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'' Tyrannosaurus'' is one of the most iconic dinosaurs and is known from numerous specimens, some of which have individually acquired notability due to their scientific significance and media coverage.


Specimen data


''Manospondylus'': AMNH 3982

The first-named fossil specimen which can be attributed to ''Tyrannosaurus rex'' consists of two partial vertebrae (one of which has been lost) found by
Edward Drinker Cope Edward Drinker Cope (July 28, 1840 – April 12, 1897) was an American zoologist, paleontologist, comparative anatomist, herpetologist, and ichthyologist. Born to a wealthy Quaker family, Cope distinguished himself as a child prodigy interested ...
in 1892. Cope believed that they belonged to an "agathaumid" (
ceratopsid Ceratopsidae (sometimes spelled Ceratopidae) is a family of ceratopsian dinosaurs including ''Triceratops'', ''Centrosaurus'', and ''Styracosaurus''. All known species were quadrupedal herbivores from the Upper Cretaceous. All but one species are k ...
) dinosaur, and named them ''Manospondylus gigas'', meaning "giant porous vertebra" in reference to the numerous openings for blood vessels he found in the bone. The ''M. gigas'' remains were later identified as those of a theropod rather than a ceratopsid, and H.F. Osborn recognized the similarity between ''M. gigas'' and ''Tyrannosaurus rex'' as early as 1917. However, due to the fragmentary nature of the ''Manospondylus'' vertebrae, Osborn did not synonymize the two genera.


''Dynamosaurus'': BMNH R7994

The holotype of ''Tyrannosaurus rex'', a partial skull and skeleton originally called AMNH 973 (AMNH stands for
American Museum of Natural History The American Museum of Natural History (abbreviated as AMNH) is a natural history museum on the Upper West Side of Manhattan in New York City. In Theodore Roosevelt Park, across the street from Central Park, the museum complex comprises 26 inter ...
), was discovered in the U.S. state of Montana in 1902 and excavated over the next three years. Another specimen (AMNH 5866), found in Wyoming in 1900, was described in the same paper under the name ''Dynamosaurus imperiosus''. At the time of their initial description and naming, these specimens had not been fully prepared and the type specimen of ''T. rex'' had not even been fully recovered. In 1906, after further preparation and examination, Henry Fairfield Osborn recognized both skeletons as belonging to the same species. Because the name ''Tyrannosaurus rex'' had appeared just one page earlier than ''Dynamosaurus'' in Osborn's 1905 work, it was considered the older name and has been used since. Had it not been for page order, ''Dynamosaurus'' would have become the official name.


Holotype: CM 9380

CM 9380 is the
type Type may refer to: Science and technology Computing * Typing, producing text via a keyboard, typewriter, etc. * Data type, collection of values used for computations. * File type * TYPE (DOS command), a command to display contents of a file. * Ty ...
specimen used to describe '' Tyrannosaurus rex''. Fragments of (then) AMNH 973 were first found in 1902 by
Barnum Brown Barnum Brown (February 12, 1873 – February 5, 1963), commonly referred to as Mr. Bones, was an American paleontologist. Named after the circus showman P. T. Barnum, he discovered the first documented remains of ''Tyrannosaurus'' during a career ...
, assistant curator of the American Museum of Natural History and a famous paleontologist in his own right. He forwarded news of it to Osborn; it would be three years before they found the rest of it. In 1905 when the type was described by Osborn, previous knowledge of dinosaur predators at the time were based on Jurassic carnosaurs, so the short fore-arms of the ''Tyrannosaurus'' were treated with extreme caution, with suspicion that bones of a smaller theropod had become jumbled with the remains of the bigger fossil. Following the 1941 entry of the United States into World War II, the holotype was sold to the Carnegie Museum of Natural History in Pittsburgh for protection against possible bombing raids. The specimen, now labeled CM 9380, is still mounted in Pittsburgh, at first with the tail acting as a tripod in the old-fashioned kangaroo pose. It has since received a modernization of its posture (mounted by Phil Fraley and crew) and can now be found balancing with tail outstretched. Along with a more lifelike posture, the specimen also now includes a composite reconstruction of the skull by Michael Holland. It has been reconstructed in recent years, it measured an estimated 11.9 meters in length and an estimated weight of 7.4–14.6 metric tonnes, 9.1 metric tonnes being the average estimate in that study, although most earlier studies have suggested lower weight figures.


