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The Phanerozoic Eon is the current
geologic eon The geologic time scale, or geological time scale, (GTS) is a representation of time based on the rock record of Earth. It is a system of chronological dating that uses chronostratigraphy (the process of relating strata to time) and geochronol ...
in the
geologic time scale The geologic time scale, or geological time scale, (GTS) is a representation of time based on the rock record of Earth. It is a system of chronological dating that uses chronostratigraphy (the process of relating strata to time) and geochrono ...
, and the one during which abundant animal and plant life has existed. It covers 538.8 million years to the present, and it began with the
Cambrian The Cambrian Period ( ; sometimes symbolized C with bar, Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million ...
Period, when animals first developed hard shells preserved in the fossil record. The time before the Phanerozoic, called the ''
Precambrian The Precambrian (or Pre-Cambrian, sometimes abbreviated pꞒ, or Cryptozoic) is the earliest part of Earth's history, set before the current Phanerozoic Eon. The Precambrian is so named because it preceded the Cambrian, the first period of the ...
'', is now divided into the
Hadean The Hadean ( ) is a Eon (geology), geologic eon of History of Earth, Earth history preceding the Archean. On Earth, the Hadean began with the Formation of the Earth, planet's formation about 4.54 billion years ago (although the start of the H ...
, Archaean and
Proterozoic The Proterozoic () is a geological eon spanning the time interval from 2500 to 538.8million years ago. It is the most recent part of the Precambrian "supereon". It is also the longest eon of the Earth's geologic time scale, and it is subdivided ...
eons. The time span of the Phanerozoic starts with the sudden appearance of fossilised evidence of a number of animal
phyla Phyla, the plural of ''phylum'', may refer to: * Phylum, a biological taxon between Kingdom and Class * by analogy, in linguistics, a large division of possibly related languages, or a major language family which is not subordinate to another Phyl ...
; the evolution of those phyla into diverse forms; the emergence and development of complex plants; the evolution of fish; the emergence of insects and tetrapods; and the development of modern fauna. Plant life on land appeared in the early Phanerozoic eon. During this time span, tectonic forces which move the continents had collected them into a single landmass known as Pangaea (the most recent supercontinent), which then separated into the current continental landmasses.


Etymology

The term ''Phanerozoic'' derives from the Ancient Greek words (), meaning ''visible'', and (), meaning ''life''; since it was once believed that life began in the
Cambrian The Cambrian Period ( ; sometimes symbolized C with bar, Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million ...
, the first period of this eon. The term "Phanerozoic" was coined in 1930 by the American geologist George Halcott Chadwick (1876–1953).


Proterozoic-Phanerozoic boundary

The
Proterozoic The Proterozoic () is a geological eon spanning the time interval from 2500 to 538.8million years ago. It is the most recent part of the Precambrian "supereon". It is also the longest eon of the Earth's geologic time scale, and it is subdivided ...
-Phanerozoic boundary is at 538.8 million years ago. In the 19th century, the boundary was set at time of appearance of the first abundant animal ( metazoan) fossils but several hundred groups ( taxa) of metazoa of the preceding Proterozoic eon have been identified since the systematic study of those forms started in the 1950s.


Eras of the Phanerozoic

The Phanerozoic is divided into three
eras Eras is a humanist sans-serif typeface designed by Albert Boton and Albert Hollenstein and was released by the International Typeface Corporation (ITC) in 1976. Eras is licensed by the Linotype type foundry. A distinct and curious feature of E ...
: the Paleozoic, Mesozoic, and
Cenozoic The Cenozoic ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterised by the dominance of mammals, birds and flowering plants, a cooling and drying climate, and the current configura ...
, which are further subdivided into 12 periods. The Paleozoic features the evolution of fish, amphibians and reptiles. The Mesozoic features the evolution of lizards, crocodiles, snakes, turtles, mammals, and dinosaurs (including birds). The Cenozoic begins with the extinction of the non-avian dinosaurs, and features evolution of great diversity in birds and mammals. Humans appeared and evolved during the most recent part of the Cenozoic.


