Euornithes (from

Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...

' meaning "true birds") is a natural group which includes the most recent common ancestor of all

avialan Avialae ("bird wings") is a clade containing the only living dinosaurs, the birds. It is usually defined as all theropod dinosaurs more closely related to birds (Aves) than to deinonychosaurs, though alternative definitions are occasionally used ...

s closer to modern

bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...

s than to ''

Sinornis ''Sinornis'' is a genus of enantiornithean birds from the Lower Cretaceous Jiufotang Formation of the People's Republic of China. When it was described in 1992, this 120 million-year-old sparrow-sized skeleton represented a new avian sharing "pr ...

''.

Description

Clarke ''et al''. (2006) found that the most primitive known euornithians (the Yanornithiformes) had a mosaic of advanced and primitive features. These species retained primitive features like

gastralia Gastralia (singular gastralium) are dermal bones found in the ventral body wall of modern crocodilians and tuatara, and many prehistoric tetrapods. They are found between the sternum and pelvis, and do not articulate with the vertebrae. In thes ...

and a

pubic symphysis The pubic symphysis is a secondary cartilaginous joint between the left and right superior rami of the pubis of the hip bones. It is in front of and below the urinary bladder. In males, the suspensory ligament of the penis attaches to the pubi ...

. They also showed the first fully modern

pygostyle Pygostyle describes a skeletal condition in which the final few caudal vertebrae are fused into a single ossification, supporting the tail feathers and musculature. In modern birds, the rectrices attach to these. The pygostyle is the main compone ...

s, and the type specimen of ''

Yixianornis ''Yixianornis'' (meaning "Yixian Formation bird") is a bird genus from the early Cretaceous period. Its remains have been found in the Jiufotang Formation at Chaoyang (People's Republic of China) dated to the early Aptian age, around 120 millio ...

'' (IVPP 13631) preserves eight elongated

rectrices Flight feathers (''Pennae volatus'') are the long, stiff, asymmetrically shaped, but symmetrically paired pennaceous feathers on the wings or tail of a bird; those on the wings are called remiges (), singular remex (), while those on the tail ...

(tail feathers) in a modern arrangement. No earlier

pygostylia Pygostylia is a group of avialans which includes the Confuciusornithidae and all of the more advanced species, the Ornithothoraces. Definition The group Pygostylia was intended to encompass all avialans with a short, stubby tail, as opposed to ...

ns are known which preserve a fan of tail feathers of this sort; instead, they showed only paired plumes or a tuft of short feathers.

Classification

The name Euornithes has been used for a wide variety of avialan groups since it was first named by Edward Drinker Cope in 1889. It was first defined as a clade in 1998 by

Paul Sereno Paul Callistus Sereno (born October 11, 1957) is a professor of paleontology at the University of Chicago and a National Geographic "explorer-in-residence" who has discovered several new dinosaur species on several continents, including at sites ...

, who made it the group of all animals closer to birds than to Enantiornithes (represented by ''Sinornis''). This definition currently includes similar content as another widely used name, Ornithuromorpha, named and defined by Luis Chiappe in 1999 as the common ancestor of '' Patagopteryx'', ''

Vorona ''Vorona'' ( ; Malagasy for "bird", ''V. berivotrensis'', "from Berivotra") is a monotypic genus of prehistoric birds. It was described from fossils found in a Maevarano Formation quarry near the village of Berivotra, Mahajanga Province, Madag ...

'', and Ornithurae, plus all of its descendants. Because one definition is node-based and the other branch-based, Ornithuromorpha is a slightly less inclusive group.

Relationships

The cladogram below follows the results of a phylogenetic analysis by Lee ''et al.'', 2014:

Other genera

The following is a list of primitive euornithian genera and those that cannot be confidently referred to any subgroups, following Holtz (2011) unless otherwise noted.Holtz, Thomas R. Jr. (2012) ''Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages,'
Winter 2011 Appendix.
/ref> *†'' Alamitornis'' *†'' Changmaornis'' *†'' Changzuiornis''Huang, J., Wang, X., Hu, Y., Liu, J., Peteya, J. A., & Clarke, J. A. (2016). A new ornithurine from the Early Cretaceous of China sheds light on the evolution of early ecological and cranial diversity in birds. ''PeerJ'', 4: e1765. *†'' Dingavis'' *†'' Gargantuavis'' *†'' Horezmavis'' *†'' Iteravis'' *†'' Juehuaornis'' *†'' Platanavis'' *†'' Wyleyia''? *†'' Yumenornis'' *'' Xinghaiornis'' *†'' Zhyraornis'' Note that Holtz also included the genera '' Eurolimnornis'' and '' Piksi'' as euornitheans, though they have since been re-identified as

pterosaur Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 ...

References

External links

{{Taxonbar, from=Q2752642 Evolution of birds Extant Early Cretaceous first appearances Taxa named by Edward Drinker Cope