AMNH 5027

AMNH 5027 was discovered and excavated in 1908 by
Barnum Brown Barnum Brown (February 12, 1873 – February 5, 1963), commonly referred to as Mr. Bones, was an American paleontologist. Named after the circus showman P. T. Barnum, he discovered the first documented remains of ''Tyrannosaurus'' during a career ...
in Montana, and described by Osborn in 1912 and 1916. At the time of discovery, a complete
cervical In anatomy, cervical is an adjective that has two meanings: # of or pertaining to any neck. # of or pertaining to the female cervix: i.e., the ''neck'' of the uterus. *Commonly used medical phrases involving the neck are **cervical collar **cervic ...
(neck vertebrae) series for ''Tyrannosaurus'' was not previously known, so it was this specimen that brought the short, stocky tyrannosaur neck to light. Compared to later specimens ( BMNH R7994 and FMNH PR2081, for instance) the cervical series of AMNH 5027 is much more gracile, so with later discoveries the distinction between tyrannosaurid necks and the necks of carnosaurs became more obvious. This specimen also provided the first complete skull of ''Tyrannosaurus rex''. In total, Brown found five partial ''Tyrannosaurus'' skeletons. The skeleton of this specimen was used as the iconic symbol for the
Jurassic Park film series ''Jurassic Park'', later also referred to as ''Jurassic World'', is an American science fiction media franchise created by Michael Crichton and centered on a disastrous attempt to create a theme park of De-extinction#Cloning, cloned dinosaurs. ...
. Osborn planned to mount the similarly sized AMNH 5027 and AMNH 973 together in dynamic poses. Designed by E.S. Christman, the scene was to depict a rearing ''Tyrannosaurus'' (AMNH 5027) snapping at another cowering one (AMNH 973), as they fought over the remains of a hadrosaur, described at the time as '' Trachodon''. However, technical difficulties prevented the mount from being executed. One obvious problem was that the Cretaceous Dinosaur Hall was too small to accommodate this dramatic display, and AMNH 5027 was already mounted by itself as the central attraction of the hall. The forearms of ''Tyrannosaurus'' were not well documented and the hands were unknown, so for the sake of the display, the forearms of AMNH 5027 were given three fingers, based on the forelimbs of '' Allosaurus'' (the more allosaur-like arms were replaced several years later when better fossils of tyrannosaurid arms were found). The mount retained a rearing pose similar to the initial proposal. By the 1980s it was generally accepted that such a pose would have been anatomically impossible in life, and the skeleton was re-mounted in a more accurate, horizontal pose during a renovation of the museum's dinosaur halls in the early 1990s. The mount can still be seen on display on the fourth floor of the American Museum. The American Museum of Natural History features AMNH 5027 in its famed Hall of Saurischian Dinosaurs to this day.


''Nanotyrannus'': CMNH 7541

A small but nearly complete skull of ''Nanotyrannus lancensis'', frequently considered to be a juvenile ''T. rex'', was recovered from Montana in 1942. This skull, Cleveland Museum of Natural History (CMNH) 7541, measures in length and was originally classified as a species of '' Gorgosaurus'' (''G. lancensis'') by
Charles W. Gilmore Charles Whitney Gilmore (March 11, 1874 – September 27, 1945) was an American paleontologist who gained renown in the early 20th century for his work on vertebrate fossils during his career at the United States National Museum (now the N ...
in 1946. In 1988, the specimen was re-described by
Robert T. Bakker Robert Thomas Bakker (born March 24, 1945) is an American paleontologist who helped reshape modern theories about dinosaurs, particularly by adding support to the theory that some dinosaurs were endothermic (warm-blooded). Along with his mentor J ...
, Phil Currie, and Michael Williams, then the curator of paleontology at the Cleveland Museum of Natural History, where the original specimen was housed and is now on display. Their initial research indicated that the skull bones were fused, and that it therefore represented an adult specimen. In light of this, Bakker and colleagues assigned the skull to a new genus, named ''Nanotyrannus'' for its apparently small adult size. The specimen is estimated to have been around long when it died. However, a detailed analysis of the specimen by Thomas Carr in 1999 showed that the specimen was, in fact, a juvenile, leading Carr and many other paleontologists to consider it a juvenile specimen of ''T. rex''. The current classification of CMNH 7541 is not universal.


