Little Blue Heron
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The little blue heron (''Egretta caerulea'') is a small
heron The herons are long-legged, long-necked, freshwater and coastal birds in the family Ardeidae, with 72 recognised species, some of which are referred to as egrets or bitterns rather than herons. Members of the genera ''Botaurus'' and ''Ixobrychu ...
of the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
'' Egretta''. It is a small, darkly colored heron with a two-toned bill. Juveniles are entirely white, bearing resemblance to the
snowy egret The snowy egret (''Egretta thula'') is a small white heron. The genus name comes from Provençal French for the little egret, , which is a diminutive of , 'heron'. The species name ''thula'' is the Araucano term for the black-necked swan, a ...
. During the breeding season, adults develop different coloration on the head, legs, and feet. They have a range that encompasses much of the
Americas The Americas, which are sometimes collectively called America, are a landmass comprising the totality of North America, North and South America. The Americas make up most of the land in Earth's Western Hemisphere and comprise the New World. ...
, from the United States to northern South America. Some populations are migratory.
Climate change In common usage, climate change describes global warming—the ongoing increase in global average temperature—and its effects on Earth's climate system. Climate change in a broader sense also includes previous long-term changes to ...
will probably cause their distribution to spread north. They can be found in both
saltwater Saline water (more commonly known as salt water) is water that contains a high concentration of dissolved salts (mainly sodium chloride). On the United States Geological Survey (USGS) salinity scale, saline water is saltier than brackish water, ...
and
freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does incl ...
ecosystems. Their preference for either one depends on where they live. Nesting behaviors are documented by numerous sources. The adults build nests in trees, in colonies with other bird species. The number of
eggs Humans and human ancestors have scavenged and eaten animal eggs for millions of years. Humans in Southeast Asia had domesticated chickens and harvested their eggs for food by 1,500 BCE. The most widely consumed eggs are those of fowl, especial ...
laid varies from place to place. The young mature quickly, requiring little attention from adults after about nineteen days of age. Both young and adults are sometimes preyed on by other species. Adults hunt fish,
crabs Crabs are decapod crustaceans of the infraorder Brachyura, which typically have a very short projecting "tail" (abdomen) ( el, βραχύς , translit=brachys = short, / = tail), usually hidden entirely under the thorax. They live in all the ...
, and other small animals. As with clutch sizes, diet can vary regionally. The population of ''E. caerulea'' is declining. Many possible reasons for this have been proposed. Exposure to
heavy metals upright=1.2, Crystals of osmium, a heavy metal nearly twice as dense as lead">lead.html" ;"title="osmium, a heavy metal nearly twice as dense as lead">osmium, a heavy metal nearly twice as dense as lead Heavy metals are generally defined as ...
has been found to have detrimental effects on young birds.


Taxonomy

The little blue heron is part of the
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Ardeidae The herons are long-legged, long-necked, freshwater and coastal birds in the family Ardeidae, with 72 recognised species, some of which are referred to as egrets or bitterns rather than herons. Members of the genera ''Botaurus'' and ''Ixobrychu ...
, a group whose members can be found throughout much of the world, including the
Americas The Americas, which are sometimes collectively called America, are a landmass comprising the totality of North America, North and South America. The Americas make up most of the land in Earth's Western Hemisphere and comprise the New World. ...
,
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
,
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an are ...
, and
Oceania Oceania (, , ) is a geographical region that includes Australasia, Melanesia, Micronesia, and Polynesia. Spanning the Eastern and Western hemispheres, Oceania is estimated to have a land area of and a population of around 44.5 million ...
. It was first described as ''Ardea caerulea'' by
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, ...
in his 10th edition of ''Systema Naturae''. It is now a member of the genus '' Egretta''. It may be closely related to the
snowy egret The snowy egret (''Egretta thula'') is a small white heron. The genus name comes from Provençal French for the little egret, , which is a diminutive of , 'heron'. The species name ''thula'' is the Araucano term for the black-necked swan, a ...
, another member of its genus, which it greatly resembles when young. Variations of the name include ''Ardea coerulea'', ''Florida caerulea'', and ''Hydranassa caerulea''. Young birds found in a little blue heron nest in
North Dakota North Dakota () is a U.S. state in the Upper Midwest, named after the indigenous Dakota Sioux. North Dakota is bordered by the Canadian provinces of Saskatchewan and Manitoba to the north and by the U.S. states of Minnesota to the east, ...
, at a site heavily populated by cattle egrets (''Bubulcus ibis''), which displayed traits of both the former and latter, are believed to be an example of
hybridization Hybridization (or hybridisation) may refer to: *Hybridization (biology), the process of combining different varieties of organisms to create a hybrid *Orbital hybridization, in chemistry, the mixing of atomic orbitals into new hybrid orbitals *Nu ...
between the two species. Other species they are known to hybridize with include the tricolored heron, little egret, snowy egret, and
black-crowned night heron The black-crowned night heron (''Nycticorax nycticorax''), or black-capped night heron, commonly shortened to just night heron in Eurasia, is a medium-sized heron found throughout a large part of the world, including parts of Europe, Asia, and N ...
. Of these four, only the black-crowned night heron is not a member of ''Egretta''.