Paleozoic Era

The Paleozoic is a time in Earth's history when complex life forms evolved, took their first breath of oxygen on dry land, and when the forerunners of all multicelular life on Earth began to diversify. There are six periods in the Paleozoic era:
Cambrian The Cambrian Period ( ; sometimes symbolized C with bar, Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million ...
, Ordovician,
Silurian The Silurian ( ) is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at million years ago ( Mya), to the beginning of the Devonian Period, Mya. The Silurian is the shortest period of the Paleozo ...
,
Devonian The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, whe ...
,
Carboniferous The Carboniferous ( ) is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period million years ago ( Mya), to the beginning of the Permian Period, million years ago. The name ''Carbonifero ...
and Permian.


Cambrian Period

The Cambrian is the first period of the Paleozoic Era and ran from 539 million to 485 million years ago. The Cambrian sparked a rapid expansion in the diversity of animals, in an event known as the
Cambrian explosion The Cambrian explosion, Cambrian radiation, Cambrian diversification, or the Biological Big Bang refers to an interval of time approximately in the Cambrian Period when practically all major animal phyla started appearing in the fossil recor ...
, during which the greatest number of animal body plans evolved in a single period in the history of Earth. Complex algae evolved, and the fauna was dominated by armoured arthropods, such as trilobites. Almost all
phyla Phyla, the plural of ''phylum'', may refer to: * Phylum, a biological taxon between Kingdom and Class * by analogy, in linguistics, a large division of possibly related languages, or a major language family which is not subordinate to another Phyl ...
of marine animals evolved in this period. During this time, the super-continent
Pannotia Pannotia (from Greek: '' pan-'', "all", '' -nótos'', "south"; meaning "all southern land"), also known as the Vendian supercontinent, Greater Gondwana, and the Pan-African supercontinent, was a relatively short-lived Neoproterozoic supercontinent ...
began to break up, most of which later recombined into the super-continent
Gondwana Gondwana () was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic (about 550 million years ago) and began to break up during the Jurassic period (about 180 million years ago). The final stages ...
.


Ordovician Period

The Ordovician spans from 485 million to 444 million years ago. The Ordovician was a time in Earth's history in which many groups still prevalent today evolved or diversified, such as primitive
cephalopod A cephalopod is any member of the molluscan class Cephalopoda (Greek plural , ; "head-feet") such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral body symmetry, a prominent head ...
s, fish, and corals. This process is known as the
Great Ordovician Biodiversification Event The Great Ordovician Biodiversification Event (GOBE), was an evolutionary radiation of animal life throughout the Ordovician period, 40 million years after the Cambrian explosion, whereby the distinctive Cambrian fauna fizzled out to be replaced wi ...
, or GOBE. Trilobites began to be replaced by
articulate brachiopod Rhynchonelliformea is a major subphylum and clade of brachiopods. It is equivalent to the former class Articulata, which was used previously in brachiopod taxonomy. Articulate brachiopods have many anatomical differences relative to "inarticulate" ...
s, and crinoids also became an increasingly important part of the fauna. The first arthropods crept ashore to colonise
Gondwana Gondwana () was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic (about 550 million years ago) and began to break up during the Jurassic period (about 180 million years ago). The final stages ...
, a continent empty of animal life. By the end of the Ordovician, Gondwana had moved from the equator to the South Pole, and Laurentia had collided with Baltica, closing the
Iapetus Ocean The Iapetus Ocean (; ) was an ocean that existed in the late Neoproterozoic and early Paleozoic eras of the geologic timescale (between 600 and 400 million years ago). The Iapetus Ocean was situated in the southern hemisphere, between the paleoco ...
. The glaciation of Gondwana resulted in a major drop in sea level, killing off all life that had established along its coast. Glaciation caused an icehouse Earth, leading to the Ordovician–Silurian extinction, during which 60% of marine invertebrates and 25% of families became extinct. Though one of the deadliest mass extinctions in earth's history, the O–S extinction did not cause profound ecological changes between the periods.


