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Wildebeest ( , , ), also called gnu ( or ), are
antelope The term antelope is used to refer to many species of even-toed ruminant that are indigenous to various regions in Africa and Eurasia. Antelope comprise a wastebasket taxon defined as any of numerous Old World grazing and browsing hoofed mammals ...
s of the genus ''Connochaetes'' and native to Eastern and Southern Africa. They belong to the family Bovidae, which includes true antelopes, cattle, goats, sheep, and other even-toed horned ungulates. There are two species of wildebeest: the
black wildebeest The black wildebeest or white-tailed gnu (''Connochaetes gnou'') is one of the two closely related wildebeest species. It is a member of the genus '' Connochaetes'' and family Bovidae. It was first described in 1780 by Eberhard August Wilhelm ...
or white-tailed gnu (''C. gnou''), and the blue wildebeest or brindled gnu (''C. taurinus''). Fossil records suggest these two species diverged about one million years ago, resulting in a northern and a southern species. The blue wildebeest remained in its original range and changed very little from the ancestral species, while the black wildebeest changed more as adaptation to its open grassland habitat in the south. The most obvious ways of telling the two species apart are the differences in their colouring and in the way their horns are oriented. In
East Africa East Africa, Eastern Africa, or East of Africa, is the eastern subregion of the African continent. In the United Nations Statistics Division scheme of geographic regions, 10-11-(16*) territories make up Eastern Africa: Due to the historical ...
, the blue wildebeest is the most abundant big-game species; some populations perform an annual migration to new grazing grounds, but the black wildebeest is merely nomadic. Breeding in both takes place over a short period of time at the end of the rainy season and the calves are soon active and are able to move with the herd, a fact necessary for their survival. Nevertheless, some fall prey to large carnivores, especially the
spotted hyena The spotted hyena (''Crocuta crocuta''), also known as the laughing hyena, is a hyena species, currently classed as the sole extant member of the genus ''Crocuta'', native to sub-Saharan Africa. It is listed as being of least concern by the IUC ...
. Wildebeest often graze in mixed herds with zebra, which gives heightened awareness of potential
predators Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill the ...
. They are also alert to the warning signals emitted by other animals such as baboons. Wildebeest are a tourist attraction but compete with domesticated livestock for pasture and are sometimes blamed by farmers for transferring diseases and parasites to their cattle. Illegal hunting does take place but the population trend is fairly stable. Wildebeest can also be found in national parks or on private land. The International Union for Conservation of Nature lists both kinds of wildebeest as least-concern species.


Etymology

is Dutch for "wild beast" or "wild cattle" in Afrikaans (''
bees Bees are winged insects closely related to wasps and ants, known for their roles in pollination and, in the case of the best-known bee species, the western honey bee, for producing honey. Bees are a monophyletic lineage within the superfamil ...
'' " cattle"), while ''Connochaetes'' derives from the Ancient Greek words , , " beard", and χαίτη, , "flowing hair", " mane". Some sources claim the name "gnu" originates from the Khoekhoe name for these animals, ''t'gnu''. Others contend the name and its pronunciation in English go back to the word ''!nu:''used for the
black wildebeest The black wildebeest or white-tailed gnu (''Connochaetes gnou'') is one of the two closely related wildebeest species. It is a member of the genus '' Connochaetes'' and family Bovidae. It was first described in 1780 by Eberhard August Wilhelm ...
by the San people.