Description

Males and females have the same coloration. The adults are darkly colored, with purple-
maroon Maroon ( US/ UK , Australia ) is a brownish crimson color that takes its name from the French word ''marron'', or chestnut. "Marron" is also one of the French translations for "brown". According to multiple dictionaries, there are vari ...
heads and blue bodies. During the breeding season, their heads turn dark red. They have two-toned bills, which are a light blue at the base, with black tips. Their eyes are yellow and their legs are greenish. Juveniles are almost completely white, although the upper
primaries Primary elections, or direct primary are a voting process by which voters can indicate their preference for their party's candidate, or a candidate in general, in an upcoming general election, local election, or by-election. Depending on the c ...
are somewhat dark in color. Like adults, their bills are two-toned. Immature birds transitioning from the juvenile to adult phase have a combination of light and dark feathers. Both sexes are about , with a wingspan of . They weigh about . The lores, which are normally a dull green become a shade of
turquoise Turquoise is an opaque, blue-to-green mineral that is a hydrated phosphate of copper and aluminium, with the chemical formula . It is rare and valuable in finer grades and has been prized as a gemstone and ornamental stone for thousands of year ...
. They also develop long plumes on the crest and back, which can stretch past the tail. The legs and feet become black. The eggs are typically smooth, light blue, and unmarked. In size, they are about , with a weight of around .