Silurian Period

The Silurian spans from 444 million to 419 million years ago, which saw a warming from an icehouse Earth. This period saw the mass evolution of fish, as jawless fish became more numerous, and early jawed and freshwater fish appeared in the fossil record. Arthropods remained abundant, and some groups, such as eurypterids, became apex predators. Fully terrestrial life established itself on land, including early arachnids, fungi, and myriapods (many-legged arthropods). The evolution of vascular plants such as '' Cooksonia'' allowed plants to gain a foothold on land as well. These early terrestrial plants are the forerunners of all plant life on land. During this time, there were four continents:
Gondwana Gondwana () was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic (about 550 million years ago) and began to break up during the Jurassic period (about 180 million years ago). The final stages ...
(Africa, South America, Australia, Antarctica, India), Laurentia (North America with parts of Europe), Baltica (the rest of Europe), and Siberia (Northern Asia).


Devonian Period

The Devonian spans from 419 million to 359 million years ago. Also informally known as the "Age of the Fish", the Devonian features a huge diversification in fish. Armored fish included jawless "agnathans", as well as jawed
placoderms Placodermi (from Greek πλάξ 'plate' and δέρμα 'skin', literally 'Plate (animal anatomy), plate-skinned') is a Class (biology), class of armoured prehistoric fish, known from fossils, which lived from the Silurian to the end of the Devoni ...
such as '' Dunkleosteus.'' The Devonian also saw a diversification of modern fish groups such as chondricthyans (sharks and kin),
osteichthyans Osteichthyes (), popularly referred to as the bony fish, is a diverse superclass of fish that have skeletons primarily composed of bone tissue. They can be contrasted with the Chondrichthyes, which have skeletons primarily composed of cartilag ...
(ray-finned fish), and
sarcopterygians Sarcopterygii (; ) — sometimes considered synonymous with Crossopterygii () — is a taxon (traditionally a class or subclass) of the bony fishes known as the lobe-finned fishes. The group Tetrapoda, a mostly terrestrial superclass includin ...
(lobe-finned fish). One lineage of sarcopterygians evolved into the first four-limbed vertebrates, which would eventually become tetrapods. On land, plant groups diversified; the first trees and seeds evolved during this period. By the Middle Devonian, shrub-like forests of early plants existed: lycophytes, horsetails, ferns, and progymnosperm. This event also allowed the diversification of arthropod life as they took advantage of the new habitat. Near the end of the Devonian, 70% of all species became extinct in a sequence of mass extinction events, collectively known as the Late Devonian extinction.


Carboniferous Period

The Carboniferous spans from 359 million to 299 million years ago. Tropical swamps dominated the Earth, and the large amounts of trees created much of the carbon that became coal deposits (hence the name Carboniferous). About 90% of all coal beds were deposited in the Carboniferous and Permian periods, which represent just 2% of the Earth's geologic history. The high oxygen levels caused by these swamps allowed massive arthropods, normally limited in size by their
respiratory systems The respiratory system (also respiratory apparatus, ventilatory system) is a biological system consisting of specific organs and structures used for gas exchange in animals and plants. The anatomy and physiology that make this happen varies grea ...
, to proliferate. Tetrapods diversified during the Carboniferous, and one lineage acquired an amniotic egg which could survive outside of the water. These tetrapods, the amniotes, included the first
reptile Reptiles, as most commonly defined are the animals in the class Reptilia ( ), a paraphyletic grouping comprising all sauropsids except birds. Living reptiles comprise turtles, crocodilians, squamates (lizards and snakes) and rhynchocephalians ( ...
s and synapsids (mammal relatives). Throughout the Carboniferous, there was a cooling pattern, which eventually led to the glaciation of
Gondwana Gondwana () was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic (about 550 million years ago) and began to break up during the Jurassic period (about 180 million years ago). The final stages ...
as much of it was situated around the south pole. This event was known as the
Permo-Carboniferous Glaciation The Permo-Carboniferous refers to the time period including the latter parts of the Carboniferous and early part of the Permian period. Permo-Carboniferous rocks are in places not differentiated because of the presence of transitional fossils, an ...
and resulted in a major loss of area for coal forests, the Carboniferous rainforest collapse.