Classification


Taxonomy and evolution

The wildebeest, genus ''Connochaetes'', is placed under the family Bovidae and subfamily
Alcelaphinae The subfamily Alcelaphinae or tribe Alcelaphini of the family Bovidae contains wildebeest, hartebeest, bonteboks, and several similar species. Depending on the classification, there are 6–10 species placed in four genera, although ''Beatragu ...
, where its closest relatives are the hartebeest ('' Alcelaphus'' spp.), the
hirola The hirola (''Beatragus hunteri''), also called the Hunter's hartebeest or Hunter's antelope, is a critically endangered antelope species found on the border between Kenya and Somalia. It was first described by the big game hunter and zoologist ...
(''Beatragus hunteri''), and species in the genus '' Damaliscus'', such as the
topi ''Damaliscus lunatus jimela'' is a subspecies of topi, and is usually just called a topi. It is a highly social and fast type of antelope found in the savannas, semi-deserts, and floodplains of sub-Saharan Africa. Names The word ''tope'' or '' ...
, the
tsessebe The common tsessebe or sassaby (''Damaliscus lunatus lunatus'') is the southern, nominate subspecies of ''Damaliscus lunatus'', although some authorities have recognised it as an independent species. It is most closely related to the Bangweulu ts ...
, the
blesbok The blesbok or blesbuck (''Damaliscus pygargus phillipsi'') is a subspecies of the bontebok antelope endemic to South Africa, Eswatini and Namibia. It has a distinctive white face and forehead which inspired the name, because ''bles'' is the Afri ...
and the
bontebok The bontebok (''Damaliscus pygargus'') is an antelope found in South Africa, Lesotho and Namibia. ''D. pygargus'' has two subspecies; the nominate subspecies (''D. p. pygargus''), occurring naturally in the Fynbos and Renosterveld areas of the W ...
. The name ''Connochaetes'' was given by German zoologist Hinrich Lichtenstein in 1812. Dutch settlers first discovered wildebeest in about 1700, on their way to the interior of South Africa. Due to their resemblance to wild cattle, these people called them "wild ox" or "wildebeest". The blue wildebeest was first known to westerners in the northern part of South Africa a century later, in the 1800s. In the early 20th century, one species of the wildebeest, ''C. albojubatus'', was identified in eastern Africa. In 1914, two separate races of the wildebeest were introduced, namely ''Gorgon a. albojubatus'' ("Athi white-bearded wildebeest") and ''G. a. mearnsi'' ("Loita white-bearded wildebeest"). However, in 1939, the two were once again merged into a single race, ''Connochaetes taurinus albojubatus''. In the mid-20th century, two separate forms were recognised, ''Gorgon taurinus hecki'' and ''G. t. albojubatus''. Finally, two distinct types of wildebeest – the blue and black wildebeest – were identified. The blue wildebeest was at first placed under a separate genus, ''Gorgon'', while the black wildebeest belonged to the genus ''Connochaetes''. Today, they are united in the single genus ''Connochaetes'', with the black wildebeest being named (''C. gnou'') and the blue wildebeest (''C. taurinus''). According to a
mitochondrial DNA Mitochondrial DNA (mtDNA or mDNA) is the DNA located in mitochondria, cellular organelles within eukaryotic cells that convert chemical energy from food into a form that cells can use, such as adenosine triphosphate (ATP). Mitochondrial D ...
analysis, the black wildebeest are estimated to have diverged from the main lineage during the Middle Pleistocene and became a distinct species around a million years ago. A divergence rate around 2% has been calculated. The split does not seem to have been driven by competition for resources, but instead because each species adopted a different ecological niche and occupied a different trophic level. Blue wildebeest fossils dating back some 2.5 million years ago are common and widespread. They have been found in the fossil-bearing caves at the
Cradle of Humankind The Cradle of Humankind is a paleoanthropological site and is located about northwest of Johannesburg, South Africa, in the Gauteng province. Declared a World Heritage Site by UNESCO in 1999, the site is home to the largest concentration of ...
north of Johannesburg. Elsewhere in South Africa, they are plentiful at such sites as
Elandsfontein Elandsfontein may refer to: * Elandsfontein, an archaeological site near Hopefield, South Africa * Elandsfontein, a farm homestead that is now a suburb of Alberton, South Africa {{disambiguation ...
, Cornelia, and Florisbad. The earliest fossils of the black wildebeest were found in sedimentary rock in Cornelia in the Free State and dated back about 800,000 years. Today, five subspecies of the blue wildebeest are recognised, while the black wildebeest has no named subspecies.


