''Urbacodon'' ("URBAC tooth") is a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...

troodontid Troodontidae is a clade of bird-like theropod dinosaurs. During most of the 20th century, troodontid fossils were few and incomplete and they have therefore been allied, at various times, with many dinosaurian lineages. More recent fossil disco ...

dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...

, a type of small

carnivore A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose food and energy requirements derive from animal tissues (mainly muscle, fat and other s ...

. It lived in

Uzbekistan Uzbekistan (, ; uz, Ozbekiston, italic=yes / , ; russian: Узбекистан), officially the Republic of Uzbekistan ( uz, Ozbekiston Respublikasi, italic=yes / ; russian: Республика Узбекистан), is a doubly landlocked co ...

during the early

Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', ...

Period, about 95 million years ago. On 9 September 2004, a lower jaw of a small

theropod Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally c ...

was uncovered by Anton Sergeevich Rezwiy near Itemir in the IT-01 quarry. The

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specime ...

, ''Urbacodon itemirensis'', was named by Alexandr Averianov and Hans-Dieter Sues in 2007. The first part of the generic name ''Urbacodon'' is an

acronym An acronym is a word or name formed from the initial components of a longer name or phrase. Acronyms are usually formed from the initial letters of words, as in ''NATO'' (''North Atlantic Treaty Organization''), but sometimes use syllables, as ...

, honouring the Uzbek, Russian, British, American and Canadian scientists who participated in its discovery. This acronym was combined with a Greek ὀδών, ''odon'', "tooth". The specific name refers to the provenance from Itemir. The name was based on the

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of sever ...

ZIN PH 944/16, a single left

dentary In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower teeth in place. The mandible sits beneath the maxilla. It is the only movable bone ...

with preserved replacement teeth from the Cenomanian Dzharakuduk Formation. Averianov and Sues also identified teeth and other material, earlier described by Lev Nesov, as a ''Urbacodon'' sp. from the nearby

Turonian The Turonian is, in the ICS' geologic timescale, the second age in the Late Cretaceous Epoch, or a stage in the Upper Cretaceous Series. It spans the time between 93.9 ± 0.8 Ma and 89.8 ± 1 Ma (million years ago). The Turonian is preceded b ...

Bissekty Formation The Bissekty Formation (sometimes referred to as Bissekt) is a geologic formation and Lagerstätte which crops out in the Kyzyl Kum desert of Uzbekistan, and dates to the Late Cretaceous Period. Laid down in the mid to late Turonian, it is dated ...

. The holotype dentary of ''U. itemirensis'' is 79.2 millimetres long (3.12 in) and has 32 tooth positions. It is rather straight in top view. The teeth are closely packed but between the front twenty-four teeth and the rear eight teeth, a distinctive gap is present, a

diastema A diastema (plural diastemata, from Greek διάστημα, space) is a space or gap between two teeth. Many species of mammals have diastemata as a normal feature, most commonly between the incisors and molars. More colloquially, the condition ...

. This is a unique trait but was not formally designated as an

autapomorphy In phylogenetics, an autapomorphy is a distinctive feature, known as a derived trait, that is unique to a given taxon. That is, it is found only in one taxon, but not found in any others or outgroup taxa, not even those most closely related to ...

because it might be the result of individual variation. ''Urbacodon'' resembles '' Byronosaurus'' and '' Mei'' but differs from most other

Troodontidae Troodontidae is a clade of bird-like theropod dinosaurs. During most of the 20th century, troodontid fossils were few and incomplete and they have therefore been allied, at various times, with many dinosaurian lineages. More recent fossil discov ...

in that its teeth lack serrations. ''Urbacodon'' is distinguished from ''Byronosaurus'' by a less vascularized lateral dentary groove and more bulbous anterior tooth crowns, and from ''Mei'' by considerably larger size. Averianov and Sues viewed ''Urbacodon'' as more plesiomorphic than ''

Troodon ''Troodon'' ( ; ''Troödon'' in older sources) is a wastebasket taxon and a dubious genus of relatively small, bird-like dinosaurs known definitively from the Campanian age of the Late Cretaceous period (about 77 mya). It includes at leas ...

'' and ''

Saurornithoides ''Saurornithoides'' ( ) is a genus of troodontid maniraptoran dinosaur, which lived during the Late Cretaceous period. These creatures were predators, which could run fast on their hind legs and had excellent sight and hearing. The name is deri ...

'' in having a straight dentary with fewer teeth, but did not attempt to place it on a cladogram. In 2010, a

cladistic Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups ("clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived char ...

analysis showed it as a close relative of '' Byronosaurus'' and ''

Xixiasaurus ''Xixiasaurus'' () is a genus of troodontid dinosaur that lived during the Late Cretaceous Period in what is now China. The only known specimen was discovered in Xixia County, Henan Province, in central China, and became the holotype of th ...

''.Lü J.-C, Xu L., Liu Y.-Q., Zhang X.-L., Jia S. & Ji Q., 2010, "A new troodontid (Theropoda: Troodontidae) from the Late Cretaceous of central China, and the radiation of Asian troodontids", ''Acta Palaeontologica Polonica'' 55(3): 381-388

References

{{Taxonbar, from=Q1950821 Troodontids Turonian life Late Cretaceous dinosaurs of Asia Fossils of Uzbekistan Bissekty Formation Fossil taxa described in 2007 Taxa named by Hans-Dieter Sues

See also

References