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''Bradysaurus'' was a large, early and common
pareiasaur Pareiasaurs (meaning "cheek lizards") are an extinct clade of large, herbivorous parareptiles. Members of the group were armoured with scutes which covered large areas of the body. They first appeared in southern Pangea during the Middle Permian, ...
, the fossils of which are known from the '' Tapinocephalus''
Assemblage Zone Biostratigraphy is the branch of stratigraphy which focuses on correlating and assigning relative ages of rock strata by using the fossil assemblages contained within them.Hine, Robert. “Biostratigraphy.” ''Oxford Reference: Dictionary of Bio ...
( Capitanian age) of the South African
Karoo The Karoo ( ; from the Afrikaans borrowing of the South Khoekhoe !Orakobab or Khoemana word ''ǃ’Aukarob'' "Hardveld") is a semi-desert natural region of South Africa. No exact definition of what constitutes the Karoo is available, so its ext ...
. Along with the similarly large dinocephalia, the bradysaurs constituted the herbivorous
megafauna In terrestrial zoology, the megafauna (from Greek μέγας ''megas'' "large" and New Latin ''fauna'' "animal life") comprises the large or giant animals of an area, habitat, or geological period, extinct and/or extant. The most common threshold ...
of the late Middle Permian Period. In life they were probably slow, clumsy and inoffensive animals, that had evolved a covering of armoured scutes to protect them against their predators, the gorgonopsians.


Description

''Bradysaurus'' was in length and half a tonne to a tonne in weight. The skull was large (about 42 to 48 centimeters long), broad and rounded at the front. It was coarsely sculptured and knobby, with the sutures between the bones not clearly visible. The marginal teeth were high-crowned, with only a few cusps, which is a primitive characteristic. The feet were short and broad, the phalangeal count being 2,3,3,3,2 on the fore-foot and 2,3,3,4,3 on the hind. The whole body is protected by dermal scutes, although these are not as thick or heavy as in more advanced forms.


Classification and species

''Bradysaurus'' is the only member of the subfamily Bradysaurinae. It is the most primitive known pareiasaur and can be considered a good ancestral type from which the others developed. Its large dimensions show that, even very early in their evolutionary history, these strange animals had already attained an optimal size. Even later, more advanced forms, like ''
Scutosaurus ''Scutosaurus'' ("shield lizard") is an extinct genus of pareiasaur parareptiles. Its genus name refers to large plates of armor scattered across its body. It was a large anapsid reptile that, unlike most reptiles, held its legs underneath its b ...
'', were no larger. The advantage of large size was to provide defense against predators and to maintain a stable body temperature ( gigantothermy).
Kuhn Kuhn is a surname of German origin. It may refer to the following: * Abraham Kuhn (banker) (1819–1892), German-American founder of Kuhn, Loeb & Co. * Abraham Kuhn (otolarynologist) (1838–1900), Alsatian otolaryngologist * Adam Kuhn (1741–181 ...
1969 lists no fewer than nine species for this genus, but this is certainly an excessive number. Boonstra 1969 distinguishes only four species on the basis of tooth structure, two of which Kuhn places in the genus ''Embrithosaurus''. The genera ''Brachypareia'', ''Bradysuchus'', ''Koalemasaurus'', and ''Platyoropha'' are synonyms of ''Bradysaurus''. ''B. baini'' (Seeley, 1892) is from the ''Tapinocephalus'' zone, Lower Beaufort Beds, Karoo basin, South Africa. This is the type species for the genus. The quadra-jugal region (cheek-bones) were only moderately developed. The snout was broad and rounded and there were 15 or 16 pairs of overlapping teeth in each jaw. This animal could be considered a generic early pareiasaur. According to Lee, 1997, the available material of ''B. baini'' lacks distinguishing autapomorphies or characteristics. ''B. seeleyi'' (Haughton and Boonstra, 1929) is from the ''Tapinocephalus'' zone, Lower Beaufort Beds, Karoo basin, South Africa. This is a less common form. Boonstra, 1969, considered this a valid species of ''Bradysaurus'' and Lee, 1997, considers this animal a sister group to more advanced pareiasaurs. ''B. seelyi'' seems to be closely related to ''Nochelesaurus'' and ''
Embrithosaurus ''Embrithosaurus'' was a pareiasaur from the Permian of South Africa. Description ''Embrithosaurus'' was in length and in weight. The skull is relatively deep and narrow. The body is lightly armoured with thin, smooth dermal scutes. Species * ...
''. In contrast to the more numerous but similarly sized ''B. baini'', the cheekbones were heavy and greatly enlarged. There were 19 or 20 pairs of strongly overlapping teeth on each jaw.


References

* Boonstra, L. D. 1969, "The Fauna of the Tapinoephalus Zone (Beaufort Beds of the Karoo)," Ann. S. Afr. Mus. 56 (1) 1-73, pp. 29–32 *
Edwin H. Colbert Edwin Harris "Ned" Colbert (September 28, 1905 – November 15, 2001)O'Connor, Anahad ''The New York Times'', November 25, 2001. was a distinguished American vertebrate paleontologist and prolific researcher and author. Born in Clarinda, Iowa, he ...
, 1965, The Age of Reptiles, The World Naturalist, Weidenfeld and Nicolson, London, pp. 52–3 * Barry Cox, R.J.G.Savage, Brian Gardiner,
Dougal Dixon Dougal Dixon (born 1 March 1947) is a Scottish geologist, palaeontologist, educator and author. Dixon has written well over a hundred books on geology and palaeontology, many of them for children, which have been credited with attracting many to ...
, 1988 ''Illustrated Encyclopaedia of Dinosaurs and Prehistoric Animals'' *
Carroll Lane Fenton Carroll Lane Fenton (February 12, 1900, Butler County, Iowa – November 16, 1969, New Brunswick, New Jersey) was a geologist, paleontologist, neoichnologist, and historian of science. Fenton was the author and illustrator of numerous books o ...
and
Mildred Adams Fenton Mildred Adams Fenton (November 14, 1899 – December 7, 1995) trained in paleontology and geology at the University of Iowa. She coauthored dozens of general science books with her husband, Carroll Lane Fenton, including ''Records of Evolution' ...
, 1958, ''The Fossil Book'', Doubleday & Co., Garden City, New York, p. 306 * Kuhn, O, 1969, Cotylosauria, part 6 of ''Handbuch der Palaoherpetologie'' (Encyclopedia of Palaeoherpetology), Gustav Fischer Verlag, Stuttgart & Portland, * Lee, MSY (1997), Pareiasaur phylogeny and the origin of turtles. ''Zool. J. Linnean Soc.'', 120: 197-280


External links


Bradysaurs
at Palaeos {{Taxonbar, from=Q137709 Pareiasaurs Permian reptiles of Africa Fossil taxa described in 1914 Capitanian genus first appearances Capitanian genus extinctions Prehistoric reptile genera