Genetics and hybrids

The diploid number of chromosomes in the wildebeest is 58. Chromosomes were studied in a male and a female wildebeest. In the female, all except a pair of very large submetacentric chromosomes were found to be acrocentric. Metaphases were studied in the male's chromosomes, and very large submetacentric chromosomes were found there, as well, similar to those in the female both in size and morphology. Other chromosomes were acrocentric. The X chromosome is a large acrocentric and the Y chromosome a minute one. The two species of the wildebeest are known to hybridise. Male black wildebeest have been reported to mate with female blue wildebeest and vice versa. The differences in social behaviour and habitats have historically prevented interspecific hybridisation between the species, but hybridisation may occur when they are both confined within the same area. The resulting offspring are usually fertile. A study of these hybrid animals at
Spioenkop Dam Nature Reserve Spioenkop Dam Nature Reserve or Spion Kop Nature Reserve is a protected area in KwaZulu-Natal, South Africa. It lies close to Ladysmith with Winterton being the closest town, and is about . The historic battlefield site can be reached by roa ...
in South Africa revealed that many had disadvantageous abnormalities relating to their teeth, horns, and the
wormian bones Wormian bones, also known as intrasutural bones or sutural bones, are extra bone pieces that can occur within a suture (joint) in the skull. These are irregular isolated bones that can appear in addition to the usual centres of ossification of the ...
in the skull. Another study reported an increase in the size of the hybrid as compared to either of its parents. In some animals, the tympanic part of the temporal bone is highly deformed, and in others, the radius and ulna are fused.


Characteristics of the species

Both species of wildebeest are even-toed, horned, greyish-brown ungulates resembling cattle. Males are larger than females and both have heavy forequarters compared to their hindquarters. They have broad muzzles, Roman noses, and shaggy manes and tails. The most striking morphological differences between the black and blue wildebeest are the orientation and curvature of their horns and the colour of their coats. The blue wildebeest is the bigger of the two species. In males, blue wildebeest stand tall at the shoulder and weigh around , while the black wildebeest stands tall and weighs about . In females, blue wildebeest have a shoulder height of and weigh while black wildebeest females stand at the shoulder and weigh . The horns of blue wildebeest protrude to the side, then curve downwards before curving up back towards the skull, while the horns of the black wildebeest curve forward then downward before curving upwards at the tips. Blue wildebeest tend to be a dark grey colour with stripes, but may have a bluish sheen. The black wildebeest has brown-coloured hair, with a mane that ranges in colour from cream to black, and a cream-coloured tail. The blue wildebeest lives in a wide variety of habitats, including woodlands and grasslands, while the black wildebeest tends to reside exclusively in open grassland areas. In some areas, the blue wildebeest migrates over long distances in the winter, whereas the black wildebeest does not. The milk of the female black wildebeest contains a higher protein, lower fat, and lower lactose content than the milk of the blue wildebeest. Wildebeest can live more than 40 years, though their average lifespan is around 20 years.


