''Theiophytalia'' is a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

herbivorous A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpart ...

iguanodont Iguanodontia (the iguanodonts) is a clade of herbivorous dinosaurs that lived from the Middle Jurassic to Late Cretaceous. Some members include ''Camptosaurus'', ''Dryosaurus'', ''Iguanodon'', '' Tenontosaurus'', and the hadrosaurids or "duck-bil ...

ian

dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...

from the lower

Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...

period Period may refer to: Common uses * Era, a length or span of time * Full stop (or period), a punctuation mark Arts, entertainment, and media * Period (music), a concept in musical composition * Periodic sentence (or rhetorical period), a concept ...

(

Aptian The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous Epoch or Series and encompasses the time from 121.4 ± 1.0 Ma to 113.0 ± 1.0 Ma (million years ago), a ...

Albian The Albian is both an age of the geologic timescale and a stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early/Lower Cretaceous Epoch/Series. Its approximate time range is 113.0 ± 1.0 Ma to 100.5 ± 0.9 M ...

stage, about 112 million years ago) of

Colorado Colorado (, other variants) is a state in the Mountain West subregion of the Western United States. It encompasses most of the Southern Rocky Mountains, as well as the northeastern portion of the Colorado Plateau and the western edge of t ...

, USA. It contains a single species, ''T. kerri''.

Description

Detailed comparisons by Brill and Carpenter (2006) also showed that the skull differed in a number of key features from that of ''Camptosaurus'', namely: a longer, heavier, and more rugose snout; a wider dorsal process on the maxilla; a proportionally smaller

antorbital fenestra An antorbital fenestra (plural: fenestrae) is an opening in the skull that is in front of the eye sockets. This skull character is largely associated with archosauriforms, first appearing during the Triassic Period. Among extant archosaurs, birds ...

; and stouter quadrate, with a bulbous articulation for the lower jaw. Compare the skull image with that of ''

Camptosaurus ''Camptosaurus'' ( ) is a genus of plant-eating, beaked ornithischian dinosaurs of the Late Jurassic period of western North America and possibly also Europe. The name means 'flexible lizard' ( Greek (') meaning 'bent' and (') meaning 'li ...

''. Therefore, they put it into its own genus and species.

Discovery

In 1878, a student of

Colorado College Colorado College is a private liberal arts college in Colorado Springs, Colorado. It was founded in 1874 by Thomas Nelson Haskell in his daughter's memory. The college enrolls approximately 2,000 undergraduates at its campus. The college offer ...

named James Kerr found a partial Ornithopod skull at Garden of the Gods Park in El Paso County,

. The skull, YPM 1887, was referred by

O.C. Marsh Othniel Charles Marsh (October 29, 1831 – March 18, 1899) was an American professor of Paleontology in Yale College and President of the National Academy of Sciences. He was one of the preeminent scientists in the field of paleontology. Among h ...

(affirmed by Gilmore, 1909), to whom the skull was given in 1886, as that of ''

amplus''. Gilmore used the skull to reconstruct the skull of ''Camptosaurus'' assuming that it came from the

Morrison Formation The Morrison Formation is a distinctive sequence of Late Jurassic, Upper Jurassic sedimentary rock found in the western United States which has been the most fertile source of dinosaur fossils in North America. It is composed of mudstone, sandsto ...

. However, microscopic comparisons of thin-sections of the Mesozoic formations in the

Garden of the Gods Garden of the Gods (Arapaho: ''Ho3o’uu Niitko’usi’i'') is a public park located in Colorado Springs, Colorado, United States. It was designated a National Natural Landmark in 1971. Name The area now known as Garden of the Gods was fir ...

showed that the specimen actually came from the Lytle Member of the Purgatoire Formation; therefore, the skull was Early Cretaceous in age. The generic name is, from Greek, θειος, ''theios'': "divine" + Greek φυταλία, ''phytalia'': "garden", or "garden of the gods".

is a park in

Colorado Springs Colorado Springs is a home rule municipality in, and the county seat of, El Paso County, Colorado, United States. It is the largest city in El Paso County, with a population of 478,961 at the 2020 United States Census, a 15.02% increase since ...

, Colorado, where a skull, the only fossil of the genus yet to be discovered, was found in 1878. The specific name ''kerri'' honors James Hutchinson Kerr, who found the specimen.

Classification

The article describing the find classified ''Theiophytalia'' as intermediate in derivation between ''Camptosaurus'' and ''

Iguanodon ''Iguanodon'' ( ; meaning 'iguana-tooth'), named in 1825, is a genus of iguanodontian dinosaur. While many species have been classified in the genus ''Iguanodon'', dating from the late Jurassic Period to the early Cretaceous Period of Asia, Eu ...

''. The

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...

is ''Theiophytalia kerri''. In 2010 and 2011

cladistic Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups (" clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived char ...

analyses of McDonald and colleagues, ''Theiophytalia'' has been recovered as a basal member of the Styracosterna and its closest relative was '' Hippodraco''.

References

External links

Dinosaur Mailing List entry, which announces the new genus
{{Taxonbar, from=Q1655417 Early Cretaceous dinosaurs of North America Iguanodonts Fossil taxa described in 2006 Taxa named by Kenneth Carpenter Paleontology in Colorado Ornithischian genera