Green kingfisher
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The green kingfisher (''Chloroceryle americana'') is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
of "water kingfisher" in
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus Genus ( plural genera ) is a taxonomic rank used in the biological classifica ...
Cerylinae of family Alcedinidae. It is found from southern
Texas Texas (, ; Spanish: ''Texas'', ''Tejas'') is a state in the South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2020, it is the second-largest U.S. state by ...
in the
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country Continental United States, primarily located in North America. It consists of 50 U.S. state, states, a Washington, D.C., ...
south through
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
, in every mainland
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the sou ...
n country except
Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the eas ...
, and on
Trinidad and Tobago Trinidad and Tobago (, ), officially the Republic of Trinidad and Tobago, is the southernmost island country in the Caribbean. Consisting of the main islands Trinidad and Tobago, and numerous much smaller islands, it is situated south of ...
.HBW and BirdLife International (2022) Handbook of the Birds of the World and BirdLife International digital checklist of the birds of the world. Version 7. Available at: http://datazone.birdlife.org/userfiles/file/Species/Taxonomy/HBW-BirdLife_Checklist_v7_Dec22.zip retrieved 13 December 2022Remsen, J. V., Jr., J. I. Areta, E. Bonaccorso, S. Claramunt, A. Jaramillo, D. F. Lane, J. F. Pacheco, M. B. Robbins, F. G. Stiles, and K. J. Zimmer. Version 24 July 2022. Species Lists of Birds for South American Countries and Territories. https://www.museum.lsu.edu/~Remsen/SACCCountryLists.htm retrieved 24 July 2022


Taxonomy and systematics

The green kingfisher was formally described in 1788 by the German naturalist
Johann Friedrich Gmelin , fields = , workplaces = University of GöttingenUniversity of Tübingen , alma_mater = University of Tübingen , doctoral_advisor = Philipp Friedrich Gmelin Ferdinand Christoph Oetinger , academic_advisors = , docto ...
in his revised and expanded edition of
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, ...
's ''
Systema Naturae ' (originally in Latin written ' with the ligature æ) is one of the major works of the Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the system, now known as binomial ...
''. He placed it with the other kingfishers in the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
''
Alcedo ''Alcedo'' is a genus of birds in the kingfisher subfamily Alcedininae. The genus was introduced by Carl Linnaeus in 1758 in the 10th edition of his ''Systema Naturae''. The type species is the common kingfisher (''Alcedo ispida'', now ''Alcedo ...
'' and coined the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bo ...
''Alcedo americana''. Gmelin based his description on "Le martin-pêcheur vert et blanc" from
Cayenne Cayenne (; ; gcr, Kayenn) is the capital city of French Guiana, an overseas region and department of France located in South America. The city stands on a former island at the mouth of the Cayenne River on the Atlantic coast. The city's m ...
that had been described and illustrated in 1780 by the French polymath
Comte de Buffon Georges-Louis Leclerc, Comte de Buffon (; 7 September 1707 – 16 April 1788) was a French naturalist, mathematician, cosmologist, and encyclopédiste. His works influenced the next two generations of naturalists, including two prominent F ...
and also the "white and green kingfisher" that had been described in 1782 by the English ornithologist John Latham. The green kingfisher is now placed in the genus ''
Chloroceryle The American green kingfishers are the kingfisher genus ''Chloroceryle'', which are native to tropical Central and South America, with one species extending north to south Texas. Species There are four species: The American green kingfishers b ...
'' that was introduced in 1848 by
Johann Jakob Kaup Johann Jakob von Kaup (10 April 1803 – 4 July 1873) was a German naturalist. A proponent of natural philosophy, he believed in an innate mathematical order in nature and he attempted biological classifications based on the Quinarian system. Kau ...
. A
molecular phylogenetic Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...
study published in 2006 found that the green kingfisher is a
sister species In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...
to the larger
green-and-rufous kingfisher The green-and-rufous kingfisher (''Chloroceryle inda'') is a species of "water kingfisher" in subfamily Cerylinae of family Alcedinidae. It is found in the American tropics from Nicaragua to Panama and in every mainland South American country ...
(''C. inda''). Five
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. Not all specie ...
are recognized: * ''C. a. hachisukai'' ( Laubmann, 1941) * ''C. a. septentrionalis'' ( Sharpe, 1892) * ''C. a. americana'' (Gmelin, JF, 1788) * ''C. a. mathewsii'' Laubmann, 1927 * ''C. a. cabanisii'' ( Tschudi, 1846) Some populations of ''C. a. septentrionalis'' have in the past been separated as subspecies ''C. a. vanrossemi'' and ''C. a. isthmica''. The population on Trinidad and Tobago, usually included with ''C. a. americana'', has in the past been treated as subspecies ''C. a. croteta''. Other populations of ''C. a. americana'' have been suggested to be split as ''C. a. hellmayri'' and ''C. a. bottomeana''. None of these subspecies are currently (2022) recognized by major worldwide taxonomic systems.Moskoff, W. (2020). Green Kingfisher (''Chloroceryle americana''), version 1.0. In Birds of the World (A. F. Poole and F. B. Gill, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.grnkin.01 retrieved December 13, 2022Clements, J. F., T. S. Schulenberg, M. J. Iliff, T. A. Fredericks, J. A. Gerbracht, D. Lepage, S. M. Billerman, B. L. Sullivan, and C. L. Wood. 2022. The eBird/Clements checklist of birds of the world: v2022. Downloaded from https://www.birds.cornell.edu/clementschecklist/download/ retrieved November 10, 2022


