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Aculeata
Aculeata is an infraorder of Hymenoptera containing ants, bees, and stinging wasps. The name is a reference to the defining feature of the group, which is the modification of the ovipositor into a stinger. However, many members of the group cannot sting, either retaining the ovipositor, or having lost it altogether. A large part of the clade is parasitic. This group includes all of the eusocial Hymenopterans. The oldest aculeates are known from the Late Jurassic Karabastau Formation of Kazakhstan, represented by the family Bethylonymidae, which may be para- or polyphyletic. Classification The use of the name Aculeata has a long history at the rank of infraorder or division. The Aculeata are a monophyletic, or good natural group, containing all the descendants of a single common ancestor. The Aculeata are therefore maintained as a taxon, either at infraorder or division rank or as an unranked clade In biology, a clade (), also known as a Monophyly, monophyletic group ...
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Apocrita
Apocrita is a suborder of insects in the order Hymenoptera. It includes wasps, bees, and ants, and consists of many families. It contains the most advanced hymenopterans and is distinguished from Symphyta by the narrow "waist" ( petiole) formed between the first two segments of the actual abdomen; the first abdominal segment is fused to the thorax, and is called the propodeum. Therefore, it is general practice, when discussing the body of an apocritan in a technical sense, to refer to the mesosoma and metasoma (or gaster) rather than the "thorax" and "abdomen", respectively. The evolution of a constricted waist was an important adaption for the parasitoid lifestyle of the ancestral apocritan, allowing more maneuverability of the female's ovipositor. The ovipositor either extends freely or is retracted, and may be developed into a stinger for both defense and paralyzing prey. Larvae are legless and blind, and either feed inside a host (plant or animal) or in a nest cell prov ...
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Parasitic Wasp
Parasitoid wasps are a large group of hymenopteran superfamilies, with all but the wood wasps ( Orussoidea) being in the wasp-waisted Apocrita. As parasitoids, they lay their eggs on or in the bodies of other arthropods, sooner or later causing the death of these host (biology), hosts. Different species specialise in hosts from different insect orders, most often Lepidoptera, though some select Coleoptera, beetles, Diptera, flies, or Hemiptera, bugs; the spider wasps (Pompilidae) exclusively attack spiders. Parasitoid wasp species differ in which host life-stage they attack: eggs, larvae, pupae, or adults. They mainly follow one of two major strategies within parasitism: either they are endoparasitic, developing inside the host, and koinobiont, allowing the host to continue to feed, develop, and moult; or they are ectoparasitic, developing outside the host, and idiobiont, paralysing the host immediately. Some endoparasitic wasps of the superfamily Ichneumonoidea have a mutuali ...
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Chrysidoidea
The superfamily Chrysidoidea is a very large cosmopolitan group, all of which are parasitoids or cleptoparasites of other insects. There are three large, common families (Bethylidae, Chrysididae, and Dryinidae) and four small, rare families (Embolemidae, Plumariidae, Sclerogibbidae, and Scolebythidae). Most species are small (7 mm or less), almost never exceeding 15 mm. This superfamily is traditionally considered to be the basal taxon within the Aculeata, and, as such, some species can sting, though the venom is harmless to humans. Members of the families Dryinidae and Embolemidae are the only parasitoids among the Hymenoptera to have a life cycle in which the wasp larva begins its life inside the body of the host, and then later forms a sac (called a ''thylacium'') that protrudes out of the host's abdomen. The closely related family Sclerogibbidae contains more traditional ectoparasitoids, attacking the nymph (biology), nymphs of webspinners. The extinct, monotypic ...
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Apoidea
The superfamily Apoidea is a major group (of over 30 000 species) within the Hymenoptera, which includes two traditionally recognized lineages, the "sphecoid" wasps, and the bees. Molecular phylogeny demonstrates that the bees arose from within the traditional " Crabronidae", so that grouping is paraphyletic, and this has led to a reclassification to produce monophyletic families.Manuela Sann, Oliver Niehuis, Ralph S. Peters, Christoph Mayer, Alexey Kozlov, Lars Podsiadlowski, Sarah Bank, Karen Meusemann, Bernhard Misof, Christoph Bleidorn and Michael Ohl (2018) Phylogenomic analysis of Apoidea sheds new light on the sister group of bees. ''BMC Evolutionary Biology'' 18:71. doi:10.1186/s12862-018-1155-8 Diagnostic features Apoid wasps and bees have several traits in common:O'Neill, K.M. (2008). Apoid Wasps (Hymenoptera: Apoidea: Spheciformes). In: Capinera, J.L. (eds) Encyclopedia of Entomology. Springer, Dordrecht. https://doi.org/10.1007/978-1-4020-6359-6_10300 * The ...
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Hymenoptera
Hymenoptera is a large order of insects, comprising the sawflies, wasps, bees, and ants. Over 150,000 living species of Hymenoptera have been described, in addition to over 2,000 extinct ones. Many of the species are parasitic. Females typically have a special ovipositor for inserting eggs into hosts or places that are otherwise inaccessible. This ovipositor is often modified into a stinger. The young develop through holometabolism (complete metamorphosis)—that is, they have a wormlike larval stage and an inactive pupal stage before they reach adulthood. Etymology The name Hymenoptera refers to the wings of the insects, but the original derivation is ambiguous. All references agree that the derivation involves the Ancient Greek πτερόν (''pteron'') for wing. The Ancient Greek ὑμήν (''hymen'') for membrane provides a plausible etymology for the term because species in this order have membranous wings. However, a key characteristic of this order is that the hi ...