Permian Period

The Permian spans from 298 million to 251 million years ago and was the last period of the Paleozoic era. At its beginning, all continents came together to form the super-continent Pangaea, surrounded by one ocean called Panthalassa. The Earth was relatively dry compared to the Carboniferous, with harsh seasons, as the climate of the interior of Pangaea was not moderated by large bodies of water.
Amniote Amniotes are a clade of tetrapod vertebrates that comprises sauropsids (including all reptiles and birds, and extinct parareptiles and non-avian dinosaurs) and synapsids (including pelycosaurs and therapsids such as mammals). They are disti ...
s flourished and diversified in the new dry climate, particularly synapsids such as ''
Dimetrodon ''Dimetrodon'' ( or ,) meaning "two measures of teeth,” is an extinct genus of non-mammalian synapsid that lived during the Cisuralian (Early Permian), around 295–272 million years ago (Mya). It is a member of the family Sphenacodontid ...
,'' '' Edaphosaurus,'' and the ancestors of modern mammals. The first conifers evolved during this period, then dominated the terrestrial landscape. The Permian ended with at least one mass extinction, the Permian-Triassic mass extinction, an event sometimes known as " the Great Dying". This extinction was the largest in earth's history and led to the loss of 95% of all species of life.


Mesozoic Era

The Mesozoic ranges from 252 million to 66 million years ago. Also referred to as the Age of Reptiles or Age of Conifers, the Mesozoic featured the appearance of many modern tetrapods, as reptiles ascended to ecological dominance over synapsids. The Mesozoic is subdivided into three periods: the Triassic, Jurassic, and Cretaceous.


Triassic Period

The Triassic ranges from 252 million to 201 million years ago. The Triassic is a transitional time in Earth's history between the
Permian Extinction The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleozo ...
and the lush Jurassic Period. It has three major epochs: Early Triassic, Middle Triassic. and Late Triassic. The Early Triassic lasted between 252 million to 247 million years ago, and was a hot and arid epoch in the aftermath of the Permian Extinction. Many tetrapods during this epoch represented a disaster fauna, a group of animals with low diversity and cosmopolitanism (wide geographic ranges). Temnospondyli recovered and rediversified into large aquatic predators during the Triassic. Reptiles also diversified rapidly, with aquatic reptiles such as
ichthyosaur Ichthyosaurs (Ancient Greek for "fish lizard" – and ) are large extinct marine reptiles. Ichthyosaurs belong to the order known as Ichthyosauria or Ichthyopterygia ('fish flippers' – a designation introduced by Sir Richard Owen in 1842, altho ...
s and sauropterygians proliferating in the seas. On land, the first true
archosaur Archosauria () is a clade of diapsids, with birds and crocodilians as the only living representatives. Archosaurs are broadly classified as reptiles, in the cladistic sense of the term which includes birds. Extinct archosaurs include non-avian d ...
s appeared, including pseudosuchians (crocodile relatives) and avemetatarsalians (bird/dinosaur relatives). The Middle Triassic spans from 247 million to 237 million years ago. The Middle Triassic featured the beginnings of the break-up of Pangaea as rifting commenced in north Pangaea. The northern part of the Tethys Ocean, the Paleotethys Ocean, had become a passive basin, but a spreading center was active in the southern part of the Tethys Ocean, the Neotethys Ocean. Phytoplankton, coral, crustaceans, and many other invertebrates recovered from the Permian extinction by the end of the Middle Triassic. Meanwhile, on land, reptiles continued to diversify, conifer forests flourished, as well as the first flies. The Late Triassic spans from 237 million to 201 million years ago. Following the bloom of the Middle Triassic, the Late Triassic was warm and arid, with a strong monsoon climate and with most precipitation limited to coastal regions and high latitudes. The first true dinosaurs appeared early in the Late Triassic, and pterosaurs evolved a bit later. Other large reptilian competitors to the dinosaurs were wiped out by the Triassic–Jurassic extinction event, in which most archosaurs (excluding
crocodylomorphs Crocodylomorpha is a group of pseudosuchian archosaurs that includes the crocodilians and their extinct relatives. They were the only members of Pseudosuchia to survive the end-Triassic extinction. During Mesozoic and early Cenozoic times, cr ...
, pterosaurs, and dinosaurs), many synapsids, and almost all large amphibians became extinct, as well as 34% of marine life in the fourth mass extinction event. The cause of the extinction is debated, but likely resulted from eruptions of the
CAMP Camp may refer to: Outdoor accommodation and recreation * Campsite or campground, a recreational outdoor sleeping and eating site * a temporary settlement for nomads * Camp, a term used in New England, Northern Ontario and New Brunswick to descri ...
large igneous province.