LACM 23844

In 1966, a crew working for the Natural History Museum of Los Angeles County under the direction of Harley Garbani discovered another ''T. rex'' (LACM 23844) which included most of the skull of a very large, mature animal. When it was put on display in Los Angeles, LACM 23844 was the largest ''T. rex'' skull on exhibit anywhere.Horner, J.R. & Lessem, D. 1993. ''The Complete T. rex'' New York: Simon & Schuster. 238pp.


"Black Beauty": RTMP 81.6.1

"Black Beauty" (specimen number RTMP 81.6.1) is a well-preserved fossil of ''Tyrannosaurus rex''. The nickname stems from the apparent shiny dark color of the fossil bones, which occurred during fossilization by the presence of minerals in the surrounding rock; it was the first ''Tyrannosaurus rex'' specimen to receive a nickname, beginning a trend that continues with most major ''T. rex'' finds. Black Beauty was found in 1980 by a high school student, Jeff Baker, while on a fishing trip with a friend in the region of the Crowsnest Pass, Alberta. A large bone was found in the
riverbank Riverbank or river bank may refer to: * Bank (geography), the bank of a river Places * Riverbank, California *Riverbank, former name of Bryte, California Enterprises and organizations *Riverbank Academy, a special school in Coventry, England *Ri ...
and shown to their teacher. Soon afterward, the Royal Tyrrell Museum was contacted, and excavation of the sandstone matrix surrounding the fossils began in 1982. The dig site where the fossil was found is located at approximately 114^\circ ~ \textrm near the confluence of the Crowsnest and Willow Rivers, and consisted of rock belonging to the
Willow Creek Formation The Willow Creek Formation is a stratigraphic unit of Late Cretaceous to Early Paleocene age in the Western Canada Sedimentary Basin of southwestern Alberta. It was first described by George Mercer Dawson in 1883 along the Willow Creek, a tribut ...
. The specimen is housed in the Royal Tyrrell Museum in Drumheller, Alberta, Canada. In 2009, a paper by
Jack Horner Jack Horner may refer to: *''Little Jack Horner'', a nursery rhyme People * Jack Horner (baseball) (1863–1910), American professional baseball player *Jack Horner (journalist) (1912–2005), Gordon John Horner, Minnesota sportscaster * Jack B. H ...
and colleagues illustrated the concept of parasitic infections in dinosaurs by analysing the lesions found on the cranial bones of Black Beauty. The specimen has been used to study comparative morphology between tyrannosaurids and ''Tyrannosaurus'' individuals. Replicas of Black Beauty have been shown in some exhibitions and museums, including both simple skull montages and complete skeletons. Casts are on display in museums around the world.


"Stan": BHI 3033

Stan is the nickname given to a fossil found in Hell Creek Formation, South Dakota, close to Buffalo in 1987 by
Stan Sacrison Stan or STAN may refer to: People * Stan (given name), a list of people with the given name ** Stan Laurel (1890–1965), English comic actor, part of duo Laurel and Hardy * Stan (surname), a Romanian surname * Stan! (born 1964), American author, ...
, who also discovered the '' Tyrannosaurus'' specimen nicknamed "Duffy". The original fossils are now housed at
Black Hills Institute of Geological Research The Black Hills Institute of Geological Research, Inc. (BHI) is a private corporation specializing in the excavation and preparation of fossils, as well as the sale of both original fossil material and museum-quality replicas. Founded in 1974 and b ...
, Inc. center. It is a well known specimen, and one of the most complete, with 199 bones recovered. About 30 casts of the original fossil have been sold worldwide, each for a price of about $100,000. Stan's skeleton was auctioned for $31.8 million in a 2020 Christie's New York sale, making it a record-breaking dinosaur sale, with the buyer eventually being revealed as the under construction Natural History Museum Abu Dhabi in the Saadiyat Cultural District. Like many other fossils of ''Tyrannosaurus rex'', the skeleton of Stan shows many broken and healed bones. These include broken
rib In vertebrate anatomy, ribs ( la, costae) are the long curved bones which form the rib cage, part of the axial skeleton. In most tetrapods, ribs surround the chest, enabling the lungs to expand and thus facilitate breathing by expanding the ches ...
s and damages in the skull. One of the most prominent injuries are in the neck and the skull. A piece of bone is missing at the rear, and the skull also bears a hole 1 inch wide, probably made by another '' Tyrannosaurus''. Also, two of the
cervical vertebrae In tetrapods, cervical vertebrae (singular: vertebra) are the vertebrae of the neck, immediately below the skull. Truncal vertebrae (divided into thoracic and lumbar vertebrae in mammals) lie caudal (toward the tail) of cervical vertebrae. In ...
are fused, and another has additional bone growth. This could have been caused by another ''Tyrannosaurus'' bite. The bite marks are healed, indicating that Stan survived the wounds. Stan could also have been infected by ''
Trichomonas ''Trichomonas'' is a genus of anaerobic excavate parasites of vertebrates. It was first discovered by Alfred François Donné in 1836 when he found these parasites in the pus of a patient suffering from vaginitis, an inflammation of the vagina. ...
''-like parasites.


