler ''A. coibensis''. This subspecies is endemic to the Azuero Peninsula in Panama. The Azuero howler is distinguished primarily by its golden flanks and loins, and browner appearance on the rest of its body. Although generally considered a subspecies of ''A. coibensis'', there is some debate within the scientific community as to whether ''A. coibensis'' itself is a valid species. If not, ''A. c. trabeata'' would be considered subspecies of the mantled howler The mantled howler (''Alouatta palliata'') is a species of howler monkey, a type of New World monkey, from Central and South America. It is one of the monkey species most often seen and heard in the wild in Central America. It takes its "mantle ..., ''A. palliata''. In that case its trinomial name would be ''A. p. trabeata''. References Primates of Central America Howler monkeys Mammals described in ...
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Mantled Howler
The mantled howler (''Alouatta palliata'') is a species of howler monkey, a type of New World monkey, from Central and South America. It is one of the monkey species most often seen and heard in the wild in Central America. It takes its "mantled" name from the long guard hairs on its sides. The mantled howler is one of the largest Central American monkeys, and males can weigh up to . It is the only Central American monkey that eats large quantities of leaves; it has several adaptations to this folivorous diet. Since leaves are difficult to digest and provide less energy than most foods, the mantled howler spends the majority of each day resting and sleeping. The male mantled howler has an enlarged hyoid bone, a hollow bone near the vocal cords, which amplifies the calls made by the male, and is the reason for the name "howler". Howling allows the monkeys to locate each other without expending energy on moving or risking physical confrontation. The mantled howler lives in gro ...
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