Arachnopulmonata
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Arachnopulmonata
Arachnids are arthropods in the class Arachnida () of the subphylum Chelicerata. Arachnida includes, among others, spiders, scorpions, ticks, mites, pseudoscorpions, harvestmen, camel spiders, whip spiders and vinegaroons. Adult arachnids have eight legs attached to the cephalothorax. In some species the frontmost pair of legs has converted to a sensory function, while in others, different appendages can grow large enough to take on the appearance of extra pairs of legs. Almost all extant arachnids are terrestrial, living mainly on land. However, some inhabit freshwater environments and, with the exception of the pelagic zone, marine environments as well. They comprise over 110,000 named species, of which 51,000 are species of spiders. The term is derived from the Greek word (''aráchnē'', 'spider'), from the myth of the hubristic human weaver Arachne, who was turned into a spider. Morphology Almost all adult arachnids have eight legs, unlike adult insects which all ...
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Scorpion
Scorpions are predatory arachnids of the Order (biology), order Scorpiones. They have eight legs and are easily recognized by a pair of Chela (organ), grasping pincers and a narrow, segmented tail, often carried in a characteristic forward curve over the back and always ending with a stinger. The evolutionary history of scorpions goes back Silurian, 435 million years. They mainly live in deserts but have adapted to a wide range of environmental conditions, and can be found on all continents except Antarctica. There are over 2,500 described species, with 22 extant (living) families recognized to date. Their Taxonomy (biology), taxonomy is being revised to account for 21st-century genomic studies. Scorpions primarily prey on insects and other invertebrates, but some species hunt vertebrates. They use their pincers to restrain and kill prey, or to prevent their own predation. The Scorpion sting, venomous sting is used for offense and defense. During courtship, the male and female ...
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Opiliones
The Opiliones (formerly Phalangida) are an Order (biology), order of arachnids, Common name, colloquially known as harvestmen, harvesters, harvest spiders, or daddy longlegs (see below). , over 6,650 species of harvestmen have been discovered worldwide, although the total number of extant taxon, extant species may exceed 10,000. The order Opiliones includes five suborders: Cyphophthalmi, Eupnoi, Dyspnoi, Laniatores, and Tetrophthalmi, which were named in 2014. Representatives of each extant suborder can be found on all continents except Antarctica. Well-preserved fossils have been found in the 400-million-year-old Rhynie cherts of Scotland, and 305-million-year-old rocks in France. These fossils look surprisingly modern, indicating that their basic body shape developed very early on, and, at least in some taxa, has changed little since that time. Their Phylogenetics, phylogenetic position within the Arachnida is disputed; their closest relatives may be camel spiders (Solifugae ...
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Acari
Mites are small arachnids (eight-legged arthropods) of two large orders, the Acariformes and the Parasitiformes, which were historically grouped together in the subclass Acari. However, most recent genetic analyses do not recover the two as each other's closest relative within Arachnida, rendering the group invalid as a clade. Most mites are tiny, less than in length, and have a simple, unsegmented body plan. The small size of most species makes them easily overlooked; some species live in water, many live in soil as decomposers, others live on plants, sometimes creating galls, while others are predators or parasites. This last type includes the commercially destructive '' Varroa'' parasite of honey bees, as well as scabies mites of humans. Most species are harmless to humans, but a few are associated with allergies or may transmit diseases. The scientific discipline devoted to the study of mites is called acarology. Evolution and taxonomy Mites are not a defined taxon, b ...
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Trigonotarbida
The order Trigonotarbida is a group of extinct arachnids whose fossil record extends from the late Silurian to the early Permian ( Pridoli to Sakmarian).Dunlop, J. A., Penney, D. & Jekel, D. 2020A summary list of fossil spiders and their relatives In World Spider Catalog. Natural History Museum Bern, online at http://wsc.nmbe.ch , version 20.5 These animals are known from several localities in Europe and North America, as well as a single record from Argentina. Trigonotarbids can be envisaged as spider-like arachnids, but without silk-producing spinnerets. They ranged in size from a few millimetres to a few centimetres in body length and had segmented abdomens (opisthosoma), with the dorsal exoskeleton ( tergites) across the backs of the animals' abdomens, which were characteristically divided into three or five separate plates. Probably living as predators on other arthropods, some later trigonotarbid species were quite heavily armoured and protected themselves with spines and tub ...
