Kryptops
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''Kryptops'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
abelisaurid Abelisauridae (meaning "Abel's lizards") is a family (or clade) of ceratosaurian theropod dinosaurs. Abelisaurids thrived during the Cretaceous period, on the ancient southern supercontinent of Gondwana, and today their fossil remains are foun ...
theropod Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally c ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
from the
Early Cretaceous The Early Cretaceous ( geochronological name) or the Lower Cretaceous (chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145  Ma to 100.5 Ma. Geology Pro ...
of Niger. It is known from a partial skeleton found at the
Gadoufaoua The Elrhaz Formation is a geological formation in Niger, central Africa. Its strata date back to the Early Cretaceous, about 125 to 112 million years ago. Dinosaur remains are among the fossils that have been recovered from the formation, alongsi ...
locality in the western
Ténéré Desert The Ténéré (Tuareg: Tenere, literally: "desert") is a desert region in the south central Sahara. It comprises a vast plain of sand stretching from northeastern Niger into western Chad, occupying an area of over . The Ténéré's boundaries a ...
, in rocks of the
Aptian The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous Epoch or Series and encompasses the time from 121.4 ± 1.0 Ma to 113.0 ± 1.0 Ma (million years ago), a ...
-
Albian The Albian is both an age of the geologic timescale and a stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early/Lower Cretaceous Epoch/Series. Its approximate time range is 113.0 ± 1.0 Ma to 100.5 ± 0.9 M ...
age
Elrhaz Formation The Elrhaz Formation is a geological formation in Niger, central Africa. Its strata date back to the Early Cretaceous, about 125 to 112 million years ago. Dinosaur remains are among the fossils that have been recovered from the formation, alongsi ...
. This dinosaur was described by
paleontologists Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of fossi ...
Paul Sereno Paul Callistus Sereno (born October 11, 1957) is a professor of paleontology at the University of Chicago and a National Geographic "explorer-in-residence" who has discovered several new dinosaur species on several continents, including at sites ...
and Stephen Brusatte in 2008. The genus name means "covered face", in reference to evidence that the face bore a tightly adhering covering. The
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
is ''K. palaios,'' which means "old".


Discovery

The
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
skeleton, MNN GAD1, includes a
maxilla The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The t ...
(main tooth-bearing bone of the upper jaw),
vertebra The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic ...
e, ribs, and articulated
pelvic girdle The pelvis (plural pelves or pelvises) is the lower part of the trunk, between the abdomen and the thighs (sometimes also called pelvic region), together with its embedded skeleton (sometimes also called bony pelvis, or pelvic skeleton). The p ...
and
sacrum The sacrum (plural: ''sacra'' or ''sacrums''), in human anatomy, is a large, triangular bone at the base of the spine that forms by the fusing of the sacral vertebrae (S1S5) between ages 18 and 30. The sacrum situates at the upper, back part ...
, belonging to an adult about . According to the describers, this specimen represents one of the earliest known abelisaurids, and is notable for the heavily textured surface of the maxilla; the presence of pits and impressions of blood vessels indicates that there was a covering firmly attached to the face, perhaps of
keratin Keratin () is one of a family of structural fibrous proteins also known as ''scleroproteins''. Alpha-keratin (α-keratin) is a type of keratin found in vertebrates. It is the key structural material making up scales, hair, nails, feathers, ho ...
. Sereno and Brusatte performed a
cladistic Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups (" clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived char ...
analysis that found ''Kryptops'' to be the most basal
abelisaurid Abelisauridae (meaning "Abel's lizards") is a family (or clade) of ceratosaurian theropod dinosaurs. Abelisaurids thrived during the Cretaceous period, on the ancient southern supercontinent of Gondwana, and today their fossil remains are foun ...
. This was based on several features, including a maxilla textured externally by impressed vascular grooves and a narrow antorbital fossa, that clearly place ''Kryptops palaios'' within Abelisauridae as its oldest known member. In 2012 Matthew Carrano and colleagues, on the other hand, considered ''Kryptops palaios'' to be a
chimera Chimera, Chimaera, or Chimaira (Greek for " she-goat") originally referred to: * Chimera (mythology), a fire-breathing monster of Ancient Lycia said to combine parts from multiple animals * Mount Chimaera, a fire-spewing region of Lycia or Cilicia ...
, and state that its postcranial remains (especially a pelvis and sacrum), found some 15 metres from the holotypic maxilla, actually belong to a
carcharodontosaurid Carcharodontosauridae (carcharodontosaurids; from the Greek καρχαροδοντόσαυρος, ''carcharodontósauros'': "shark-toothed lizards") is a group of carnivorous theropod dinosaurs. In 1931, Ernst Stromer named Carcharodontosauridae ...
(possibly to '' Eocarcharia dinops'').


See also

*
Timeline of ceratosaur research This timeline of ceratosaur research is a chronological listing of events in the history of paleontology focused on the ceratosaurs, a group of relatively primitive, often horned, predatory theropod dinosaurs that became the apex predators of ...


References

Abelisaurids Early Cretaceous dinosaurs of Africa Albian life Aptian life Cretaceous Niger Fossils of Niger Fossil taxa described in 2008 Taxa named by Paul Sereno Taxa named by Stephen L. Brusatte {{theropod-stub