"Wankel Rex": MOR 555

In 1988, local rancher Kathy Wankel discovered another ''Tyrannosaurus rex'' in Hell Creek sediments on an island in the
Charles M. Russell National Wildlife Refuge The Charles M. Russell National Wildlife Refuge (abbreviated as the CMR NWR) is a National Wildlife Refuge in the U.S. state of Montana on the Missouri River. The refuge surrounds Fort Peck Reservoir and is in size.U.S. Fish and Wildlife Servi ...
of Montana. This specimen was excavated by a team from the
Museum of the Rockies Museum of the Rockies is a museum in Bozeman, Montana. Originally affiliated with Montana State University in Bozeman, and now also, the Smithsonian Institution, the museum is largely known for its paleontological collections. The Museum houses ...
led by paleontologist
Jack Horner Jack Horner may refer to: *''Little Jack Horner'', a nursery rhyme People * Jack Horner (baseball) (1863–1910), American professional baseball player *Jack Horner (journalist) (1912–2005), Gordon John Horner, Minnesota sportscaster * Jack B. H ...
, with assistance from the U.S. Army Corps of Engineers. The specimen, given the number MOR 555 but informally called the "Wankel rex," includes approximately 46 percent of the skeleton, including the skull, as well as what at the time was the first complete ''T. rex'' forelimb. It has an estimated length of around and a weight between and in newer figures. It is estimated that the "Wankel rex" was 18 years old when it died, an adult but not completely grown. The "Wankel rex" was also one of the first fossil dinosaur skeletons studied to see if biological molecules still existed within the fossilized bones. Doctoral candidate Mary Schweitzer found heme, a biological form of iron that makes up hemoglobin (the red pigment in blood).Schontzler, Gail. "Montana ''T. rex'' Heading to Smithsonian." ''Bozeman Daily Chronicle.'' 28 June 2013.
Accessed 2013-06-28.
It was long on exhibit at the Museum of the Rockies in
Bozeman, Montana Bozeman is a city and the county seat of Gallatin County, Montana, United States. Located in southwest Montana, the 2020 census put Bozeman's population at 53,293, making it the fourth-largest city in Montana. It is the principal city of th ...
. In June 2013, the Corps loaned the specimen to the National Museum of Natural History, Smithsonian Institution museum in Washington, D.C., for 50 years. (The Museum of the Rockies continues to display a cast reconstruction of the skull by Michael Holland). The specimen went on temporary display on National Fossil Day, 16 October 2013, and was exhibited until the museum's dinosaur hall exhibit closed for renovation in the spring of 2014. The skeleton, named "The Nation's T. rex" became the centerpiece of the dinosaur hall when it reopened in 2019. Casts of MOR 555 are on display at the
National Museum of Scotland The National Museum of Scotland in Edinburgh, Scotland, was formed in 2006 with the merger of the new Museum of Scotland, with collections relating to Scottish antiquities, culture and history, and the adjacent Royal Scottish Museum (opened in ...
, the
Australian Fossil and Mineral Museum The Australian Fossil and Mineral Museum - Home of the Somerville Collection is located in the city of Bathurst in regional New South Wales Australia and was opened in July 2004. The collection is housed in a group of heritage buildings, the o ...
, and the University of California Museum of Paleontology. A bronze cast of the specimen, known as "Big Mike", stands outside the Museum of the Rockies.