Distribution and habitat

Wildebeest inhabit the plains and open woodlands of parts of Africa south of the Sahara. The black wildebeest is native to the southernmost parts of the continent. Its historical range included South Africa, Eswatini, and Lesotho, but in the latter two countries, it was hunted to extinction in the 19th century. It has now been reintroduced to them and also introduced to Namibia, where it has become well established. It inhabits open plains, grasslands, and Karoo shrublands in both steep mountainous regions and lower undulating hills at altitudes varying between . In the past, it inhabited the highveld temperate grasslands during the dry winter season and the arid Karoo region during the rains. However, as a result of widespread hunting, the black wildebeest no longer occupies its historical range or makes migrations, and is now largely limited to game farms and protected reserves. The blue wildebeest is native to eastern and southern Africa. Its range includes Kenya, Tanzania, Botswana, Zambia, Zimbabwe, Mozambique, South Africa,
Eswatini Eswatini ( ; ss, eSwatini ), officially the Kingdom of Eswatini and formerly named Swaziland ( ; officially renamed in 2018), is a landlocked country in Southern Africa. It is bordered by Mozambique to its northeast and South Africa to its no ...
and Angola. It is no longer found in Malawi but has been successfully reintroduced into Namibia. Blue wildebeest are mainly found in short grass plains bordering bush-covered acacia savannas, thriving in areas that are neither too wet nor too dry. They can be found in habitats that vary from overgrazed areas with dense bush to open woodland floodplains. In East Africa, the blue wildebeest is the most abundant big
game A game is a structured form of play (activity), play, usually undertaken for enjoyment, entertainment or fun, and sometimes used as an educational tool. Many games are also considered to be work (such as professional players of spectator s ...
species, both in population and biomass. It is a notable feature of the Serengeti National Park in Tanzania, the Maasai Mara National Reserve in Kenya and the
Liuwa Plain National Park Liuwa Plain National Park is a national park in Zambia's Western Province. "Liuwa" means "plain" in the local Liuwa language, a dialect of Lozi language, and the plains originally served as a hunting ground for Lubosi Lewanika, the Litunga (ki ...
in Zambia.


Migration

Not all wildebeest are migratory. Black wildebeest herds are often nomadic or may have a regular home range of . Bulls may occupy territories, usually about apart, but this spacing varies according to the quality of the habitat. In favourable conditions, they may be as close as , or they may be as far apart as in poor habitat. Female herds have home ranges of about in size. Herds of nonterritorial bachelor males roam at will and do not seem to have any restrictions on where they wander. Blue wildebeest have both migratory and sedentary populations. In the
Ngorongoro The Ngorongoro Conservation Area (, ) is a protected area and a UNESCO World Heritage Site located in Ngorongoro District, west of Arusha City in Arusha Region, within the Crater Highlands geological area of northern Tanzania. The area is nam ...
, most animals are sedentary and males maintain a network of territories throughout the year, though breeding is seasonal in nature. Females and young form groups of about 10 individuals or join in larger aggregations, and nonterritorial males form bachelor groups. In the Serengeti and
Tarangire The Tarangire River is a perennial river located in central Manyara Region in the eastern branch of the East African Rift Valley, within northern Tanzania. Course The headwaters of the Tarangire River are in the highlands and escarpments of Baba ...
ecosystems, populations are mostly migratory, with herds consisting of both sexes frequently moving, but resident subpopulations also exist. During the rutting season, the males may form temporary territories for a few hours or a day or so, and attempt to gather together a few females with which to mate, but soon they have to move on, often moving ahead to set up another temporary territory. In the
Maasai Mara Maasai Mara, also sometimes spelled Masai Mara and locally known simply as The Mara, is a large national game reserve in Narok, Kenya, contiguous with the Serengeti National Park in Tanzania. It is named in honor of the Maasai people, the ancestr ...
game reserve, a non-migratory population of blue wildebeest had dwindled from about 119,000 animals in 1977 to about 22,000 in 1997. The reason for the decline is thought to be the increasing competition between cattle and wildebeest for a diminishing area of grazing land as a result of changes in agricultural practices, and possibly fluctuations in rainfall. Each year, some East African populations of blue wildebeest have a long-distance migration, seemingly timed to coincide with the annual pattern of rainfall and grass growth. The timing of their migrations in both the rainy and dry seasons can vary considerably (by months) from year to year. At the end of the wet season (May or June in East Africa), wildebeest migrate to dry-season areas in response to a lack of surface (drinking) water. When the rainy season begins again (months later), animals quickly move back to their wet-season ranges. Factors suspected to affect migration include food abundance, surface water availability, predators, and phosphorus content in grasses. Phosphorus is a crucial element for all life forms, particularly for lactating female bovids. As a result, during the rainy season, wildebeest select grazing areas that contain particularly high phosphorus levels. One study found, in addition to phosphorus, wildebeest select ranges containing grass with relatively high nitrogen content. Aerial photography has revealed that a level of organisation occurs in the movement of the herd that cannot be apparent to each individual animal; for example, the migratory herd exhibits a wavy front, and this suggests that some degree of local decision-making is taking place. Numerous documentaries feature wildebeest crossing rivers, with many being eaten by
crocodile Crocodiles (family (biology), family Crocodylidae) or true crocodiles are large semiaquatic reptiles that live throughout the tropics in Africa, Asia, the Americas and Australia. The term crocodile is sometimes used even more loosely to inclu ...
s or drowning in the attempt. While having the appearance of a frenzy, recent research has shown a herd of wildebeest possesses what is known as a "
swarm intelligence Swarm intelligence (SI) is the collective behavior of decentralized, self-organized systems, natural or artificial. The concept is employed in work on artificial intelligence. The expression was introduced by Gerardo Beni and Jing Wang in 1989, in ...
", whereby the animals systematically explore and overcome the obstacle as one.