Distribution and habitat

''Egretta caerulea'' can be found regularly in the United States, Mexico,
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
, northern South America (including Venezuela, Colombia, and Peru), and numerous
Caribbean islands Almost all of the Caribbean islands are in the Caribbean Sea, with only a few in inland lakes. The largest island is Cuba. Other sizable islands include Hispaniola, Jamaica, Puerto Rico and Trinidad and Tobago. Some of the smaller islands a ...
(including Cuba, Jamaica, and
Hispaniola Hispaniola (, also ; es, La Española; Latin and french: Hispaniola; ht, Ispayola; tnq, Ayiti or Quisqueya) is an island in the Caribbean that is part of the Greater Antilles. Hispaniola is the most populous island in the West Indies, and th ...
). They have been recorded as a
vagrant Vagrancy is the condition of homelessness without regular employment or income. Vagrants (also known as bums, vagabonds, rogues, tramps or drifters) usually live in poverty and support themselves by begging, scavenging, petty theft, temporar ...
(a species that appears far outside its natural range) in Greenland, Portugal, and South Africa. Whether or not their range is declining is unknown. In the United States, they can be found from Missouri to Virginia to Florida. They are more common in peninsular Florida than the
Florida Panhandle The Florida Panhandle (also West Florida and Northwest Florida) is the northwestern part of the U.S. state of Florida; it is a salient roughly long and wide, lying between Alabama on the north and the west, Georgia on the north, and the ...
. They can occasionally roam as far north as Canada. Individuals in central
Alabama (We dare defend our rights) , anthem = " Alabama" , image_map = Alabama in United States.svg , seat = Montgomery , LargestCity = Huntsville , LargestCounty = Baldwin County , LargestMetro = Greater Birmingham , area_total_km2 = 135,7 ...
tend to migrate towards South America and the Caribbean, while those from the
Mississippi River The Mississippi River is the List of longest rivers of the United States (by main stem), second-longest river and chief river of the second-largest Drainage system (geomorphology), drainage system in North America, second only to the Hudson B ...
west travel to Mexico and Central America. One study found that of seven migratory wading bird species, the little blue heron had the greatest mean dispersal distance, of . Future
climate change In common usage, climate change describes global warming—the ongoing increase in global average temperature—and its effects on Earth's climate system. Climate change in a broader sense also includes previous long-term changes to ...
is projected to increase its overall range. If global warming continues at its current rate, by the year 2080, its summer range will have increased by 87%. Of its current range, it is expected to lose only 1%. These gains would spread its summer distribution well into more northern parts of the US, such as Michigan and Minnesota, and even into southern Canada. The little blue heron can be found in
freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does incl ...
and
marine Marine is an adjective meaning of or pertaining to the sea or ocean. Marine or marines may refer to: Ocean * Maritime (disambiguation) * Marine art * Marine biology * Marine debris * Marine habitats * Marine life * Marine pollution Military ...
environments. These include
mangrove forest Mangrove forests, also called mangrove swamps, mangrove thickets or mangals, are productive wetlands that occur in coastal intertidal zones. Mangrove forests grow mainly at tropical and subtropical latitudes because mangroves cannot withstand fre ...
s,
bogs A bog or bogland is a wetland that accumulates peat as a deposit of dead plant materials often mosses, typically sphagnum moss. It is one of the four main types of wetlands. Other names for bogs include mire, mosses, quagmire, and muskeg; a ...
,
swamps A swamp is a forested wetland.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p. Swamps are considered to be transition zones because both land and water play a role in ...
,
salt marshes A salt marsh or saltmarsh, also known as a coastal salt marsh or a tidal marsh, is a coastal ecosystem in the upper coastal intertidal zone between land and open saltwater or brackish water that is regularly flooded by the tides. It is domina ...
,
tidal flats Mudflats or mud flats, also known as tidal flats or, in Ireland, slob or slobs, are coastal wetlands that form in intertidal areas where sediments have been deposited by tides or rivers. A global analysis published in 2019 suggested that tidal f ...
,
estuaries An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environme ...
,
streams A stream is a continuous body of surface water flowing within the bed and banks of a channel. Depending on its location or certain characteristics, a stream may be referred to by a variety of local or regional names. Long large streams a ...
, and flooded fields. They are usually found at low elevations, but can be seen at heights of in the
Andes The Andes, Andes Mountains or Andean Mountains (; ) are the longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range is long, wide (widest between 18°S – 20°S ...
. In
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and th ...
, they tend to favor freshwater habitats, while in the Caribbean, they are more often found in saltwater. Towards the southern extent of their range, in
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
, they are found almost exclusively along the coast, rarely venturing inland at all.


Regional variations

Juveniles in San Blas, in the
Mexican state The states of Mexico are first-level administrative territorial entities of the country of Mexico, which is officially named United Mexican States. There are 32 federal entities in Mexico (31 states and the capital, Mexico City, as a separate en ...
of
Nayarit Nayarit (), officially the Free and Sovereign State of Nayarit ( es, Estado Libre y Soberano de Nayarit), is one of the 31 states that, along with Mexico City, comprise the Federal Entities of Mexico. It is divided in 20 municipalities and its ...
have an atypical color-scheme. In these birds, the top of the head is
chestnut The chestnuts are the deciduous trees and shrubs in the genus ''Castanea'', in the beech family Fagaceae. They are native to temperate regions of the Northern Hemisphere. The name also refers to the edible nuts they produce. The unrel ...
colored and the wings tips are much darker. Initially, it was suggested that they may be hybrids, however further study concluded they were most likely a natural variation. No other geographic varieties have been observed.


Behavior and ecology

Little blue herons are prefer to stay still and wait when hunting, rather than chase after prey. They walk slowly and search for fish and other prey items, flying to different spots if needed. They tend to move slower than other related species, which can help distinguish them. They are not usually found in large numbers at any body of water. Occasionally, however, they will gather with other
herons The herons are long-legged, long-necked, freshwater and coastal birds in the family Ardeidae, with 72 recognised species, some of which are referred to as egrets or bitterns rather than herons. Members of the genera ''Botaurus'' and ''Ixobrychus ...
, especially if they have found a school of fish trapped in shallow water. They sometimes also feeds in grassy fields.


Reproduction and life cycle

During
courtship Courtship is the period wherein some couples get to know each other prior to a possible marriage. Courtship traditionally may begin after a betrothal and may conclude with the celebration of marriage. A courtship may be an informal and private m ...
, both males and females practice bill-nibling. Males also use a neck-stretch to attract mates.