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Tetrapulmonata
Tetrapulmonata is a Taxonomic rank, non-ranked Order (biology), supra-ordinal clade of arachnids. It is composed of the Extant taxon, extant orders Uropygi (whip scorpions), Schizomida (short-tailed whip scorpions), Amblypygi (tail-less whip scorpions) and Spider, Araneae (spiders). It is the only supra-ordinal group of arachnids that is Resampling (statistics), strongly supported in molecular phylogenetic Research, studies. Two extinct orders are also placed in this clade, Haptopoda and Uraraneida. In 2016, a newly described fossil arachnid, ''Idmonarachne'', was also included in the Tetrapulmonata; it has not been assigned to an order. Etymology It receives its International Code of Zoological Nomenclature, name from the presence of paired book lungs occupying the second and third opisthosomal segments, although the posterior pair is absent in Schizomida and most Araneomorphae, araneomorph spiders. Previous synonym (taxonomy), synonyms of this lineage are rejected; "Caulogastra ...
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Schizotarsata
Tetrapulmonata is a non-ranked supra-ordinal clade of arachnids. It is composed of the extant orders Uropygi (whip scorpions), Schizomida (short-tailed whip scorpions), Amblypygi (tail-less whip scorpions) and Araneae (spiders). It is the only supra-ordinal group of arachnids that is strongly supported in molecular phylogenetic studies. Two extinct orders are also placed in this clade, Haptopoda and Uraraneida. In 2016, a newly described fossil arachnid, ''Idmonarachne'', was also included in the Tetrapulmonata; it has not been assigned to an order. Etymology It receives its name from the presence of paired book lungs occupying the second and third opisthosomal segments, although the posterior pair is absent in Schizomida and most araneomorph spiders. Previous synonyms of this lineage are rejected; "Caulogastra Pocock, 1893" refers to pedicel, which is symplesiomorphic for the lineage and convergent with Solifugae, and "Arachnidea Van der Hammen, 1977" is easily confused ...
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Haptopoda
''Plesiosiro'' is an extinct arachnid genus known exclusively from nine specimens from the Upper Carboniferous of Coseley, Staffordshire, United Kingdom. The genus is monotypic, represented only by the species ''Plesiosiro madeleyi'' described by Reginald Innes Pocock in his important 1911 monograph on British Carboniferous arachnids. It is the only known member of the order Haptopoda. The original fossils have been redescribed in detail by Alexander Petrunkevitch in 1949 and Dunlop in 1999. A supposed example from the Coal Measures of Lancashire is a misidentification. Relationships ''Plesiosiro'' means "close to '' Siro''", which is a genus of cyphophthalmid (Cyphophthalmi); the most primitive group of the living harvestmen (Opiliones). These harvestmen do, in some ways, resemble the reconstructed body plan of the haptopods. Revisions have confirmed that Haptopoda should be treated as a separate and independent order. A 2007 study tentatively recognised a group named Schizot ...
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Serikodiastida
Tetrapulmonata is a non-ranked supra-ordinal clade of arachnids. It is composed of the extant orders Uropygi (whip scorpions), Schizomida (short-tailed whip scorpions), Amblypygi (tail-less whip scorpions) and Araneae (spiders). It is the only supra-ordinal group of arachnids that is strongly supported in molecular phylogenetic studies. Two extinct orders are also placed in this clade, Haptopoda and Uraraneida. In 2016, a newly described fossil arachnid, ''Idmonarachne'', was also included in the Tetrapulmonata; it has not been assigned to an order. Etymology It receives its name from the presence of paired book lungs occupying the second and third opisthosomal segments, although the posterior pair is absent in Schizomida and most araneomorph spiders. Previous synonyms of this lineage are rejected; "Caulogastra Pocock, 1893" refers to pedicel, which is symplesiomorphic for the lineage and convergent with Solifugae, and "Arachnidea Van der Hammen, 1977" is easily confused ...
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