Ecology


Predators

Major predators that feed on wildebeest include the
lion The lion (''Panthera leo'') is a large Felidae, cat of the genus ''Panthera'' native to Africa and India. It has a muscular, broad-chested body; short, rounded head; round ears; and a hairy tuft at the end of its tail. It is sexually dimorphi ...
,
hyena Hyenas, or hyaenas (from Ancient Greek , ), are feliform carnivoran mammals of the family Hyaenidae . With only four extant species (each in its own genus), it is the fifth-smallest family in the Carnivora and one of the smallest in the clas ...
, African wild dog, cheetah,
leopard The leopard (''Panthera pardus'') is one of the five extant species in the genus '' Panthera'', a member of the cat family, Felidae. It occurs in a wide range in sub-Saharan Africa, in some parts of Western and Central Asia, Southern Russia, a ...
, and
crocodile Crocodiles (family (biology), family Crocodylidae) or true crocodiles are large semiaquatic reptiles that live throughout the tropics in Africa, Asia, the Americas and Australia. The term crocodile is sometimes used even more loosely to inclu ...
, which seem to favour the wildebeest over other prey. Wildebeest, however, are very strong, and can inflict considerable injury even to a lion. Wildebeest have a maximum running speed of around . The primary defensive tactic is herding, where the young animals are protected by the older, larger ones, while the herd runs as a group. Typically, the predators attempt to isolate a young or ill animal and attack without having to worry about the herd. Wildebeest have developed additional sophisticated cooperative behaviours, such as animals taking turns sleeping while others stand guard against a night attack by invading predators. Wildebeest migrations are closely followed by vultures, as wildebeest carcasses are an important source of food for these scavengers. The vultures consume about 70% of the wildebeest carcasses available. Decreases in the number of migrating wildebeest have also had a negative effect on the vultures. In the Serengeti ecosystem, Tanzania, wildebeest may help facilitate the migration of other, smaller-bodied grazers, such as Thomson's gazelles (''Eudorcas thomsonii''), which eat the new-growth grasses stimulated by wildebeest foraging.


Interactions with nonpredators

Zebras and wildebeest group together in open savannah environments with high chances of predation. This grouping strategy reduces predation risk because larger groups decrease each individual's chance of being hunted, and predators are more easily seen in open areas. The seasonal presence of thousands of migratory wildebeests reduces local lion predation on giraffe calves, resulting in greater survival of giraffes. Wildebeest can also listen in on the alarm calls of other species, and by doing so, can reduce their risk of predation. One study showed, along with other ungulates, wildebeests responded more strongly to the baboon alarm calls compared to the baboon contest calls, though both types of calls had similar patterns, amplitudes, and durations. The alarm calls were a response of the baboons to lions, and the contest calls were recorded when a dispute between two males occurred. Wildebeest compete with domesticated livestock for pasture and are sometimes blamed by farmers for transferring diseases and parasites to their cattle.