Nesting

Little blue herons typically nest in trees alongside other roosting birds. They are colonial nesters (nesting in groups). Examples of species they may nest alongside include the
scarlet ibis The scarlet ibis (''Eudocimus ruber'') is a species of ibis in the bird family Threskiornithidae. It inhabits tropical South America and part of the Caribbean. In form, it resembles most of the other twenty-seven extant species of ibis, but ...
, yellow-crowned night heron,
great egret The great egret (''Ardea alba''), also known as the common egret, large egret, or (in the Old World) great white egret or great white heron is a large, widely distributed egret. The four subspecies are found in Asia, Africa, the Americas, an ...
, black-crowned night heron, and snowy egret. During nest construction, males bring twigs to females, who use them to build the nest. Both males and females help incubate their clutch. They begin incubation after two eggs have been laid, which will cause any later eggs to hatch out of sync. The chicks that hatch later tend to not receive as much food as early-hatching ones, which limits their growth. Clutch sizes vary significantly throughout their range. In
Trinidad Trinidad is the larger and more populous of the two major islands of Trinidad and Tobago. The island lies off the northeastern coast of Venezuela and sits on the continental shelf of South America. It is often referred to as the southernmos ...
, there are usually 2-5 eggs, while in Costa Rica, only 2-4 are laid on average. In North America, the mean is 2.67-4.4. The very lowest values are seen in southeastern Brazil and the US states Florida and Georgia, where no more than three are generally laid. Young herons are able to start climbing around the branches by their nests at 15 days old. Due to the young age at which they develop
motor skills A motor skill is a function that involves specific movements of the body's muscles to perform a certain task. These tasks could include walking, running, or riding a bike. In order to perform this skill, the body's nervous system, muscles, and br ...
in their legs, the young do not rely on their parents for anything besides feeding after 19 days, at which point the adults begin foraging away from the nest. By 20–25 days, they can climb to the very top of the tree their nest is built on, or even into other trees. They can fly short distances at around 30 days of age (some take 35–38), but will still be dependent on adults for about two weeks after that. It is in their second year of life that juveniles begin to lose their white feathers.


Predation

There is circumstantial evidence that young black-crowned night herons and crab-eating raccoons prey on nestling little blue herons. Adults have been observed driving a yellow-headed caracara away from their nests. In the presence of a Harris' hawk, however, the little blue herons fled.


Parasites

Twenty-four different species of parasitic worms were found on 33 of 35 little blue herons examined in
South Florida South Florida is the southernmost region of the U.S. state of Florida. It is one of Florida's three most commonly referred to directional regions; the other two are Central Florida and North Florida. South Florida is the southernmost part of ...
. These included
trematodes Trematoda is a class of flatworms known as flukes. They are obligate internal parasites with a complex life cycle requiring at least two hosts. The intermediate host, in which asexual reproduction occurs, is usually a snail. The definitive host ...
,
nematodes The nematodes ( or grc-gre, Νηματώδη; la, Nematoda) or roundworms constitute the phylum Nematoda (also called Nemathelminthes), with plant-parasitic nematodes also known as eelworms. They are a diverse animal phylum inhabiting a broa ...
, acanthocephalans, and one
cestode Cestoda is a class of parasitic worms in the flatworm phylum (Platyhelminthes). Most of the species—and the best-known—are those in the subclass Eucestoda; they are ribbon-like worms as adults, known as tapeworms. Their bodies consist of ...
. The most common trematode was '' Posthodiplostomum macrocotyle'', and the most common nematodes were '' Contracaecum multipapillatum'' and '' Contracaecum microcephalum''. The acanthocephalan and cestode species could not be identified (in the latter, neither could the genus).


