Pinophyta
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Pinophyta
Conifers are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class, Pinopsida. All extant conifers are perennial woody plants with secondary growth. The great majority are trees, though a few are shrubs. Examples include cedars, Douglas-firs, cypresses, firs, junipers, kauri, larches, pines, hemlocks, redwoods, spruces, and yews.Campbell, Reece, "Phylum Coniferophyta". Biology. 7th. 2005. Print. P. 595 As of 1998, the division Pinophyta was estimated to contain eight families, 68 genera, and 629 living species. Although the total number of species is relatively small, conifers are ecologically important. They are the dominant plants over large areas of land, most notably the taiga of the Northern Hemisphere, but also in similar cool climates in mountains further south. Boreal conifers have many wintertime ad ...
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Pinales
The order Pinales in the division Pinophyta, class Pinopsida, comprises all the extant conifers. The distinguishing characteristic is the reproductive structure known as a cone produced by all Pinales. All of the extant conifers, such as cedar, celery-pine, cypress, fir, juniper, larch, pine, redwood, spruce, yew and '' Araucaria araucana'' ("Monkey tail tree" or "Monkey puzzle tree") are included here. Some fossil conifers, however, belong to other distinct orders within the division Pinophyta. Multiple molecular studies indicate this order being paraphyletic with respect to Gnetales, with studies recovering Gnetales as either a sister group to Pinaceae or being more derived than Pinaceae but sister to the rest of the group. Taxonomy History Brown (1825) first discerned that there were two groups of seed plants, distinguished by the form of seed development, based on whether the ovules were exposed, receiving pollen directly, or enclosed, which do not. Shortly afterwa ...
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Spermatophyte
A spermatophyte (; ), also known as phanerogam (taxon Phanerogamae) or phaenogam (taxon Phaenogamae), is any plant that produces seeds, hence the alternative name seed plant. Spermatophytes are a subset of the embryophytes or land plants. They include most familiar types of plants, including all flowers and most trees, but exclude some other types of plants such as ferns, mosses, algae. The term ''phanerogams'' or ''phanerogamae'' is derived from the Greek (), meaning "visible", in contrast to the cryptogamae (), together with the suffix (), meaning "to marry". These terms distinguished those plants with hidden sexual organs (cryptogamae) from those with visible sexual organs (phanerogamae). Description The extant spermatophytes form five divisions, the first four of which are traditionally grouped as gymnosperms, plants that have unenclosed, "naked seeds": * Cycadophyta, the cycads, a subtropical and tropical group of plants, * Ginkgophyta, which includes a single liv ...
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Gymnosperm
The gymnosperms ( lit. revealed seeds) are a group of seed-producing plants that includes conifers, cycads, '' Ginkgo'', and gnetophytes, forming the clade Gymnospermae. The term ''gymnosperm'' comes from the composite word in el, γυμνόσπερμος ( el, γυμνός, translit=gymnos, lit=naked, label=none and el, σπέρμα, translit=sperma, lit=seed, label=none), literally meaning 'naked seeds'. The name is based on the unenclosed condition of their seeds (called ovules in their unfertilized state). The non-encased condition of their seeds contrasts with the seeds and ovules of flowering plants ( angiosperms), which are enclosed within an ovary. Gymnosperm seeds develop either on the surface of scales or leaves, which are often modified to form cones, or solitary as in yew, '' Torreya'', '' Ginkgo''. Gymnosperm lifecycles involve alternation of generations. They have a dominant diploid sporophyte phase and a reduced haploid gametophyte phase which is dep ...
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Araucariales
Araucariaceae – also known as araucarians – is an extremely ancient family of coniferous trees. The family achieved its maximum diversity during the Jurassic and Cretaceous periods and the early Cenozoic, when it was distributed almost worldwide. Most of the Araucariaceae in the Northern Hemisphere vanished in the Cretaceous–Paleogene extinction event, and they are now largely confined to the Southern Hemisphere, except for a few species of ''Agathis'' in Southeast Asia. Description Members of Araucariaceae are typically extremely tall evergreen trees, reaching heights of or more. They can also grow very large stem diameters; a New Zealand kauri tree (''Agathis australis'') named ''Tāne Mahuta'' ("The Lord of the Forest") has been measured at tall with a diameter at breast height of . Its total wood volume is calculated to be , making it the third-largest conifer after '' Sequoia'' and ''Sequoiadendron'' (both from the Cupressaceae subfamily Sequoioideae). The trunk ...
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Araucariaceae
Araucariaceae – also known as araucarians – is an extremely ancient family of coniferous trees. The family achieved its maximum diversity during the Jurassic and Cretaceous periods and the early Cenozoic, when it was distributed almost worldwide. Most of the Araucariaceae in the Northern Hemisphere vanished in the Cretaceous–Paleogene extinction event, and they are now largely confined to the Southern Hemisphere, except for a few species of '' Agathis'' in Southeast Asia. Description Members of Araucariaceae are typically extremely tall evergreen trees, reaching heights of or more. They can also grow very large stem diameters; a New Zealand kauri tree ('' Agathis australis'') named '' Tāne Mahuta'' ("The Lord of the Forest") has been measured at tall with a diameter at breast height of . Its total wood volume is calculated to be , making it the third-largest conifer after '' Sequoia'' and '' Sequoiadendron'' (both from the Cupressaceae subfamily Sequoioideae). The ...
