δ13C
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δ13C
In geochemistry, paleoclimatology, and paleoceanography ''δ''13C (pronounced "delta thirteen c") is an isotopic signature, a measure of the ratio of the two stable isotopes of carbon— 13C and 12C—reported in parts per thousand (per mil, ‰). The measure is also widely used in archaeology for the reconstruction of past diets, particularly to see if marine foods or certain types of plants were consumed. The definition is, in per mille: :\delta \ce = \left( \frac - 1 \right) \times 1000 where the standard is an established reference material. ''δ''13C varies in time as a function of productivity, the signature of the inorganic source, organic carbon burial, and vegetation type. Biological processes preferentially take up the lower mass isotope through kinetic fractionation. However some abiotic processes do the same. For example, methane from hydrothermal vents can be depleted by up to 50‰. Reference standard The standard established for carbon-13 work was the ...
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Isotopic Signature
An isotopic signature (also isotopic fingerprint) is a ratio of non-radiogenic ' stable isotopes', stable radiogenic isotopes, or unstable radioactive isotopes of particular elements in an investigated material. The ratios of isotopes in a sample material are measured by isotope-ratio mass spectrometry against an isotopic reference material. This process is called isotope analysis. Stable isotopes The atomic mass of different isotopes affect their chemical kinetic behavior, leading to natural isotope separation processes. Carbon isotopes For example, different sources and sinks of methane have different affinity for the 12C and 13C isotopes, which allows distinguishing between different sources by the 13C/12C ratio in methane in the air. In geochemistry, paleoclimatology and paleoceanography this ratio is called δ13C. The ratio is calculated with respect to Pee Dee Belemnite (PDB) standard: :\delta \ce_\mathrm = \left(\frac - 1\right) \cdot 1000 ‰ Similarly, c ...
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Carbon-13
Carbon-13 (13C) is a natural, stable isotope of carbon with a nucleus containing six protons and seven neutrons. As one of the environmental isotopes, it makes up about 1.1% of all natural carbon on Earth. Detection by mass spectrometry A mass spectrum of an organic compound will usually contain a small peak of one mass unit greater than the apparent molecular ion peak (M) of the whole molecule. This is known as the M+1 peak and comes from the few molecules that contain a 13C atom in place of a 12C. A molecule containing one carbon atom will be expected to have an M+1 peak of approximately 1.1% of the size of the M peak, as 1.1% of the molecules will have a 13C rather than a 12C. Similarly, a molecule containing two carbon atoms will be expected to have an M+1 peak of approximately 2.2% of the size of the M peak, as there is double the previous likelihood that any molecule will contain a 13C atom. In the above, the mathematics and chemistry have been simplified, however it ...
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Paleocene–Eocene Thermal Maximum
The Paleocene–Eocene thermal maximum (PETM), alternatively ”Eocene thermal maximum 1 (ETM1)“ and formerly known as the "Initial Eocene" or “Late Paleocene thermal maximum", was a geologically brief time interval characterized by a global average temperature rise and massive input of carbon into the ocean and atmosphere. The event began, now formally codified, at the precise time boundary between the Paleocene and Eocene geological Epoch (geology), epochs. The exact age and duration of the PETM remain uncertain, but it occurred around 55.8 million years ago (Ma) and lasted about 200 thousand years (Ka). The PETM arguably represents our best past analogue for which to understand how global warming and the carbon cycle operate in a greenhouse world. The time interval is marked by a prominent negative excursion in carbon stable isotope () records from around the globe; more specifically, a large decrease in the 13C/Carbon-12, 12C ratio of marine and terrestrial carbon ...
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Methane Clathrate
Methane clathrate (CH4·5.75H2O) or (4CH4·23H2O), also called methane hydrate, hydromethane, methane ice, fire ice, natural gas hydrate, or gas hydrate, is a solid clathrate compound (more specifically, a clathrate hydrate) in which a large amount of methane is trapped within a crystal structure of water, forming a solid similar to ice. Originally thought to occur only in the outer regions of the Solar System, where temperatures are low and water ice is common, significant deposits of methane clathrate have been found under sediments on the ocean floors of the Earth (around 1100m below the sea level). Methane hydrate is formed when hydrogen-bonded water and methane gas come into contact at high pressures and low temperatures in oceans. Methane clathrates are common constituents of the shallow marine geosphere and they occur in deep sedimentary structures and form outcrops on the ocean floor. Methane hydrates are believed to form by the precipitation or crystallisation of metha ...
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Benthic Foraminifera
The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from the Ancient Greek word (), meaning "the depths". Organisms living in this zone are called benthos and include microorganisms (e.g., bacteria and fungi) as well as larger invertebrates, such as crustaceans and polychaetes. Organisms here, known as bottom dwellers, generally live in close relationship with the substrate and many are permanently attached to the bottom. The benthic boundary layer, which includes the bottom layer of water and the uppermost layer of sediment directly influenced by the overlying water, is an integral part of the benthic zone, as it greatly influences the biological activity that takes place there. Examples of contact soil layers include sand bottoms, rocky outcrops, coral, and bay mud. Description Oceans The benthic region of the ocean begins at the sho ...
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Peedee Formation
The Peedee Formation is a Formation (geology), geologic formation in North Carolina, North and South Carolina. A marine deposit representing an inner Neritic zone, neritic environment, named for exposures along the Great Peedee River, it preserves invertebrate (primarily Belemnitida, belemnites, Echinoderm, echinoderms and foraminifera) and vertebrate (primarily shark teeth, with some marine reptile remains) fossils dating to the Late Cretaceous (Maastrichtian). The formation is notable for its occurrence of ''Belemnitella americana'', known as the Pee Dee Belemnite (PDB), a long-standing standard in Delta13c, stable carbon isotope research. A single pterosaur femur, possibly an Azhdarchidae, Azhdarchid, from the Peedee formation is one of the few pterosaur body fossils found in Appalachia (landmass), Eastern North America. The stratigraphy of the formation spans from the early Maastrichtian (in South Carolina) to the late Maastrichtian shortly before the Cretaceous–Paleogene ...
