Pilgerodendron uviferum
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''Pilgerodendron'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of conifer belonging to the cypress family Cupressaceae. It has only one
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
, ''Pilgerodendron uviferum'', which is
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found else ...
to the
Valdivian temperate rain forests The Valdivian temperate forests (NT0404) is an ecoregion on the west coast of southern South America, in Chile and Argentina. It is part of the Neotropical realm. The forests are named after the city of Valdivia. The Valdivian temperate rainforest ...
and
Magellanic subpolar forests The Magellanic subpolar forests () are a terrestrial ecoregion of southernmost South America, covering parts of southern Chile and Argentina, and are part of the Neotropical realm. It is a temperate broadleaf and mixed forests ecoregion, and ...
of southern
Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the east a ...
and southwestern
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
. It grows from 40 to 55°S in
Tierra del Fuego Tierra del Fuego (, ; Spanish for "Land of the Fire", rarely also Fireland in English) is an archipelago off the southernmost tip of the South American mainland, across the Strait of Magellan. The archipelago consists of the main island, Isla ...
, where it is the southernmost conifer in the world. It is a member of subfamily
Callitroideae Cupressaceae is a conifer family, the cypress family, with worldwide distribution. The family includes 27–30 genera (17 monotypic), which include the junipers and redwoods, with about 130–140 species in total. They are monoecious, subdioecio ...
, a group of distinct Southern Hemisphere genera associated with the
Antarctic flora Antarctic flora are a distinct community of vascular plants which evolved millions of years ago on the supercontinent of Gondwana. Presently, species of Antarctica flora reside on several now separated areas of the Southern Hemisphere, includin ...
.Farjon, A. (2005). ''Monograph of Cupressaceae and Sciadopitys''. Royal Botanic Gardens, Kew. Flora Chilena
''Pilgerodendron uviferum''
/ref> ''Pilgerodendron'' is very closely related to the
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...
and New Caledonian genus '' Libocedrus'', and many botanists treat ''P. uviferum'' within this genus, as ''Libocedrus uvifera'' (D.Don) Pilg.Eckenwalder, J. E. (1976). Re-evaluation of Cupressaceae and Taxodiaceae: a proposed merger. ''Madroño'' 23 (5): 237-256. It is also a
taxonomical In biology, taxonomy () is the scientific study of naming, defining ( circumscribing) and classifying groups of biological organisms based on shared characteristics. Organisms are grouped into taxa (singular: taxon) and these groups are given ...
synonym for ''Libocedrus tetragona'' (Hooker). The species is known locally as ciprés de las GuaitecasChilebosque
''Pilgerodendron uviferum''
/ref> (after the
Guaitecas Archipelago Guaitecas Archipelago is a sparsely populated archipelago in the Aisén region of Chile. The archipelago is made up of eight main islands and numerous smaller ones. The eight largest islands are from northwest to southeast: Gran Guaiteca, Ascenc ...
), and elsewhere by its scientific name, as pilgerodendron. The genus is named after
Robert Knud Friedrich Pilger Robert Knud Friedrich Pilger (3 July 1876, in Helgoland – 1 September 1953, in Berlin)
. ''P. uviferum'' is a dioecious, slow-growing, narrowly conical
evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, whic ...
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
that grows from 2–20 m in height, with a trunk up to 1.5 m in diameter. Taller specimens and wider trunks (reportedly up to 3 m in diameter) are known to have existed before it was overexploited. The leaves are scale-like and arranged in decussate pairs, with each leaf equal in size, giving the
shoot In botany, a plant shoot consists of any plant stem together with its appendages, leaves and lateral buds, flowering stems, and flower buds. The new growth from seed germination that grows upward is a shoot where leaves will develop. In the sp ...
s a square cross-section (unlike the ''Libocedrus'' species, where pairs of larger leaves alternate with pairs of smaller leaves, giving a somewhat flattened shoot). The
seed cones A conifer cone (in formal botanical usage: strobilus, plural strobili) is a seed-bearing organ on gymnosperm plants. It is usually woody, ovoid to globular, including scales and bracts arranged around a central axis, especially in conifers an ...
are 5–12 mm long and 4–6 mm broad, with four scales, two sterile basal scales and two fertile scales; each scale has a slender spine-like bract, and each fertile scale has two winged
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiospe ...
s 3–4 mm long. The pollen cones are 5–10 mm long and 2 mm broad, with 12–20 scales. It is found in the evergreen coastal lowland forests along the
Pacific The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the contine ...
coast of southern
Patagonia Patagonia () refers to a geographical region that encompasses the southern end of South America, governed by Argentina and Chile. The region comprises the southern section of the Andes Mountains with lakes, fjords, temperate rainforests, and g ...
, in association with the broadleaf evergreens ''
Nothofagus betuloides ''Nothofagus betuloides'', Magellan's beech or ''guindo'', is a tree native to southern Patagonia. In 1769, Sir Joseph Banks collected a specimen of the tree in Tierra del Fuego during Captain Cook's first voyage. Its occurrence on Hornos Isl ...
'' and ''
Drimys winteri ''Drimys winteri'', the winter's bark or canelo, is a slender tree in the family Winteraceae, growing up to tall. It is native to the Magellanic and Valdivian temperate rain forests of Chile and Argentina, where it is a dominant tree in the coas ...
''. It is also found in open stands in sheltered bogs farther inland, where it is often locally dominant, and ranges as far as the eastern slopes of the
Andes The Andes, Andes Mountains or Andean Mountains (; ) are the longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range is long, wide (widest between 18°S – 20°S ...
in southwestern Argentina. At the northern end of its range it is found in association with '' Fitzroya cupressoides''. It has also been planted along the northern part of the Pacific Coast of the
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territori ...
. At present much ''Pilgerodendron uviferum'' grow in the Andes and in the
Chilean Coast Range The Chilean Coastal Range ( es, Cordillera de la Costa) is a mountain range that runs from north to south along the Pacific coast of South America parallel to the Andean Mountains, extending from Morro de Arica in the north to Taitao Peninsula, ...
. However, during the
interstadial Stadials and interstadials are phases dividing the Quaternary period, or the last 2.6 million years. Stadials are periods of colder climate while interstadials are periods of warmer climate. Each Quaternary climate phase is associated with a Ma ...
s of the region's last glacial period, ''P. uviferum'' grew in lowland areas such as the Central Valley, where it is now absent. Remaining lowland populations are thought to be
relicts A relict is a surviving remnant of a natural phenomenon. Biology A relict (or relic) is an organism that at an earlier time was abundant in a large area but now occurs at only one or a few small areas. Geology and geomorphology In geology, a r ...
that have survived the warmer climate of the
Holocene The Holocene ( ) is the current geological epoch. It began approximately 11,650 cal years Before Present (), after the Last Glacial Period, which concluded with the Holocene glacial retreat. The Holocene and the preceding Pleistocene togeth ...
. The
wood Wood is a porous and fibrous structural tissue found in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulose fibers that are strong in tension and embedded in a matrix of lignin ...
of ''P. uviferum'' is yellow-reddish and has a distinct spicy-resinous smell, and is highly resistant to decay, which has made it very valuable as a source of
timber Lumber is wood that has been processed into dimensional lumber, including beams and planks or boards, a stage in the process of wood production. Lumber is mainly used for construction framing, as well as finishing (floors, wall panels, w ...
for building construction in its native range. Much of its original lowland habitat has been cleared for this and other reasons. Listed as Vulnerable (VU A2cd v2.3) Due to
over-exploitation Overexploitation, also called overharvesting, refers to harvesting a renewable resource to the point of diminishing returns. Continued overexploitation can lead to the destruction of the resource, as it will be unable to replenish. The term ap ...
, the species is now much scarcer than formerly, and is accordingly listed under
CITES CITES (shorter name for the Convention on International Trade in Endangered Species of Wild Fauna and Flora, also known as the Washington Convention) is a multilateral treaty to protect endangered plants and animals from the threats of interna ...
Appendix I, meaning commercial international trade is prohibited. ''P. uviferum'' is considered threatened by the
World Conservation Monitoring Centre The UN Environment Programme World Conservation Monitoring Centre (UNEP-WCMC) is a collaboration centre of UN Environment Programme, based in Cambridge in the United Kingdom. UNEP-WCMC has been part of UN Environment Programme since 2000, and has ...
and vulnerable by the
International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of nat ...
.