Breeding and reproduction

Wildebeest do not form permanent pair bonds and during the mating season, or rut, the males establish temporary territories and try to attract females into them. These small territories are about , with up to 300 territories per . The males defend these territories from other males while competing for females that are coming into oestrus. The males use grunts and distinctive behaviour to entice females into their territories. Wildebeest usually breed at the end of the rainy season when the animals are well fed and at their peak of fitness. This usually occurs between May and July, and birthing usually takes place between January and March, at the start of the wet season. Wildebeest females breed seasonally and ovulate spontaneously. The estrous cycle is about 23 days and the gestation period lasts 250 to 260 days. The calves weigh about at birth and scramble to their feet within minutes, being able to move with the herd soon afterwards, a fact on which their survival relies. The main predator of the calves is the
spotted hyena The spotted hyena (''Crocuta crocuta''), also known as the laughing hyena, is a hyena species, currently classed as the sole extant member of the genus ''Crocuta'', native to sub-Saharan Africa. It is listed as being of least concern by the IUC ...
. The calving peak period lasts for 2–3 weeks, and in small subpopulations and isolated groups, mortality of calves may be as high as 50%. However, in larger aggregations, or small groups living near large herds, mortality rates may be under 20%. Groups of wildebeest females and young live in the small areas established by the male. When groups of wildebeest join, the female to male ratio is higher because the females choose to move to the areas held by a smaller number of males. This female-dominated sex ratio may be due to illegal hunting and human disturbance, with higher male mortality having been attributed to hunting.


Threats and conservation

The
black wildebeest The black wildebeest or white-tailed gnu (''Connochaetes gnou'') is one of the two closely related wildebeest species. It is a member of the genus '' Connochaetes'' and family Bovidae. It was first described in 1780 by Eberhard August Wilhelm ...
has been classified as a least-concern species by the International Union for Conservation of Nature in the '' IUCN Red List''. The populations of this species are on an increase. Now, more than 18,000 individuals are believed to remain, 7,000 of which are in Namibia, outside their natural range, and where it is farmed. Around 80% of the wildebeest occurs in private areas, while the other 20% is confined in protected areas. Its introduction into Namibia has been a success and numbers have increased substantially there from 150 in 1982 to 7,000 in 1992. The blue wildebeest has also been rated as of least concern. The population trend is stable, and their numbers are estimated to be around 1,500,000 – mainly due to the increase of the populations in Serengeti National Park (Tanzania) to 1,300,000 as of 1998. However, the numbers of one of the subspecies, the eastern white-bearded wildebeest (''C. t. albojubatus'') have seen a steep decline. Population density ranges from 0.15/km2. in Hwange and Etosha National Parks to 35/km2 in Ngorongoro Conservation Area and Serengeti National Park. Overland migration as a biological process requires large, connected landscapes, which are increasingly difficult to maintain, particularly over the long term, when human demands on the landscape compete. The most acute threat comes from migration barriers, such as fences and roads. In one of the more striking examples of the consequences of fence-building on terrestrial migrations, Botswanan authorities placed thousands of kilometres of fences across the Kalahari that prevented wildebeests from reaching watering holes and grazing grounds, resulting in the deaths of tens of thousands of individuals, reducing the wildebeest population to less than 10% of its previous size. Illegal hunting is a major conservation concern in many areas, along with natural threats posed by main predators (which include lions, leopards, African hunting dogs, cheetahs and hyenas). Where the black and blue wildebeest share a common range, the two can hybridise, and this is regarded as a potential threat to the black wildebeest.


