Pygostylia
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Pygostylia
Pygostylia is a group of avialans which includes the Confuciusornithidae and all of the more advanced species, the Ornithothoraces. Definition The group Pygostylia was intended to encompass all avialans with a short, stubby tail, as opposed to the longer, unfused tails of more primitive species like ''Archaeopteryx lithographica''. It was named by Sankar Chatterjee in 1997.Chatterjee, S. 1997. ''The Rise of Birds.'' Baltimore: Johns Hopkins University Press. 312 pp. Luis Chiappe later defined Pygostylia as a node-based clade, "the common ancestor of the Confuciusornithidae and Neornithes plus all its descendants". In 2001, Jacques Gauthier and Kevin de Queiroz recommended that Chatterjee's original apomorphy-based clade concept be used instead of Chiappe's node-based definition,Gauthier, J., & De Queiroz, K. (2001). Feathered dinosaurs, flying dinosaurs, crown dinosaurs, and the name" Aves. ''New perspectives on the origin and early evolution of birds'', 7-41. but this recommendat ...
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Jinguofortisidae
Jinguofortisidae is an extinct family of early pygostylian avialans known from the Early Cretaceous, found in northern China. Description The Jinguofortisidae are distinguished from other non- ornithothoracine avialans by the following features: the scapula and coracoid fused into a scapulocoracoid; a boomeranged-shaped furcula without a hypocleidium; proximal margin of humerus concave centrally; deltopectoral crest large and not perforated; minor metacarpal markedly bowed caudally; alular digit terminating at the level of the distal end of major metacarpal; and proximal phalanx of the major digit longer than the intermediate phalanx. Phylogeny Wang et al. (2018) erected Jinguofortisidae after finding '' Chongmingia'' (previously considered Avebrevicauda ''incertae sedis'' by Wang et al. 2016) to be sister to their new taxon ''Jinguofortis''. In 2023, a third jinguofortisid was described by Li ''et al''. They recovered their new taxon, '' Cratonavis'', as the sister taxon to ' ...
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Confuciusornithidae
Confuciusornithidae is an extinct family of pygostylian avialans known from the Early Cretaceous, found in northern China. They are commonly placed as a sister group to Ornithothoraces, a group that contains all extant birds along with their closest extinct relatives. Confuciusornithidae contains four genera, possessing both shafted and non-shafted (downy) feathers. They are also noted for their distinctive pair of ribbon-like tail feathers of disputed function. The wing anatomy of confuciusornithids suggests an unusual flight behavior, due to anatomy that implies conflicting abilities. They possessed feathers similar to those of fast-flapping birds, which rely on quick flapping of their wings to stay aloft. At the same time, their wing anatomy also suggests a lack of flapping ability. Confuciusornithids are also noted for their beak and lack of teeth, similar to modern birds. Both predators and prey, confuciusornithid fossils have been observed with fish remains in their digestive ...
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Confuciusornithiformes
Confuciusornithidae is an extinct family of pygostylian avialans known from the Early Cretaceous, found in northern China. They are commonly placed as a sister group to Ornithothoraces, a group that contains all extant birds along with their closest extinct relatives. Confuciusornithidae contains four genera, possessing both shafted and non-shafted (downy) feathers. They are also noted for their distinctive pair of ribbon-like tail feathers of disputed function. The wing anatomy of confuciusornithids suggests an unusual flight behavior, due to anatomy that implies conflicting abilities. They possessed feathers similar to those of fast-flapping birds, which rely on quick flapping of their wings to stay aloft. At the same time, their wing anatomy also suggests a lack of flapping ability. Confuciusornithids are also noted for their beak and lack of teeth, similar to modern birds. Both predators and prey, confuciusornithid fossils have been observed with fish remains in their digestive ...
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Euornithes
Euornithes (from Greek ' meaning "true birds") is a natural group which includes the most recent common ancestor of all avialans closer to modern birds than to ''Sinornis''. Description Clarke ''et al''. (2006) found that the most primitive known euornithians (the Yanornithiformes) had a mosaic of advanced and primitive features. These species retained primitive features like gastralia and a pubic symphysis. They also showed the first fully modern pygostyles, and the type specimen of ''Yixianornis'' (IVPP 13631) preserves eight elongated rectrices (tail feathers) in a modern arrangement. No earlier pygostylians are known which preserve a fan of tail feathers of this sort; instead, they showed only paired plumes or a tuft of short feathers. Classification The name Euornithes has been used for a wide variety of avialan groups since it was first named by Edward Drinker Cope in 1889. It was first defined as a clade in 1998 by Paul Sereno, who made it the group of all animals closer t ...
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Early Cretaceous
The Early Cretaceous ( geochronological name) or the Lower Cretaceous (chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145  Ma to 100.5 Ma. Geology Proposals for the exact age of the Barremian-Aptian boundary ranged from 126 to 117 Ma until recently (as of 2019), but based on drillholes in Svalbard the defining early Aptian Oceanic Anoxic Event 1a (OAE1a) was carbon isotope dated to 123.1±0.3 Ma, limiting the possible range for the boundary to c. 122–121 Ma. There is a possible link between this anoxic event and a series of Early Cretaceous large igneous provinces (LIP). The Ontong Java-Manihiki-Hikurangi large igneous province, emplaced in the South Pacific at c. 120 Ma, is by far the largest LIP in Earth's history. The Ontong Java Plateau today covers an area of 1,860,000 km2. In the Indian Ocean another LIP began to form at c. 120 Ma, the Kerguelen P ...