Jurassic Period

The Jurassic ranges from 201 million to 145 million years ago, and features three major epochs: Early Jurassic,
Middle Jurassic The Middle Jurassic is the second epoch of the Jurassic Period. It lasted from about 174.1 to 163.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relatively rare, but geological formations co ...
, and Late Jurassic. The Early Jurassic Epoch spans from 201 million to 174 million years ago. The climate was much more humid than during the Triassic, and as a result, the world was warm and partially tropical, though possibly with short colder intervals.
Plesiosaur The Plesiosauria (; Greek: πλησίος, ''plesios'', meaning "near to" and ''sauros'', meaning "lizard") or plesiosaurs are an order or clade of extinct Mesozoic marine reptiles, belonging to the Sauropterygia. Plesiosaurs first appeared ...
s, ichthyosaurs and
ammonite Ammonoids are a group of extinct marine mollusc animals in the subclass Ammonoidea of the class Cephalopoda. These molluscs, commonly referred to as ammonites, are more closely related to living coleoids (i.e., octopuses, squid and cuttlefish) ...
s dominated the seas, while dinosaurs and other reptiles dominated the land, with species such as '' Dilophosaurus'' at the apex. Crocodylomorphs evolved into aquatic forms, pushing the large amphibians to near extinction. True mammals were present during the Jurassic but remained small, with average body masses of less than until the end of the Cretaceous. The Middle and Late Jurassic Epochs span from 174 million to 145 million years ago. Conifer savannahs made up a large portion of the world's forests. In the oceans, plesiosaurs were quite common, and ichthyosaurs were flourishing. The Late Jurassic Epoch spans from 163 million to 145 million years ago. The Late Jurassic featured a severe extinction of sauropods in northern continents, alongside many ichthyosaurs. However, the Jurassic-Cretaceous boundary did not strongly impact most forms of life.