Description

The green kingfisher is about long and weighs about ; females are larger and heavier than males. Birds in the northern and southern parts of its range, and those west of the Andes, are larger and heavier than the others, but the differences tend to be clinal. The species has the typical kingfisher shape with long heavy bill (heaviest in Trinidad and Tobago); in contrast to many other kingfishers, however, it does not have an obvious crest. The bill is black with some horn color at the base of the
mandible In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower teeth in place. The mandible sits beneath the maxilla. It is the only movable bone ...
and its legs and feet are dark gray. Both sexes have green upperparts with two or more rows of white spots on the flight feathers. Their underparts are mostly white with green spots on the sides and flanks. Their tail is green with much white on the outer feathers that shows best in flight. Adult males have a white collar and a rufous breast. Adult females have a white collar, a buffy throat and breast, and a band of green speckles across the breast and upper belly. Juveniles resemble females but are duller and have small buff spots on their crown and wing
coverts A covert feather or tectrix on a bird is one of a set of feathers, called coverts (or ''tectrices''), which, as the name implies, cover other feathers. The coverts help to smooth airflow over the wings and tail. Ear coverts The ear coverts are ...
. One vocalization "resembles striking of two pebbles together, usually single or double". Another is "a harsh, buzzy scold, described variously as ''tsheersh'', ''tseelp'' or ''zchrrk''". Authors differ on their interpretations of whether the vocalizations are
calls Call or Calls may refer to: Arts, entertainment, and media Games * Call, a type of betting in poker * Call, in the game of contract bridge, a bid, pass, double, or redouble in the bidding stage Music and dance * Call (band), from Lahore, Pak ...
or
songs A song is a musical composition intended to be performed by the human voice. This is often done at distinct and fixed pitches (melodies) using patterns of sound and silence. Songs contain various forms, such as those including the repetition ...
, and if they are calls what their purpose is. The subspecies differ somewhat in the shade of the upperparts' green, the amount of white on the wings and tail, the amount and size of the flank markings, and the extent of the breast bands (complete or not). However, there is much individual variation within each subspecies and the differences are to some extent clinal, so the differences among the subspecies tend to be obscured.