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Pubis (bone)
In vertebrates, the pubic region ( la, pubis) is the most forward-facing (ventral and anterior) of the three main regions making up the coxal bone. The left and right pubic regions are each made up of three sections, a superior ramus, inferior ramus, and a body. Structure The pubic region is made up of a ''body'', ''superior ramus'', and ''inferior ramus'' (). The left and right coxal bones join at the pubic symphysis. It is covered by a layer of fat, which is covered by the mons pubis. The pubis is the lower limit of the suprapubic region. In the female, the pubic region is anterior to the urethral sponge. Body The body forms the wide, strong, middle and flat part of the pubic region. The bodies of the left and right pubic regions join at the pubic symphysis. The rough upper edge is the pubic crest, ending laterally in the pubic tubercle. This tubercle, found roughly 3 cm from the pubic symphysis, is a distinctive feature on the lower part of the abdominal wall; important ...
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Synapomorphies
In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to have evolved in their most recent common ancestor. ) In cladistics, synapomorphy implies homology. Examples of apomorphy are the presence of erect gait, fur, the evolution of three middle ear bones, and mammary glands in mammals but not in other vertebrate animals such as amphibians or reptiles, which have retained their ancestral traits of a sprawling gait and lack of fur. Thus, these derived traits are also synapomorphies of mammals in general as they are not shared by other vertebrate animals. Etymology The word —coined by German entomologist Willi Hennig—is derived from the Ancient Greek words (''sún''), meaning "with, together"; (''apó''), meaning "away from"; and (''morphḗ''), meaning "shape, form". Clade analysis T ...
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Pygostyle
Pygostyle describes a skeletal condition in which the final few caudal vertebrae are fused into a single ossification, supporting the tail feathers and musculature. In modern birds, the rectrices attach to these. The pygostyle is the main component of the uropygium, a structure colloquially known as the bishop's nose, parson's nose, pope's nose, or sultan's nose. This is the fleshy protuberance visible at the posterior end of a bird (most commonly a chicken or turkey) that has been dressed for cooking. It has a swollen appearance because it also contains the uropygial gland that produces preen oil. Evolution Pygostyles probably began to evolve very early in the Cretaceous period, perhaps 140 – 130 million years ago. The earliest known species to have evolved a pygostyle were members of the Confuciusornithidae. The structure provided an evolutionary advantage, as a completely mobile tail as found in species such as '' Archaeopteryx'' is detrimental to its use for flight contro ...
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Hyposphene
The hyposphene-hypantrum articulation is an accessory joint found in the vertebrae of several fossil reptiles of the group Archosauromorpha. It consists of a process on the backside of the vertebrae, the hyposphene, that fits in a depression in the front side of the next vertebrae, the hypantrum. Hyposphene-hypantrum articulations occur in the dorsal vertebrae and sometimes also in the posteriormost cervical and anteriormost caudal vertebrae. In most tetrapods including the human, the vertebrae are connected with each other only via the centrum and the zygapophysis joints. Additional joints like the hyposphene-hypantrum articulations, which add rigidity to the vertebral column, are found in several different reptile lineages; a known example are the zygosphene-zygantrum articulations found in snakes. Hyposphene-hypantrum articulations are found in several unrelated groups within the Archosauromorpha. They occur especially in large forms, for example in rauisuchids and in silesau ...
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Hypantrum
The hyposphene-hypantrum articulation is an accessory joint found in the vertebrae of several fossil reptiles of the group Archosauromorpha. It consists of a process on the backside of the vertebrae, the hyposphene, that fits in a depression in the front side of the next vertebrae, the hypantrum. Hyposphene-hypantrum articulations occur in the dorsal vertebrae and sometimes also in the posteriormost cervical and anteriormost caudal vertebrae. In most tetrapods including the human, the vertebrae are connected with each other only via the centrum and the zygapophysis joints. Additional joints like the hyposphene-hypantrum articulations, which add rigidity to the vertebral column, are found in several different reptile lineages; a known example are the zygosphene-zygantrum articulations found in snakes. Hyposphene-hypantrum articulations are found in several unrelated groups within the Archosauromorpha. They occur especially in large forms, for example in rauisuchids and in silesau ...
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Anatomical Terms Of Location
Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position provides a definition of what is at the front ("anterior"), behind ("posterior") and so on. As part of defining and describing terms, the body is described through the use of anatomical planes and anatomical axes. The meaning of terms that are used can change depending on whether an organism is bipedal or quadrupedal. Additionally, for some animals such as invertebrates, some terms may not have any meaning at all; for example, an animal that is radially symmetrical will have no anterior surface, but can still have a description that a part is close to the middle ("proximal") or further from the middle ("distal"). International organisations have determined vocabularies that are often used as standard vocabularies for subdisciplines of anatom ...
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Sacrum
The sacrum (plural: ''sacra'' or ''sacrums''), in human anatomy, is a large, triangular bone at the base of the spine that forms by the fusing of the sacral vertebrae (S1S5) between ages 18 and 30. The sacrum situates at the upper, back part of the pelvic cavity, between the two wings of the pelvis. It forms joints with four other bones. The two projections at the sides of the sacrum are called the alae (wings), and articulate with the ilium at the L-shaped sacroiliac joints. The upper part of the sacrum connects with the last lumbar vertebra (L5), and its lower part with the coccyx (tailbone) via the sacral and coccygeal cornua. The sacrum has three different surfaces which are shaped to accommodate surrounding pelvic structures. Overall it is concave (curved upon itself). The base of the sacrum, the broadest and uppermost part, is tilted forward as the sacral promontory internally. The central part is curved outward toward the posterior, allowing greater room for the pel ...
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