Cretaceous Period

The Cretaceous is the Phanerozoic's longest period, and the last period of the Mesozoic. It spans from 145 million to 66 million years ago, and is divided into two epochs:
Early Cretaceous The Early Cretaceous ( geochronological name) or the Lower Cretaceous (chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145  Ma to 100.5 Ma. Geology Pro ...
, and Late Cretaceous. The Early Cretaceous Epoch spans from 145 million to 100 million years ago. Dinosaurs continued to be abundant, with groups such as
tyrannosauroids Tyrannosauroidea (meaning 'tyrant lizard forms') is a superfamily (or clade) of coelurosaurian theropod dinosaurs that includes the family Tyrannosauridae as well as more basal relatives. Tyrannosauroids lived on the Laurasian supercontinent be ...
,
avialans Avialae ("bird wings") is a clade containing the only living dinosaurs, the birds. It is usually defined as all theropod dinosaurs more closely related to birds (Aves) than to deinonychosaurs, though alternative definitions are occasionally used ...
(birds), marginocephalians, and ornithopods seeing early glimpses of later success. Other tetrapods, such as stegosaurs and ichthyosaurs, declined significantly, and sauropods were restricted to southern continents. The Late Cretaceous Epoch spans from 100 million to 66 million years ago. The Late Cretaceous featured a cooling trend that would continue into the
Cenozoic The Cenozoic ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterised by the dominance of mammals, birds and flowering plants, a cooling and drying climate, and the current configura ...
Era. Eventually, the tropical climate was restricted to the equator and areas beyond the tropic lines featured more seasonal climates. Dinosaurs still thrived as new species such as '' Tyrannosaurus'', '' Ankylosaurus'', '' Triceratops'' and hadrosaurs dominated the food web. Whether or not pterosaurs went into a decline as birds radiated is debated; however, many families survived until the end of the Cretaceous, alongside new forms such as the gigantic ''
Quetzalcoatlus ''Quetzalcoatlus'' is a genus of pterosaur known from the Late Cretaceous period of North America (Maastrichtian stage); its members were among the largest known flying animals of all time. ''Quetzalcoatlus'' is a member of the Azhdarchidae, ...
''. Mammals diversified despite their small sizes, with
metatheria Metatheria is a mammalian clade that includes all mammals more closely related to marsupials than to placentals. First proposed by Thomas Henry Huxley in 1880, it is a more inclusive group than the marsupials; it contains all marsupials as well ...
ns ( marsupials and kin) and eutherians (
placentals Placental mammals (infraclass Placentalia ) are one of the three extant subdivisions of the class Mammalia, the other two being Monotremata and Marsupialia. Placentalia contains the vast majority of extant mammals, which are partly distinguishe ...
and kin) coming into their own. In the oceans,
mosasaurs Mosasaurs (from Latin ''Mosa'' meaning the 'Meuse', and Greek ' meaning 'lizard') comprise a group of extinct, large marine reptiles from the Late Cretaceous. Their first fossil remains were discovered in a limestone quarry at Maastricht on the ...
diversified to fill the role of the now-extinct ichthyosaurs, alongside huge plesiosaurs such as ''
Elasmosaurus ''Elasmosaurus'' (;) is a genus of plesiosaur that lived in North America during the Campanian stage of the Late Cretaceous period, about 80.5million years ago. The first specimen was discovered in 1867 near Fort Wallace, Kansas, US, and was se ...
''. Also, the first flowering plants evolved. At the end of the Cretaceous, the
Deccan Traps The Deccan Traps is a large igneous province of west-central India (17–24°N, 73–74°E). It is one of the largest volcanic features on Earth, taking the form of a large shield volcano. It consists of numerous layers of solidified flood ...
and other volcanic eruptions were poisoning the atmosphere. As this was continued, it is thought that a large meteor smashed into Earth, creating the Chicxulub Crater and the event known as the K–Pg extinction, the fifth and most recent mass extinction event, during which 75% of life on Earth became extinct, including all non-avian dinosaurs. Every living thing with a body mass over 10 kilograms became extinct, and the age of the dinosaurs came to an end.


Cenozoic Era

The Cenozoic featured the rise of mammals as the dominant class of animals, as the end of the age of the dinosaurs left significant open niches. There are three divisions of the Cenozoic: Paleogene, Neogene and Quaternary.