"Sue": FMNH PR2081

Susan Hendrickson of the Black Hills Institute discovered the best-preserved ''Tyrannosaurus'' currently known, in the Hell Creek Formation near
Faith, South Dakota Faith is a city in the northeastern corner of Meade County, South Dakota, United States. The population was 367 at the 2020 census. The most complete ''Tyrannosaurus rex'' skeleton known, Sue, was discovered approximately 15 miles northeas ...
, on 12 August 1990. About 90% of the skeleton was recovered, allowing the first complete description of a ''Tyrannosaurus rex'' skeleton. This specimen, named "Sue" in honor of its discoverer, soon became embroiled in a legal battle over its ownership. The owner of the land the fossil was found, Maurice Williams, as well the
Sioux Tribe The Sioux or Oceti Sakowin (; Dakota: /otʃʰeːtʰi ʃakoːwĩ/) are groups of Native American tribes and First Nations peoples in North America. The modern Sioux consist of two major divisions based on language divisions: the Dakota a ...
he belonged to, claimed ownership, the Institute had considered itself to have. In 1997, the suit was settled in favor of Williams and the fossil was returned to Williams' ownership. Williams quickly offered up "Sue" for auction by Sotheby's in New York, where it was sold to the
Field Museum of Natural History The Field Museum of Natural History (FMNH), also known as The Field Museum, is a natural history museum in Chicago, Illinois, and is one of the largest such museums in the world. The museum is popular for the size and quality of its educational ...
in Chicago for US$8.4 million—the highest price ever paid for a fossil before being surpassed by
Stan Stan or STAN may refer to: People * Stan (given name), a list of people with the given name ** Stan Laurel (1890–1965), English comic actor, part of duo Laurel and Hardy * Stan (surname), a Romanian surname * Stan! (born 1964), American author ...
. Sue has a length of , stands tall at the hips, and according to the most recent studies estimated to have weighed between 8.4 and 14 metric tons when alive. It has been hypothesised that Sue's impressive size may have been achieved due to a prolonged
ontogenic Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the egg to adult. The term can also be used to refer to the stu ...
development, since it is the third oldest ''Tyrannosaurus'' known. Sue's age at the time of death was estimated by Peter Mackovicky and the University of Florida to be 28 years old, over 6–10 years older than most big ''Tyrannosaurus'' specimens, like MOR 555, AMNH 5027 or BHI 3033. The only known specimen of ''T. rex'' that is older than Sue is Trix.Koen Stein, paleontologist at the Free University of Brussels ( Vrije Universiteit Brussel)
"Oldest T. rex Trix on view in Leiden, VUB researchers established the dinosaur's age", ''VUB Today'', 23 January 2017
/ref> Preparation of "Sue" (FMNH PR2081) was completed at the Field Museum and the skeleton was placed on exhibit on 17 May 2000.
by Neal Larson. Black Hills Institute of Geological Research, Inc. 19 May 2000. Retrieved 16 July 2006.
"Sue at the Field Museum"
Field Museum of Natural History. 9 September 2005. Retrieved 16 July 2006.