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Devonian
The Devonian ( ) is a period (geology), geologic period and system (stratigraphy), system of the Paleozoic era (geology), era during the Phanerozoic eon (geology), eon, spanning 60.3 million years from the end of the preceding Silurian period at million years ago (Megaannum, Ma), to the beginning of the succeeding Carboniferous period at Ma. It is the fourth period of both the Paleozoic and the Phanerozoic. It is named after Devon, South West England, where rocks from this period were first studied. The first significant evolutionary radiation of history of life#Colonization of land, life on land occurred during the Devonian, as free-spore, sporing land plants (pteridophytes) began to spread across dry land, forming extensive coal forests which covered the continents. By the middle of the Devonian, several groups of vascular plants had evolved leaf, leaves and true roots, and by the end of the period the first seed-bearing plants (Pteridospermatophyta, pteridospermatophyt ...
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Evolution Of Plants
The evolution of plants has resulted in a wide range of complexity, from the earliest algal mats of unicellular archaeplastids evolved through endosymbiosis, through multicellular marine habitat, marine and freshwater green algae, to spore-bearing terrestrial bryophytes, lycopods and ferns, and eventually to the complex seed-bearing gymnosperms and angiosperms (flowering plants) of today. While many of the earliest groups continue to thrive, as exemplified by red algae, red and green algae in marine environments, more recently derived groups have displaced previously ecologically dominant ones; for example, the ascendance of flowering plants over gymnosperms in terrestrial environments. There is evidence that cyanobacteria and multicellular Thallus, thalloid eukaryotes lived in freshwater communities on land as early as 1 billion years ago, and that communities of complex, multicellular photosynthesizing organisms existed on land in the late Precambrian, around . Evidence of the ...
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C4 Carbon Fixation
carbon fixation or the Hatch–Slack pathway is one of three known photosynthetic processes of carbon fixation in plants. It owes the names to the 1960s discovery by Marshall Davidson Hatch and Charles Roger Slack. fixation is an addition to the ancestral and more common carbon fixation. The main carboxylating enzyme in photosynthesis is called RuBisCO, which catalyses two distinct reactions using either (carboxylation) or oxygen (oxygenation) as a substrate. RuBisCO oxygenation gives rise to phosphoglycolate, which is toxic and requires the expenditure of energy to recycle through photorespiration. photosynthesis reduces photorespiration by concentrating around RuBisCO. To enable RuBisCO to work in a cellular environment where there is a lot of carbon dioxide and very little oxygen, leaves generally contain two partially isolated compartments called mesophyll cells and bundle-sheath cells. is initially fixed in the mesophyll cells in a reaction catalysed by th ...
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C3 Carbon Fixation
carbon fixation is the most common of three metabolic pathways for carbon fixation in photosynthesis, the other two being and CAM. This process converts carbon dioxide and ribulose bisphosphate (RuBP, a 5-carbon sugar) into two molecules of 3-phosphoglycerate through the following reaction: :CO2 + H2O + RuBP → (2) 3-phosphoglycerate This reaction was first discovered by Melvin Calvin, Andrew Benson and James Bassham in 1950. C3 carbon fixation occurs in all plants as the first step of the Calvin–Benson cycle. (In and CAM plants, carbon dioxide is drawn out of malate and into this reaction rather than directly from the air.) Plants that survive solely on fixation ( plants) tend to thrive in areas where sunlight intensity is moderate, temperatures are moderate, carbon dioxide concentrations are around 200 ppm or higher, and groundwater is plentiful. The plants, originating during Mesozoic and Paleozoic eras, predate the plants and still represent approximatel ...
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Metabolic Pathway
In biochemistry, a metabolic pathway is a linked series of chemical reactions occurring within a cell (biology), cell. The reactants, products, and Metabolic intermediate, intermediates of an enzymatic reaction are known as metabolites, which are modified by a sequence of chemical reactions catalyzed by enzymes. In most cases of a metabolic pathway, the product (chemistry), product of one enzyme acts as the substrate (chemistry), substrate for the next. However, side products are considered waste and removed from the cell. Different metabolic pathways function in the position within a Eukaryotic Cell, eukaryotic cell and the significance of the pathway in the given compartment of the cell. For instance, the electron transport chain and oxidative phosphorylation all take place in the mitochondrial membrane. In contrast, glycolysis, pentose phosphate pathway, and Fatty acid synthesis, fatty acid biosynthesis all occur in the cytosol of a cell. There are two types of metabolic pathw ...
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Primary Productivity
Primary or primaries may refer to: Arts, entertainment, and media Music Groups and labels * Primary (band), from Australia * Primary (musician), hip hop musician and record producer from South Korea * Primary Music, Israeli record label Works * Primary (album), ''Primary'' (album) by Rubicon (2002) * Primary (song), "Primary" (song) by The Cure * "Primary", song by Spoon from the album ''Telephono'' Other uses in arts, entertainment, and media * Primaries or primary beams, in E. E. Smith's science-fiction series ''Lensman'' * Primary (film), ''Primary'' (film), American political documentary (1960) Computing * PRIMARY, an X Window selection * Primary data storage, computer technology used to retain digital data * Primary server, main server on the server farm Education * Primary education, the first stage of compulsory education * Primary FRCA, academic examination for anaesthetists in the U.K. * Primary school, school providing primary education Mathematics * p-group, ''p' ...
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