Paleogene Period

The Paleogene spans from the extinction of the non-avian dinosaurs, some 66 million years ago, to the dawn of the Neogene 23 million years ago. It features three epochs: Paleocene, Eocene and
Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but the ...
. The Paleocene Epoch began with the K–Pg extinction event, and the early part of the Paleocene saw the recovery of the Earth from that event. The continents began to take their modern shapes, but most continents (and India) remained separated from each other: Africa and Eurasia were separated by the Tethys Sea, and the Americas were separated by the strait of Panama, as the Isthmus of Panama had not yet formed. This epoch featured a general warming trend, and the earliest modern jungles expanded, eventually reaching the poles. The oceans were dominated by sharks, as the large reptiles that had once ruled had become extinct. Mammals diversified rapidly, but most remained small. The largest tetrapod carnivores during the Paleocene were reptiles, including crocodyliforms,
choristodera Choristodera (from the Greek χωριστός ''chōristos'' + δέρη ''dérē'', 'separated neck') is an extinct order of semiaquatic diapsid reptiles that ranged from the Middle Jurassic, or possibly Triassic, to the late Miocene (168 to 1 ...
ns, and snakes. ''
Titanoboa ''Titanoboa'' (; ) is an extinct genus of very large snakes that lived in what is now La Guajira in northeastern Colombia. They could grow up to , perhaps even long and reach a body mass of . This snake lived during the Middle to Late Paleoc ...
,'' the largest known snake, lived in South America during the Paleocene. The Eocene Epoch ranged from 56 million to 34 million years ago. In the early Eocene, most land mammals were small and living in cramped jungles, much like the Paleocene. Among them were early primates, whales and horses along with many other early forms of mammals. The climate was warm and humid, with little temperature gradient from pole to pole. In the Middle Eocene Epoch, the circum-Antarctic current between Australia and Antarctica formed, disrupting ocean currents worldwide, resulting in global cooling, and causing the jungles to shrink. More modern forms of mammals continued to diversify with the cooling climate even as more archaic forms died out. By the end of the Eocene, whales such as '' Basilosaurus'' had become fully aquatic. The late Eocene Epoch saw the rebirth of seasons, which caused the expansion of savanna-like areas with the earliest substantial grasslands. At the transition between the Eocene and Oligocene epochs there was a significant
extinction event An extinction event (also known as a mass extinction or biotic crisis) is a widespread and rapid decrease in the biodiversity on Earth. Such an event is identified by a sharp change in the diversity and abundance of multicellular organisms. I ...
, the cause of which is debated. The Oligocene Epoch spans from 34 million to 23 million years ago. The Oligocene was an important transitional period between the tropical world of the Eocene and more modern ecosystems. This period featured a global expansion of grass which led to many new species taking advantage, including the first elephants, cats, dogs, marsupials and many other species still prevalent today. Many other species of plants evolved during this epoch also, such as the evergreen trees. The long term cooling continued and seasonal rain patterns established. Mammals continued to grow larger. '' Paraceratherium'', one of the largest land mammals to ever live, evolved during this epoch, along with many other perissodactyls.


Neogene Period

The Neogene spans from 23.03 million to 2.58 million years ago. It features two epochs: the Miocene and the Pliocene. The Miocene spans from 23.03 million to 5.333 million years ago and is a period in which grass spread further across, effectively dominating a large portion of the world, diminishing forests in the process. Kelp forests evolved, leading to the evolution of new species, such as sea otters. During this time, perissodactyls thrived, and evolved into many different varieties. Alongside them were the apes, which evolved into 30 species. Overall, arid and mountainous land dominated most of the world, as did grazers. The Tethys Sea finally closed with the creation of the Arabian Peninsula and in its wake left the Black, Red, Mediterranean and Caspian Seas. This only increased aridity. Many new plants evolved, and 95% of modern seed plants evolved in the mid-Miocene. The Pliocene lasted from 5.333 million to 2.58 million years ago. The Pliocene featured dramatic climatic changes, which ultimately led to modern species and plants. The Mediterranean Sea dried up for hundreds of thousand years in the Messinian salinity crisis. Along with these major geological events, Africa saw the appearance of '' Australopithecus,'' the ancestor of '' Homo''. The isthmus of Panama formed, and animals migrated between North and South America, wreaking havoc on the local ecology. Climatic changes brought savannas that are still continuing to spread across the world, Indian monsoons, deserts in East Asia, and the beginnings of the Sahara desert. The Earth's continents and seas moved into their present shapes. The world map has not changed much since, save for changes brought about by the
glaciations A glacial period (alternatively glacial or glaciation) is an interval of time (thousands of years) within an ice age that is marked by colder temperatures and glacier advances. Interglacials, on the other hand, are periods of warmer climate betwe ...
of the Quaternary, such as the Great Lakes.


