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Jojoba (; botanical name: ''Simmondsia chinensis'')also commonly called goat nut, deer nut, pignut, wild hazel, quinine nut, coffeeberry, and gray box bushis native to the Southwestern United States. ''Simmondsia chinensis'' is the sole species of the family Simmondsiaceae, placed in the order Caryophyllales. Jojoba is grown commercially to produce jojoba oil, a liquid wax ester extracted from its seed.


Distribution

The plant is a native
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
of the Sonoran Desert,
Colorado Desert California's Colorado Desert is a part of the larger Sonoran Desert. It encompasses approximately , including the heavily irrigated Coachella and Imperial valleys. It is home to many unique flora and fauna. Geography and geology The Colorado De ...
, Baja California Desert, and
California chaparral and woodlands The California chaparral and woodlands is a terrestrial ecoregion of southwestern Oregon, northern, central, and southern California (United States) and northwestern Baja California (Mexico), located on the west coast of North America. It is a ...
habitats in the
Peninsular Ranges The Peninsular Ranges (also called the Lower California province) are a group of mountain ranges that stretch from Southern California to the southern tip of the Baja California peninsula; they are part of the North American Coast Ranges, which ...
and San Jacinto Mountains. It is found in southern California, Arizona, and Utah (U.S.), and Baja California state (Mexico). Jojoba is endemic to North America, and occupies an area of approximately between latitudes 25° and 31° North and between longitudes 109° and 117° West.


Description

''Simmondsia chinensis'', or jojoba, typically grows to tall, with a broad, dense crown, but there have been reports of plants as tall as . The
leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...
are opposite, ovalish in shape, long and broad, thick, waxy, and glaucous gray-green in color. The flowers are small and greenish-yellow, with 5–6 sepals and no petals. The plant typically blooms from March to May.


Reproduction

Each plant is
dioecious Dioecy (; ; adj. dioecious , ) is a characteristic of a species, meaning that it has distinct individual organisms (unisexual) that produce male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproductio ...
, with hermaphrodites being extremely rare. The fruit is an acorn-shaped ovoid, three-angled capsule long, partly enclosed at the base by the sepals. The mature seed is a hard oval that is dark brown and contains an oil (liquid wax) content of approximately 54%. An average-sized bush produces of
pollen Pollen is a powdery substance produced by seed plants. It consists of pollen grains (highly reduced microgametophytes), which produce male gametes (sperm cells). Pollen grains have a hard coat made of sporopollenin that protects the gametophyt ...
, to which few humans are allergic. The female plants produce seed from flowers pollinated by the male plants. Jojoba leaves have an aerodynamic shape, creating a spiral effect, which brings wind-borne pollen from the male flower to the female flower. In the Northern Hemisphere, pollination occurs during February and March. In the Southern Hemisphere, pollination occurs during August and September. Somatic cells of jojoba are tetraploid; the number of chromosomes is 2''n'' = 4''x'' = 52. File:Jojoba-weibliche-blüte.JPG, Female flower Image:Simmondsia chinensis male flower.jpg, Close-up of male ''Simmondsia chinensis'' flowers File:Jojoba.jpg, Jojoba fruits Image:Jojoba.seed.jpg, Jojoba seed


Genetics

The jojoba genome was sequenced in 2020 and reported to be 887-Mb, consisting of 26 chromosomes (2n = 26), and is predicted to have 23,490 protein-coding genes.


Taxonomy

Despite its scientific name ''Simmondsia chinensis'', the plant is not native to
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
. The botanist Johann Link originally named the species ''Buxus chinensis'', after misreading a collection label "Calif", referring to California, as "China". Jojoba was collected again in 1836 by Thomas Nuttall who described it as a new genus and species in 1844, naming it ''Simmondsia californica'', but priority rules require that the original specific epithet be used. The common name "jojoba" originated from O'odham name ''Hohowi''. The common name should not be confused with the similarly written jujube (''Ziziphus zizyphus''), an unrelated plant species, which is commonly grown in China.


Uses

Jojoba foliage provides year-round food for many animals, including deer, javelina, bighorn sheep, and livestock. Its nuts are eaten by squirrels, rabbits, other rodents, and larger birds. Only
Bailey's pocket mouse Bailey's pocket mouse (''Chaetodipus baileyi'') is a species of rodent of the subfamily Perognathinae, family Heteromyidae. It is found in Baja California, Sinaloa and Sonora in Mexico and in California, Arizona and New Mexico in the United Sta ...
, however, is known to be able to digest the wax found inside the jojoba nut. In large quantities, jojoba seed meal is toxic to many mammals. Later this effect was found to be due to simmondsin, which inhibits hunger. The indigestible wax acts as a laxative in humans.