History

Indigenous sea-faring nomads of the Chono and Kawésqars groups used ''Pilgerodendron uviferum'' as firewood as well as wood for rows, boats and houses. During the
Antonio de Vea expedition The Antonio de Vea expedition of 1675–1676 was a Spanish naval expedition to the fjords and channels of Patagonia aimed to find whether rival colonial powers—specifically, the English—were active in the region. While this was not the first ...
(1675–1676) Spanish explorers visited the Guaitecas and Chonos archipelagoes where the tree grew noticing the similarity to the "cypresses of Spain". Historically the wood was used to make railway sleepers, but eventually became popular for use as timber in all types of construction. In the 1860s, Felipe Westhoff was one of various businessmen who exported ''Pilgerodendron uviferum'' north to Chilean and Peruvian ports. Westhoff, who was based in
Ancud Ancud () is a city in southern Chile located in the northernmost part of the island and province of Chiloé, in Los Lagos Region. It is the second largest city of Chiloé Archipelago after Castro. The city was established in 1768 to function as ...
, came initially as an agent of
Ferrocarril Central Andino Ferrocarril Central Andino (FCCA) is the consortium which operates the Ferrovías Central railway in Peru linking the Pacific port of Callao and the capital Lima with Huancayo and Cerro de Pasco. As one of the Trans-Andean Railways it is the sec ...
in Peru to purchase sleepers but soon became an independent businessman. The town of
Melinka Melinka is a Chilean town in Aysén Province, Aysén Region. It is located on Ascención Island and is the administrative center of the commune of Guaitecas since 1979. The town is on a small peninsula off the main island, and shelters a small ...
was established in 1860 during the ''Pilgerodendron'' boom. This was the first permanent settlement in the Guaitecas Archipelago. After Westhoff's retirement in the 1870s, Ciriaco Álvarez, a native of
Chonchi Chonchi is a Chilean town and commune located in Los Lagos Region. The commune consists of the town of Chonchi and the following villages: * Rauco * Quinched * Vilupulli * Teupa * Terao * Petanes * Notue * Quiao * Huillinco * Cucao Demographics ...
, rose as the most prominent ''P. uviferum'' businessman in the area, and was dubbed "The King of Pilgerodendron" ( es, El Rey del Ciprés). Many of the wooden structures in settlements of the southern Chilean archipelagos are built largely from ''P. uviferum'' wood.


References

{{Taxonbar, from=Q133103 Cupressaceae Valdivian temperate rainforest Trees of mild maritime climate Trees of subpolar oceanic climate Monotypic conifer genera Garden plants of South America Ornamental trees Dioecious plants