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Pinaceae
The Pinaceae, or pine family, are conifer trees or shrubs, including many of the well-known conifers of commercial importance such as cedars, firs, hemlocks, larches, pines and spruces. The family is included in the order Pinales, formerly known as Coniferales. Pinaceae are supported as monophyletic by their protein-type sieve cell plastids, pattern of proembryogeny, and lack of bioflavonoids. They are the largest extant conifer family in species diversity, with between 220 and 250 species (depending on taxonomic opinion) in 11 genera, and the second-largest (after Cupressaceae) in geographical range, found in most of the Northern Hemisphere, with the majority of the species in temperate climates, but ranging from subarctic to tropical. The family often forms the dominant component of boreal, coastal, and montane forests. One species, ''Pinus merkusii'', grows just south of the equator in Southeast Asia. Major centres of diversity are found in the mountains of southwest ...
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Phylum
In biology, a phylum (; plural: phyla) is a level of classification or taxonomic rank below kingdom and above class. Traditionally, in botany the term division has been used instead of phylum, although the International Code of Nomenclature for algae, fungi, and plants accepts the terms as equivalent. Depending on definitions, the animal kingdom Animalia contains about 31 phyla, the plant kingdom Plantae contains about 14 phyla, and the fungus kingdom Fungi contains about 8 phyla. Current research in phylogenetics is uncovering the relationships between phyla, which are contained in larger clades, like Ecdysozoa and Embryophyta. General description The term phylum was coined in 1866 by Ernst Haeckel from the Greek (, "race, stock"), related to (, "tribe, clan"). Haeckel noted that species constantly evolved into new species that seemed to retain few consistent features among themselves and therefore few features that distinguished them as a group ("a self-contained un ...
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Cephalotaxaceae
Cephalotaxaceae is a small grouping of conifers, that included one to three genera closely allied to Taxaceae. However, members of Cephalotaxaceae are now included in Taxaceae by botanists, instead of as a distinct family, based on phylogenetic evidence and close morphological similarities between them. Included species were restricted to east Asia, except for two species of ''Torreya'' found in the southwest and southeast of the United States; fossil evidence shows a much wider prehistorical Northern Hemisphere distribution. The most notable differences between Taxaceae and Cephalotaxaceae concerned the cone aril, which fully encloses the seeds of Cephalotaxaceae, the longer maturation of Cephalotaxaceae seeds and the larger size of the mature seeds. When considered as a distinct family, members of Cephalotaxaceae are much branched, small trees and shrubs. The leaves are evergreen, spirally arranged, often twisted at the base to appear biranked. They are linear to lanceolate, an ...
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Conifer Cone
A conifer cone (in formal botanical usage: strobilus, plural strobili) is a seed-bearing organ on gymnosperm plants. It is usually woody, ovoid to globular, including scales and bracts arranged around a central axis, especially in conifers and cycads. The cone of Pinophyta (conifer clade) contains the reproductive structures. The woody cone is the female cone, which produces seeds. The male cone, which produces pollen, is usually herbaceous and much less conspicuous even at full maturity. The name "cone" derives from Greek ''konos'' (pine cone), which also gave name to the geometric cone. The individual plates of a cone are known as ''scales''. The ''umbo'' of a conifer cone refers to the first year's growth of a seed scale on the cone, showing up as a protuberance at the end of the two-year-old scale. The male cone (microstrobilus or pollen cone) is structurally similar across all conifers, differing only in small ways (mostly in scale arrangement) from species to s ...
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Pinidae
Pinidae is a subclass of Equisetopsida in the sense used by Mark W. Chase and James L. Reveal in their 2009 article "A phylogenetic classification of the land plants to accompany APG III." This subclass comprises the conifers. The Pinidae subclass is equivalent to the division Pinophyta Conifers are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class, Pinopsida. All exta ... and class Pinopsida of previous treatments. There are over 600 species of Pinidae all over the world. Phylogeny The following diagram shows a likely phylogenic relationship between subclass Pinidae and the other Equisetopsida subclasses. References Plant subclasses Equisetopsida sensu lato {{Conifer-stub ...
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Taxaceae
Taxaceae (), commonly called the yew family, is a coniferous family which includes six extant and two extinct genera, and about 30 species of plants, or in older interpretations three genera and 7 to 12 species. Description They are many-branched, small trees and shrubs. The leaves are evergreen, spirally arranged, often twisted at the base to appear 2-ranked. They are linear to lanceolate, and have pale green or white stomatal bands on the undersides. The plants are dioecious, or rarely monoecious. The catkin like male cones are long, and shed pollen in the early spring. They are sometimes externally only slightly differentiated from the branches. The fertile bracts have 2-8 pollen sacs. The female 'cones' are highly reduced. Only the upper or uppermost bracts are fertile and bear one or rarely two seeds. The ovule usually exceeds the scale, although ovules are sometimes rarely enclosed by it. They may be found on the ends of branches or on the branches. They may grow ...
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Sciadopityaceae
''Sciadopitys verticillata'', the or Japanese umbrella-pine, is a unique conifer endemic to Japan. It is the sole member of the family Sciadopityaceae and genus ''Sciadopitys'', a living fossil with no close relatives. The oldest fossils of ''Sciadopitys'' are from the Late Cretaceous of Japan, and the genus was widespread in Laurasia during most of the Cenozoic, especially in Europe until the Pliocene. Taxonomy Molecular evidence indicates that Sciadopityaceae is the sister group to a clade comprising Taxaceae and Cupressaceae, and has an extremely ancient divergence, having diverged from the rest of the conifers during the early mid- Permian; this would also make it a survivor of the Permian–Triassic extinction event. There is inconsistent evidence regarding the plant family which produced Baltic amber. Both macrofossil and microfossil evidence suggest a ''Pinus'' relative, whereas chemical and infrared microspectroscopy evidence suggest relatives of either ''Ag ...
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