Uses and interaction with humans

Wildebeest provide several useful animal products. The hide makes good-quality leather and the flesh is coarse, dry and rather hard. Wildebeest are killed for food, especially to make biltong in Southern Africa. This dried game meat is a delicacy and an important food item in Africa. The meat of females is more tender than that of males, and is the most tender during the autumn season. Wildebeest are a regular target for illegal meat hunters because their numbers make them easy to find. Cooks preparing the wildebeest carcass usually cut it into 11 pieces. The estimated price for wildebeest meat was about US$0.47 per around 2008. The silky, flowing tail of the black wildebeest is used to make fly-whisks or ''chowries''. Wildebeest benefit the ecosystem by increasing soil fertility with their excreta. They are economically important for human beings, as they are a major tourist attraction. They also provide important products, such as leather, to humans. Wildebeest, however, can also have a negative impact on humans. Wild individuals can be competitors of commercial livestock, and can transmit diseases and cause epidemics among animals, particularly domestic cattle. They can also spread ticks, lungworms, tapeworms, flies, and paramphistome flukes.


Cultural depictions

The wildebeest is depicted on the coat of arms of the
Province of Natal The Province of Natal (), commonly called Natal, was a province of South Africa from May 1910 until May 1994. Its capital was Pietermaritzburg. During this period rural areas inhabited by the black African population of Natal were organized into ...
and later
KwaZulu-Natal KwaZulu-Natal (, also referred to as KZN and known as "the garden province") is a province of South Africa that was created in 1994 when the Zulu bantustan of KwaZulu ("Place of the Zulu" in Zulu) and Natal Province were merged. It is locate ...
in South Africa. Over the years, the South African authorities have issued several stamps displaying the animal and the South African Mint has struck a two-cent piece with a prancing black wildebeest. Movies and television shows also feature wildebeests, including ''
Khumba ''Khumba'' is a 2013 South African computer-animated comedy film directed and co-produced by Anthony Silverston and written by Silverston and Raffaella Delle Donne. The film stars the voices of Jake T. Austin, Steve Buscemi, Loretta Devine, Lau ...
'' (Mama V), '' The Wild'' (Kazar and his minions), '' All Hail King Julien'' (Vigman Wildebeest), '' Phineas and Ferb'' (Newton the Gnu), '' The Great Space Coaster'' (newscaster Gary Gnu), and '' The Lion King'' (the wildebeest stampede that resulted in Mufasa's death). Michael Flanders wrote a humorous song called " The Gnu", which was very popular when he performed it, with Donald Swann, in a revue called ''
At the Drop of a Hat ''At the Drop of a Hat'' is a musical revue by Flanders and Swann, described by them as "an after-dinner farrago". In the show, they both sang on a nearly bare stage, accompanied by Swann on the piano. The songs were linked by contemporary socia ...
'', which opened in London on 31 December 1956. In the 1970s, British tea brand Typhoo ran a series of television advertisements featuring an animated anthropomorphic gnu character. The wildebeest is the mascot of the GNU Project and GNU operating system. In the ''Llama Llama'' picture-book series by
Anna Dewdney Anna Elizabeth Dewdney (née Luhrmann; December 25, 1965 – September 3, 2016) was an American author and illustrator of children's books. The first book she wrote and illustrated, ''Llama Llama Red Pajama'', received critical acclaim in 2005. ...
, an anthropomorphised wildebeest named Nelly Gnu is the main character, Llama Llama's best friend, and is also featured in a title of her own, ''Nelly Gnu and Daddy Too''. In Terry Pratchett's '' Discworld'' series, Discworld analogues of wildebeests, which inhabited Howonderland, were called "bewilderbeest".


See also

*
Keystone species A keystone species is a species which has a disproportionately large effect on its natural environment relative to its abundance, a concept introduced in 1969 by the zoologist Robert T. Paine. Keystone species play a critical role in maintaini ...


References

{{Taxonbar, from=Q7609 Alcelaphinae Mammals of Africa Extant Pleistocene first appearances Herbivorous mammals Articles containing video clips