Native American uses

Native Americans first made use of jojoba. During the early 18th century
Jesuit , image = Ihs-logo.svg , image_size = 175px , caption = ChristogramOfficial seal of the Jesuits , abbreviation = SJ , nickname = Jesuits , formation = , founders ...
missionaries on the Baja California Peninsula observed indigenous peoples heating jojoba seeds to soften them. They then used a mortar and pestle to create a salve or buttery substance. The latter was applied to the skin and hair to heal and condition. The O'odham people of the Sonoran Desert treated burns with an
antioxidant Antioxidants are compounds that inhibit oxidation, a chemical reaction that can produce free radicals. This can lead to polymerization and other chain reactions. They are frequently added to industrial products, such as fuels and lubricant ...
salve made from a paste of the jojoba nut. Native Americans also used the salve to soften and preserve
animal hides A hide or skin is an animal skin treated for human use. The word "hide" is related to the German word "Haut" which means skin. The industry defines hides as "skins" of large animals ''e.g''. cow, buffalo; while skins refer to "skins" of smaller an ...
. Pregnant women ate jojoba seeds, believing they assisted during childbirth. Hunters and raiders ate jojoba on the trail to keep hunger at bay. The
Seri Seri or SERI may refer to: People *Jean Michaël Seri, an Ivorian professional footballer Places *Seri Yek-e Zarruk, Iran *Seri, Bheri, Nepal *Seri, Karnali, Nepal *Seri, Mahakali, Nepal *Seri, Raebareli, a village in Uttar Pradesh, India Oth ...
, who utilize nearly every edible plant in their domain, do not regard the beans as real food and in the past ate it only in emergencies.


Introduction to Europe

Archibald Menzies was the botanist with the Vancouver Expedition that arrived in Santa Barbara, California in November 1793. He was given fruit and plants of the jojoba by padre of the San Diego Mission. These survived the voyage back to the UK and were planted in the Royal Botanic Gardens at Kew near London.


Contemporary uses

Jojoba is grown for the liquid wax, commonly called jojoba oil, in its seeds. The oil is rare in that it is an extremely long (C36–C46) straight-chain wax ester and not a triglyceride, making jojoba and its derivative
jojoba ester Jojoba esters are the hydrogenation or transesterification product of Jojoba oil. Jojoba Esters are commonly used in cosmetic formulations as an emollient, due to its remarkable similarity to the natural oils produced by the human skin, and its hi ...
s more similar to whale oil than to traditional vegetable oils. Jojoba oil has also been discussed as a possible biodiesel fuel, but it cannot be cultivated on a scale to compete with traditional fossil fuels, so its use is restricted to personal care products..


Cultivation

Plantations of jojoba have been established in a number of desert and semi-desert areas, predominantly in Argentina, Australia, Israel, Mexico, Peru and the United States. It is currently the Sonoran Desert's second most economically valuable native plant (overshadowed only by ''
Washingtonia filifera ''Washingtonia filifera'', the desert fan palm, California fan palm, or California palm,Flora of North America Association. ''Flora of North America: North of Mexico Volume 22: Magnoliophyta: Alismatidae, Arecidae, Commelinidae (in Part), and Zi ...
''—California fan palms, used as ornamental trees). Jojoba prefers light, coarsely textured soils. Good drainage and water penetration is necessary. It tolerates
salinity Salinity () is the saltiness or amount of salt dissolved in a body of water, called saline water (see also soil salinity). It is usually measured in g/L or g/kg (grams of salt per liter/kilogram of water; the latter is dimensionless and equal ...
and nutrient-poor soils. Soil pH should be between 5 and 8. High temperatures are tolerated by jojoba, but frost can damage or kill plants. Requirements are minimal, so jojoba plants do not need intensive cultivation. Weed problems only occur during the first two years after planting and there is little damage by insects. Supplemental irrigation could maximize production where rainfall is less than 400 mm. There is no need for high
fertilisation Fertilisation or fertilization (see spelling differences), also known as generative fertilisation, syngamy and impregnation, is the fusion of gametes to give rise to a new individual organism or offspring and initiate its development. Proce ...
, but, especially in the first year, nitrogen increases growth. Jojoba is normally harvested by hand because seeds do not all mature in the same time. Yield is around 3.5 t/ha depending on the age of the plantation. Selective breeding is developing plants that produce more beans with higher wax content, as well as other characteristics that will facilitate harvesting. Its ability to withstand high salinity up to 12 ds −1 at pH 9) (deciSiemens or ECe Salt tolerance of crops) and the high value of jojoba products make jojoba an interesting plant for the use of desertification control. It has been used to Desertification#Countering desertification, combat and prevent desertification in the Thar Desert in India.


References


External links


USDA Plants Profile for ''Simmondsia chinensis'' (jojoba)

Calflora Database: ''Simmondsia chinensis'' (jojoba)

Jepson Manual (TJM93) treatment of ''Simmondsia chinensis''


€”an explanation of the scientific name



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Uses and Benefits of Jojoba Oils
{{Taxonbar, from=Q267749 Caryophyllales Crops originating from Mexico Crops originating from the United States Flora of Arizona Flora of Baja California Flora of California Flora of the California desert regions Flora of the Sonoran Deserts Flora of Utah Natural history of the California chaparral and woodlands Natural history of the Colorado Desert Natural history of the Peninsular Ranges North American desert flora Waxes Dioecious plants Flora without expected TNC conservation status