Quaternary Period

The Quaternary spans from 2.58 million years ago to present day, and is the shortest geological period in the Phanerozoic Eon. It features modern animals, and dramatic changes in the climate. It is divided into two epochs: the Pleistocene and the Holocene. The Pleistocene lasted from 2.58 million to 11,700 years ago. This epoch was marked by a series of glacial periods (ice ages) as a result of the cooling trend that started in the Mid-Eocene. There were numerous separate glaciation periods marked by the advance of ice caps as far south as 40 degrees N latitude in mountainous areas. Meanwhile, Africa experienced a trend of desiccation which resulted in the creation of the Sahara, Namib, and Kalahari deserts. Mammoths,
giant ground sloths ''Megatherium'' ( ; from Greek () 'great' + () 'beast') is an extinct genus of ground sloths endemic to South America that lived from the Early Pliocene through the end of the Pleistocene. It is best known for the elephant-sized type species ' ...
,
dire wolves The dire wolf (''Aenocyon dirus'' ) is an extinct canine. It is one of the most famous prehistoric carnivores in North America, along with its extinct competitor ''Smilodon''. The dire wolf lived in the Americas and eastern Asia during the L ...
,
sabre-toothed cat Machairodontinae is an extinct subfamily of carnivoran mammals of the family Felidae (true cats). They were found in Asia, Africa, North America, South America, and Europe from the Miocene to the Pleistocene, living from about 16 million until ...
s, and humans were common and widespread during the Pleistocene. As the Pleistocene drew to a close, a major extinction wiped out much of the world's
megafauna In terrestrial zoology, the megafauna (from Greek μέγας ''megas'' "large" and New Latin ''fauna'' "animal life") comprises the large or giant animals of an area, habitat, or geological period, extinct and/or extant. The most common threshold ...
, including non-''Homo sapiens'' human species such as '' Homo neanderthalensis''. All the continents were affected, but Africa was impacted to a lesser extent. That continent retained many large animals, such as elephants, rhinos, and hippos. The extent to which ''Homo Sapiens'' were involved in this extinction is debated. The Holocene began 11,700 years ago and lasts until the present day. All recorded history and "
Human history Human history, also called world history, is the narrative of humanity's past. It is understood and studied through anthropology, archaeology, genetics, and linguistics. Since the invention of writing, human history has been studied throug ...
" lies within the boundaries of the Holocene epoch. Human activity is blamed for an ongoing mass extinction that began roughly 10,000 years ago, though the species becoming extinct have only been recorded since the Industrial Revolution. This is sometimes referred to as the " Sixth Extinction". Hundreds of species have become extinct due to human activity since the Industrial Revolution.


Biodiversity

It has been demonstrated that changes in biodiversity through the Phanerozoic correlate much better with the hyperbolic model (widely used in demography and macrosociology) than with exponential and logistic models (traditionally used in population biology and extensively applied to fossil biodiversity as well). The latter models imply that changes in diversity are guided by a first-order positive feedback (more ancestors, more descendants) or a
negative feedback Negative feedback (or balancing feedback) occurs when some function (Mathematics), function of the output of a system, process, or mechanism is feedback, fed back in a manner that tends to reduce the fluctuations in the output, whether caused by ...
that arises from resource limitation, or both. The hyperbolic model implies a second-order positive feedback. The hyperbolic pattern of the
human population Humans (''Homo sapiens'') are the most abundant and widespread species of primate, characterized by bipedalism and exceptional cognitive skills due to a large and complex brain. This has enabled the development of advanced tools, culture, ...
growth arises from quadratic positive feedback, caused by the interaction of the population size and the rate of technological growth.See, e. g., The character of biodiversity growth in the Phanerozoic Eon can be similarly accounted for by a feedback between the diversity and community structure complexity. It has been suggested that the similarity between the curves of biodiversity and human population probably comes from the fact that both are derived from the superposition on the hyperbolic trend of cyclical and random dynamics.


See also

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Citations


General references

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External links


Phanerozoic (chronostratigraphy scale)
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