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Paraphyly
Paraphyly is a taxonomic term describing a grouping that consists of the grouping's last common ancestor and some but not all of its descendant lineages. The grouping is said to be paraphyletic ''with respect to'' the excluded subgroups. In contrast, a monophyletic grouping (a clade) includes a common ancestor and ''all'' of its descendants. The terms are commonly used in phylogenetics (a subfield of biology) and in the tree model of historical linguistics. Paraphyletic groups are identified by a combination of synapomorphies and symplesiomorphies. If many subgroups are missing from the named group, it is said to be polyparaphyletic. The term received currency during the debates of the 1960s and 1970s accompanying the rise of cladistics, having been coined by zoologist Willi Hennig to apply to well-known taxa like Reptilia (reptiles), which is paraphyletic with respect to birds. Reptilia contains the last common ancestor of reptiles and all descendants of that ancest ...
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Tiphioidea
Tiphioidea is a suggested superfamily of stinging wasps in the order Hymenoptera. There are three families in Tiphioidea, Bradynobaenidae, Tiphiidae, and Sierolomorphidae. Recent research in molecular phylogenetics has resulted in the reorganization of the infraorder Aculeata, which now contains eight superfamilies: Apoidea, Chrysidoidea The superfamily Chrysidoidea is a very large cosmopolitan group, all of which are parasitoids or cleptoparasites of other insects. There are three large, common families (Bethylidae, Chrysididae, and Dryinidae) and four small, rare families (Em ..., Formicoidea, Pompiloidea, Scolioidea, Tiphioidea, Thynnoidea, and Vespoidea. References Further reading * * * External links * Hymenoptera Apocrita superfamilies {{Apocrita-stub ...
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Vespoidea
Vespoidea is a superfamily of wasps in the order Hymenoptera. Vespoidea includes wasps with a large variety of lifestyles including eusocial, social, and solitary habits, predators, scavengers, parasitoids, and some herbivores. Description Vespoid wasp females have antennae with 10 flagellomeres, while males have 11 flagellomeres. The edge of the pronotum reaches or passes the tegula. Many species display some level of sexual dimorphism. Most species have fully developed wings, but some have reduced or absent wings in one or both sexes. As in other Aculeata, only the females are ever capable of stinging. Phylogenetics and taxonomy Research based on four nuclear genes (elongation factor-1α F2 copy, long-wavelength rhodopsin, wingless and the D2–D3 regions of 28S ribosomal RNA—2700 bp in total) suggests the historical view of family relationships need to be changed, with Rhopalosomatidae as a sister group of the Vespidae and the clade Rhopalosomatidae + Vespidae a ...
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Hymenopteran
Hymenoptera is a large order of insects, comprising the sawflies, wasps, bees, and ants. Over 150,000 living species of Hymenoptera have been described, in addition to over 2,000 extinct ones. Many of the species are parasitic. Females typically have a special ovipositor for inserting eggs into hosts or places that are otherwise inaccessible. This ovipositor is often modified into a stinger. The young develop through holometabolism (complete metamorphosis)—that is, they have a wormlike larval stage and an inactive pupal stage before they reach adulthood. Etymology The name Hymenoptera refers to the wings of the insects, but the original derivation is ambiguous. All references agree that the derivation involves the Ancient Greek πτερόν (''pteron'') for wing. The Ancient Greek ὑμήν (''hymen'') for membrane provides a plausible etymology for the term because species in this order have membranous wings. However, a key characteristic of this order is that the h ...
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Ovipositor
The ovipositor is a tube-like organ used by some animals, especially insects, for the laying of eggs. In insects, an ovipositor consists of a maximum of three pairs of appendages. The details and morphology of the ovipositor vary, but typically its form is adapted to functions such as preparing a place for the egg, transmitting the egg, and then placing it properly. For most insects, the organ is used merely to attach the egg to some surface, but for many parasitic species (primarily in wasps and other Hymenoptera), it is a piercing organ as well. Some ovipositors only retract partly when not in use, and the basal part that sticks out is known as the scape, or more specifically oviscape, the word ''scape'' deriving from the Latin word , meaning "stalk" or "shaft". In insects Grasshoppers use their ovipositors to force a burrow into the earth to receive the eggs. Cicadas pierce the wood of twigs with their ovipositors to insert the eggs. Sawflies slit the tissues of ...
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Clade
In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach to taxonomy adopted by most biological fields. The common ancestor may be an individual, a population, or a species (extinct or Extant taxon, extant). Clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently. Clades are termed ''monophyletic'' (Greek: "one clan") groups. Over the last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming Taxon, taxa that are not clades; that is, taxa that are not Monophyly, monophyletic. Some of the relationships between organisms that the molecul ...
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Sawfly
Sawflies are wasp-like insects that are in the suborder Symphyta within the order Hymenoptera, alongside ants, bees, and wasps. The common name comes from the saw-like appearance of the ovipositor, which the females use to cut into the plants where they lay their eggs. The name is associated especially with the Tenthredinoidea, by far the largest superfamily in the suborder, with about 7,000 known species; in the entire suborder, there are 8,000 described species in more than 800 genera. Symphyta is paraphyletic, consisting of several basal groups within the order Hymenoptera, each one rooted inside the previous group, ending with the Apocrita which are not sawflies. The primary distinction between sawflies and the Apocrita – the ants, bees, and wasps – is that the adults lack a "wasp waist", and instead have a broad connection between the abdomen and the thorax. Some sawflies are Batesian mimics of wasps and bees, and the ovipositor can be mistaken for a stinger. S ...
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