"Peck's Rex": MOR980

Peck's Rex (also known as "Peckrex", "Rigby's rex" and ''Tyrannosaurus'' "imperator") is the nickname given to a fossil specimen found in Montana in 1997. The discovery was made by Louis E. Tremblay on 4 July 1997 working under the supervision of
J. Keith Rigby Jr. ''J. The Jewish News of Northern California'', formerly known as ''Jweekly'', is a weekly print newspaper in Northern California, with its online edition updated daily. It is owned and operated by San Francisco Jewish Community Publications In ...
who led the excavation and bone preparation. The fossil got its nickname after Fort Peck, which is located close to the place of discovery. The fossil of Peck's Rex is relatively well preserved. The skeleton includes a relatively complete skull with jaws, multiple vertebrae of the back and tail, a well preserved
gastralium Gastralia (singular gastralium) are dermal bones found in the ventral body wall of modern crocodilians and tuatara, and many prehistoric tetrapods. They are found between the sternum and pelvis, and do not articulate with the vertebrae. In these ...
, and
hipbone The hip bone (os coxae, innominate bone, pelvic bone or coxal bone) is a large flat bone, constricted in the center and expanded above and below. In some vertebrates (including humans before puberty) it is composed of three parts: the ilium, ischi ...
with complete
ischium The ischium () form ...
and pubis. The left hindleg is relatively complete with a femur, missing only some toe bones. The forelimbs include the scapula and furcula, both humeri and right hand
phalanges The phalanges (singular: ''phalanx'' ) are digital bones in the hands and feet of most vertebrates. In primates, the thumbs and big toes have two phalanges while the other digits have three phalanges. The phalanges are classed as long bones. ...
, as well as metacarpal III. Peck's Rex has been the subject of research regarding parasitic infections in dinosaurs. The forelimbs of Peck's Rex have also been studied as they show evidence of use. This evidence includes the construction of metacarpal III, as well as repeated fractures in the furcula—possibly caused by heavy loads or pressure (Carpenter and Lipkin, 2005). Peck's Rex real fossilized bones are exhibited at The Museum of the Rockies as part of a full body mount completed with plaster replacing the missing bones, and was nicknamed Montana's Rex. This was made since their old main ''T. rex'' complete real skeleton; Wankel Rex (now nicknamed Nation's Rex) was sent to Washington DC Smithsonian's museum to occupy a central part in the museum's dinosaur hall, featuring a dynamic mount showing the apex predator devouring a ''Triceratops'' carcass. It has been stated by Pete Makovicky, the Chicago museum's lead curator of dinosaurs, that this specimen is in the same size range as "Sue" and "Scotty".


"Bucky": TCM 2001.90.1

Bucky is a fossil of a juvenile specimen on display at The Children's Museum of Indianapolis in
Indianapolis Indianapolis (), colloquially known as Indy, is the state capital and most populous city of the U.S. state of Indiana and the seat of Marion County. According to the U.S. Census Bureau, the consolidated population of Indianapolis and Marion ...
, Indiana. It is the first juvenile ''Tyrannosaurus'' ever placed on permanent exhibit in a museum. The dinosaur remains were found in 1998 in the Hell Creek Formation near the town of
Faith, South Dakota Faith is a city in the northeastern corner of Meade County, South Dakota, United States. The population was 367 at the 2020 census. The most complete ''Tyrannosaurus rex'' skeleton known, Sue, was discovered approximately 15 miles northeas ...
. The skeleton, transported by water, ended up in a low shallow valley along with bones from an ''
Edmontosaurus ''Edmontosaurus'' ( ) (meaning "lizard from Edmonton") is a genus of hadrosaurid (duck-billed) dinosaur. It contains two known species: ''Edmontosaurus regalis'' and ''Edmontosaurus annectens''. Fossils of ''E. regalis'' have been found in rocks ...
'' and '' Triceratops''. It was discovered by rancher and cowboy Bucky Derflinger. Bucky was well preserved and easily prepared by the Black Hills Institute in South Dakota. Part of The Children's Museum of Indianapolis Dinosphere exhibit, Bucky is displayed along with Stan, an adult ''Tyrannosaurus'', in a hunting scene. Both dinosaurs are attacking a ''Triceratops'' specimen known as "Kelsey". Bucky is one of the few dinosaur fossils found with a furcula; Bucky's furcula was the first one found for the genus ''Tyrannosaurus.'' Bucky also has a nearly complete set of gastralia, or belly ribs, and an ulna, or lower arm bone. As of now, 101 bones, or about 34% of Bucky's skeleton, has been discovered and verified. Bucky is the sixth-most complete ''Tyrannosaurus rex'' out of more than 40 that have been discovered.