Distribution and habitat

The subspecies of green kingfisher are distributed thus: * ''C. a. hachisukai'', from extreme southern Arizona, southwestern New Mexico, and west central Texas in the USA into northwestern Mexico to
Nayarit Nayarit (), officially the Free and Sovereign State of Nayarit ( es, Estado Libre y Soberano de Nayarit), is one of the 31 states that, along with Mexico City, comprise the Federal Entities of Mexico. It is divided in 20 municipalities and its ...
* ''C. a. septentrionalis'', from south central Texas and eastern Mexico south to northern Colombia and western Venezuela * ''C. a. americana'', South America east of the Andes from Venezuela east through
the Guianas The Guianas, sometimes called by the Spanish loan-word ''Guayanas'' (''Las Guayanas''), is a region in north-eastern South America which includes the following three territories: * French Guiana, an overseas department and region of France ...
into northern and central Brazil and south to northeastern Bolivia; also Trinidad and Tobago * ''C. a. mathewsii'', from southern Brazil and southern Bolivia through Paraguay and Uruguay to central Argentina * ''C. a. cabanisii'', west of the Andes from Colombia through Ecuador to southernmost Peru; as a
vagrant Vagrancy is the condition of homelessness without regular employment or income. Vagrants (also known as bums, vagabonds, rogues, tramps or drifters) usually live in poverty and support themselves by begging, scavenging, petty theft, temporar ...
to Chile The green kingfisher inhabits wooded shorelines of streams and freshwater ponds and lakes. It favors still or slow-moving water, and though it requires low vegetation for hunting perches it generally prefers relatively open habitat rather than dense forest. It is a year-round resident throughout its range but roams a territory that may be as much as or more of river.


Behavior


Feeding

The Amazon kingfisher usually hunts from a perch from which it dives into water for its prey. The perch is typically about high; it may be directly over water or within a few meters of its edge. In a study in Amazonia about half of the perches were bare snags and the rest were a mix of leafless and leafy trees and bushes. It usually shifts perches after each foray. Occasionally it hovers before diving, sometimes from as high as . Pairs often defend feeding territories from other green kingfishers but seldom from other kingfisher species. The prey is mostly small fish but includes crustaceans such as shrimp and also adult and nymph aquatic and terrestrial insects. The size of the fish taken varies, apparently with availabilty and the presence of other species of kingfishers. Studies have published sizes as , as averages of and at different sites, and as "seldom exceed nga length of two inches".


Breeding

The green kingfisher's breeding season varies geographically. In Central America it is during the dry season of spring and early summer. In Suriname and Guyana, it usually extends to August and occasionally to December or beyond. Breeding dates further south have not been documented. Both members of a pair excavate a nest burrow, almost always in an earthen bank of a stream or river. It is up to long with a small nest chamber at the end. The clutch size varies from two to six. The incubation period is 19 to 21 days; usually the female incubates at night and the parents alternate during the day. The young fledge 26 to 27 days after hatching and the parents chase juveniles from their territory about 29 days after fledging.


Status

The
IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
has assessed the green kingfisher as being of Least Concern. It has an extremely large range. Its estimated population of about 20 million mature individuals is, however, believed to be decreasing. No immediate threats have been identified.


References


External links


Stamps
(for
El Salvador El Salvador (; , meaning " The Saviour"), officially the Republic of El Salvador ( es, República de El Salvador), is a country in Central America. It is bordered on the northeast by Honduras, on the northwest by Guatemala, and on the south ...
, Mexico,
Nicaragua Nicaragua (; ), officially the Republic of Nicaragua (), is the largest country in Central America, bordered by Honduras to the north, the Caribbean to the east, Costa Rica to the south, and the Pacific Ocean to the west. Managua is the coun ...
,
Paraguay Paraguay (; ), officially the Republic of Paraguay ( es, República del Paraguay, links=no; gn, Tavakuairetã Paraguái, links=si), is a landlocked country in South America. It is bordered by Argentina to the south and southwest, Brazil to t ...
,
Suriname Suriname (; srn, Sranankondre or ), officially the Republic of Suriname ( nl, Republiek Suriname , srn, Ripolik fu Sranan), is a country on the northeastern Atlantic coast of South America. It is bordered by the Atlantic Ocean to the nor ...
) with Range Map at bird-stamps.org * {{Taxonbar, from=Q580159 green kingfisher green kingfisher Native birds of the Southwestern United States Kingfisher, green Birds of the Rio Grande valleys Birds of Central America Birds of South America green kingfisher green kingfisher