Lukousaurus
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''Lukousaurus'' is an
archosauromorph Archosauromorpha ( Greek for "ruling lizard forms") is a clade of diapsid reptiles containing all reptiles more closely related to archosaurs (such as crocodilians and dinosaurs, including birds) rather than lepidosaurs (such as tuataras, l ...
based on most of a small skull's snout, displaying distinctive lachrymal horns, found in the
Early Jurassic The Early Jurassic Epoch (geology), Epoch (in chronostratigraphy corresponding to the Lower Jurassic series (stratigraphy), Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic-J ...
-age Lower Lufeng Formation,
Yunnan Yunnan , () is a landlocked province in the southwest of the People's Republic of China. The province spans approximately and has a population of 48.3 million (as of 2018). The capital of the province is Kunming. The province borders the C ...
, China and was described by Chung Chien Young in 1940. The generic name refers to the
Lugou Bridge The Marco Polo Bridge or Lugou Bridge () is a stone bridge located 15 km southwest of Beijing's city center in the Fengtai District. It bridges the Yongding River, a major tributary of Hai River. Situated at the eastern end of the bridge ...
, lit. “crossroads”, near
Beijing } Beijing ( ; ; ), alternatively romanized as Peking ( ), is the capital of the People's Republic of China. It is the center of power and development of the country. Beijing is the world's most populous national capital city, with over 21 ...
, where the Sino-Japanese War started.C.-C. Young. (1940). Preliminary notes on the Lufeng vertebrate fossils. ''Bulletin of the Geological Society of China'' 20(3-4):235-239 ''L. yini'' is tentatively classified as a theropod
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
by some allied to ceratosaurs, by others a coelurosaur. Its skull is rather robust for its size though the teeth were described by the author as typically
theropod Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally c ...
an. Whatever ''Lukousaurus'' was, it was definitely an
archosauromorph Archosauromorpha ( Greek for "ruling lizard forms") is a clade of diapsid reptiles containing all reptiles more closely related to archosaurs (such as crocodilians and dinosaurs, including birds) rather than lepidosaurs (such as tuataras, l ...
.Knoll F., Rohrberg K. (2012). CT scanning, rapid prototyping and re-examination of a partial skull of a basal crocodylomorph from the Late Triassic of Germany. ''Swiss Journal of Geosciences'' 105:109–115.


History

At a locality in the town of Huangchiatien (also called Dahungtien) in
Yunnan Province Yunnan , () is a landlocked province in the southwest of the People's Republic of China. The province spans approximately and has a population of 48.3 million (as of 2018). The capital of the province is Kunming. The province borders the ...
, China, a partial anterior skull and lower jaws as well as possibly a tooth and humerus found nearby. The skull was found in the
lower Jurassic The Early Jurassic Epoch (in chronostratigraphy corresponding to the Lower Jurassic Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic-Jurassic extinction event, 201.3 Ma&nb ...
strata of the Red Beds of the
Lufeng Formation The Lufeng Formation (formerly Lower Lufeng Series) is a Lower Jurassic sedimentary rock formation found in Yunnan, China. It has two units: the lower Dull Purplish Beds/Shawan Member are of Hettangian age, and Dark Red Beds/Zhangjia'ao Member ar ...
, though at the time of its naming in 1940 by Chung Chien Young the beds were thought to date to the Triassic. Young noted that the skull is very strange, with morphologies similar to those of not just Coelurosaurs, which he thought the taxon was, but also Prosauropods and Carnosaurs. The generic name refers to the
Lugou Bridge The Marco Polo Bridge or Lugou Bridge () is a stone bridge located 15 km southwest of Beijing's city center in the Fengtai District. It bridges the Yongding River, a major tributary of Hai River. Situated at the eastern end of the bridge ...
, lit. “crossroads”, near
Beijing } Beijing ( ; ; ), alternatively romanized as Peking ( ), is the capital of the People's Republic of China. It is the center of power and development of the country. Beijing is the world's most populous national capital city, with over 21 ...
, where the Sino-Japanese War started and a symbol of the Chinese resistance against
Japanese imperialism This is a list of regions occupied or annexed by the Empire of Japan until 1945, the year of the end of World War II in Asia, after the surrender of Japan. Control over all territories except most of the Japanese mainland (Hokkaido, Honshu, Kyu ...
. The species name honors the former deputy Director of the Geological Survey of China, T. H. Yin, who kept work on the survey continuing despite the Japanese invasion. The holotype specimen is housed within the
Institute of Vertebrate Paleontology and Paleoanthropology The Institute of Vertebrate Paleontology and Paleoanthropology (IVPP; ) of China is a research institution and collections repository for fossils, including many dinosaur and pterosaur specimens (many from the Yixian Formation). As its name sugges ...
in Beijing, China under specimen number IVPP 23. Simmons, 1965 assigned a distal humerus and co-ossified tibia and fibula to ''Lukousaurus'', though there is no overlap between elements.


Classification

The classification of ''Lukousaurus'' is very uncertain due to the strange characteristics of the type specimen and its referred humerus. ''Lukousaurus'' was first described as a Coelurosaur based on its postorbital and the shaped of the orbit compared to ''
Saltopus ''Saltopus'' ("hopping foot") is a genus of very small bipedal dinosauriform containing the single species ''Saltopus elginensis'' from the late Triassic period of Scotland. It is one of the most famous Elgin Reptiles. Description ''Saltopus ...
,
Podokesaurus ''Podokesaurus'' is a genus of coelophysoid dinosaur that lived in what is now the eastern United States during the Early Jurassic Period. The first fossil was discovered by the geologist Mignon Talbot near Mount Holyoke, Massachusetts, in 1910 ...
,'' and '' Ammosaurus'', though none of these taxa are now considered Coelurosaurs. The size and "some characteristics of the skull" were also noted as being similar to ''
Palaeosaurus ''Palaeosaurus'' (or ''Paleosaurus'') is a genus of indeterminate archosaur known from two teeth found in the Bromsgrove Sandstone Formation and also either the Magnesian Conglomerate or the Avon Fissure Fill of Clifton, Bristol, England (ori ...
'', though ''Palaeosaurus'' is considered dubious. Dong Zhiming in his book on Chinese dinosaurs assigned ''Lukousaurus'' to Podokesauridae in 1992, a similar conclusion to Young's original classification with ''Podokesaurus,'' though in 1997 Kenneth Carpenter placed it as an indeterminate Theropod''.'' In 2008, Mickey Mortimer believed that ''Lukousaurus'' was either an abelisaurid ceratosaur or a sphenosuchian, but revised her position in 2011 after placing it as a basal pseudosuchian using a phylogenetic matrix developed by Sterling Nesbitt. Knoll ''et al''., 2012 found ''Lukousuchus'' to be similar to suchian archosaurs in the anatomy of the antorbital fenestra and Irmis, 2004 stated that ''Lukousuchus'' wasn't a theropod dinosaur or a dinosauromorph in general.Irmis, R. B. (2004). First report of Megapnosaurus (Theropoda: Coelophysoidea) from China. ''PaleoBios'', ''24''(3), 11-18.


References

{{Taxonbar, from=Q591610 Prehistoric theropods Early Jurassic dinosaurs of Asia Taxa named by Yang Zhongjian Fossil taxa described in 1940 Controversial dinosaur taxa