Prey

On the eastern coast of North America, little blue herons primarily feed on fish, however their diet varies significantly throughout their range. In a study of individuals in mangrove forests in southeastern Brazil, 80% of their diet during the breeding season was found to consist of
crabs Crabs are decapod crustaceans of the infraorder Brachyura, which typically have a very short projecting "tail" (abdomen) ( el, βραχύς , translit=brachys = short, / = tail), usually hidden entirely under the thorax. They live in all the ...
. Compared to the scarlet ibis, the herons preferred
arboreal Arboreal locomotion is the locomotion of animals in trees. In habitats in which trees are present, animals have evolved to move in them. Some animals may scale trees only occasionally, but others are exclusively arboreal. The habitats pose nu ...
or semi-arboreal species, such as '' Aratus pisonii'' and '' Metasesarma rubripes'', while the former preferred to take burrowing species. This demonstrates their different feeding strategies—scarlet ibises being foragers who hunt using their
sense of touch In physiology, the somatosensory system is the network of neural structures in the brain and body that produce the perception of touch ( haptic perception), as well as temperature ( thermoception), body position (proprioception), and pain. It ...
and little blue herons being visual hunters. In another mangrove forest in southwestern
Puerto Rico Puerto Rico (; abbreviated PR; tnq, Boriken, ''Borinquen''), officially the Commonwealth of Puerto Rico ( es, link=yes, Estado Libre Asociado de Puerto Rico, lit=Free Associated State of Puerto Rico), is a Caribbean island and unincorporated ...
, the entire diet was found to consist of
fiddler crabs The fiddler crab or calling crab may be any of more than one hundred species of semiterrestrial marine crabs in the family Ocypodidae, well known for their sexually dimorphic claws; the males' major claw is much larger than the minor claw, while ...
.


Conservation

The little blue heron is listed as a
least-concern species A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. T ...
by the
International Union for the Conservation of Nature The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
, although its numbers are decreasing. Historically, they were not hunted for their feathers as much as other heron species due to their lack of visually attractive plumes. The dangers faced by ''Egretta caerulea'' are not well researched. They could include development along coastlines, habitat disturbance, predators,
pesticide Pesticides are substances that are meant to control pests. This includes herbicide, insecticide, nematicide, molluscicide, piscicide, avicide, rodenticide, bactericide, insect repellent, animal repellent, microbicide, fungicide, and ...
exposure, and
parasites Parasitism is a close relationship between species, where one organism, the parasite, lives on or inside another organism, the host, causing it some harm, and is adapted structurally to this way of life. The entomologist E. O. Wilson ha ...
. The metals
cadmium Cadmium is a chemical element with the Symbol (chemistry), symbol Cd and atomic number 48. This soft, silvery-white metal is chemically similar to the two other stable metals in group 12 element, group 12, zinc and mercury (element), mercury. Li ...
and
lead Lead is a chemical element with the symbol Pb (from the Latin ) and atomic number 82. It is a heavy metal that is denser than most common materials. Lead is soft and malleable, and also has a relatively low melting point. When freshly cut, ...
have been found to lead to slower growth rates and higher death rates, respectively, of young birds. In
Sepetiba Bay Sepetiba is a neighborhood in the West Zone of Rio de Janeiro, Brazil, surrounded by Santa Cruz and Guaratiba, and by the Sepetiba Bay. It occupies an area of 1,162.13 ha, and has a population of 35,892 (according to Instituto Brasileiro de G ...
,
Rio de Janeiro Rio de Janeiro ( , , ; literally 'River of January'), or simply Rio, is the capital of the state of the same name, Brazil's third-most populous state, and the second-most populous city in Brazil, after São Paulo. Listed by the GaWC as a b ...
,
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
, little blue herons were found to have relatively high levels of metal contamination in the
liver The liver is a major organ only found in vertebrates which performs many essential biological functions such as detoxification of the organism, and the synthesis of proteins and biochemicals necessary for digestion and growth. In humans, it i ...
and
kidneys The kidneys are two reddish-brown bean-shaped organs found in vertebrates. They are located on the left and right in the retroperitoneal space, and in adult humans are about in length. They receive blood from the paired renal arteries; bloo ...
. In areas with cattle egrets, little blue herons have been found to nest for shorter amounts of time, and produce fewer young that survive to adulthood. Cattle egrets only begin pairing when most little blue herons already have eggs or live young in their nests. The former species has been observed stealing twigs from nests of the latter. This behavior sometimes leads to the young falling out of the nest or the cattle egrets removing them.


References


Notes


External links


Field guide on Flickr

Florida Bird Sounds
- Florida Museum of Natural History * * {{Taxonbar, from=Q371028 little blue heron Little blue heron Native birds of the Southeastern United States Birds of the Dominican Republic Birds of the Americas Birds described in 1758 Taxa named by Carl Linnaeus Articles containing video clips