"Jane": BMRP 2002.4.1

Jane is a fossil specimen of small tyrannosaurid dinosaur, officially known as BMRP 2002.4.1, discovered in the Hell Creek Formation in southern Montana in 2001. Despite having a typically female name, Jane's sex is unknown—the specimen was named after Burpee Museum benefactor Jane Solem. The specimen was found in the summer of 2001 by Carol Tuck and Bill Harrison on an expedition led by Burpee Museum curator Michael Henderson. After four years of preparation, Jane was put on display at
Rockford, Illinois Rockford is a city in Winnebago County, Illinois, located in the far northern part of the state. Situated on the banks of the Rock River, Rockford is the county seat of Winnebago County (a small portion of the city is located in Ogle County). ...
'
Burpee Museum of Natural History The Burpee Museum of Natural History is located along the Rock River in downtown Rockford, Illinois, United States, at 737 North Main Street. Museum history The museum was created as a Works Progress Administration project. It was established ...
as the centerpiece of an exhibit called "Jane: Diary of a Dinosaur." The Jane specimen has been central to the debate regarding the validity of the proposed tyrannosaurid genus ''
Nanotyrannus ''Tyrannosaurus'' is a genus of large theropod dinosaur. The species ''Tyrannosaurus rex'' (''rex'' meaning "king" in Latin), often called ''T. rex'' or colloquially ''T-Rex'', is one of the best represented theropods. ''Tyrannosaurus'' live ...
''. However, the Jane material has yet to be properly studied and described by scientists. Although Larson (2013) saw Jane as more identical to CMNH 7541 and LACM 28471 than to adult ''T. rex'' in having a higher tooth count, large pneumatic foramen on the center of the quadratojugal, T-shaped postorbital, and fused shoulder blade and pelvis, Yun (2015) concurred with the opinion of most workers that ''Nanotyrannus'' is a juvenile ''T. rex'', noting that a juvenile specimen of '' Tarbosaurus'' described by Tsuihiji ''et al.'' (2011) also has a T-shaped postorbital. Paleontologists who support the theory that Jane represents a juvenile believe the tyrannosaur was approximately 11 years old at its time of death, and its fully restored skeleton measured long, a bit more than half as long as the largest-known complete ''T. rex'' specimen, nicknamed " Sue," which measures long. According to Hutchinson ''et al.'' (2011), the weight of the Jane specimen in life was probably between and , being the average estimate.


"B-rex": MOR 1125

This specimen was found in the lower portion of the Hell Creek Formation near
Fort Peck Lake Fort Peck Lake, or Lake Fort Peck, is a major reservoir in Montana, formed by the Fort Peck Dam on the Missouri River. The lake lies in the eastern prairie region of Montana approximately east of Great Falls and north of Billings, reaching int ...
in the Charles M. Russell National Wildlife Refuge in Garfield County, Montana. Its discoverer was Bob Harmon, a fossil preparator for the Museum of the Rockies, and was nicknamed the "B-rex" (or "Bob-rex") in honor of Harmon. The specimen was discovered in 2000, and excavated by MOR from 2001 to 2003. Although only 37 percent of the skeleton was present, this included almost all of the skull (although the skull was nearly completely disarticulated). The specimen also includes several cervical, dorsal, sacral, and caudal vertebrae; several chevrons; some cervical and dorsal ribs; left scapula and coracoid; the furcula; the left ulna; both femora, tibiae, and ulnae; the right
calcaneum In humans and many other primates, the calcaneus (; from the Latin ''calcaneus'' or ''calcaneum'', meaning heel) or heel bone is a bone of the tarsus of the foot which constitutes the heel. In some other animals, it is the point of the hock. St ...
; right astragalus; and a number of
pes Pes (Latin for "foot") or the acronym PES may refer to: Pes * Pes (unit), a Roman unit of length measurement roughly corresponding with a foot * Pes or podatus, a * Pes (rural locality), several rural localities in Russia * Pes (river), a river ...
phalanges The phalanges (singular: ''phalanx'' ) are digital bones in the hands and feet of most vertebrates. In primates, the thumbs and big toes have two phalanges while the other digits have three phalanges. The phalanges are classed as long bones. ...
. In the March 2005 '' Science magazine'',
Mary Higby Schweitzer Mary Higby Schweitzer is an American paleontologist at North Carolina State University, who led the groups that discovered the remains of blood cells in dinosaur fossils and later discovered soft tissue remains in the ''Tyrannosaurus rex'' specime ...
of
North Carolina State University North Carolina State University (NC State) is a public land-grant research university in Raleigh, North Carolina. Founded in 1887 and part of the University of North Carolina system, it is the largest university in the Carolinas. The universit ...
and colleagues announced the recovery of soft tissue from the marrow cavity of a fossilized femur belong to B-Rex. Flexible, bifurcating blood vessels and fibrous but elastic bone matrix tissue were recognized. In addition, microstructures resembling
blood cell A blood cell, also called a hematopoietic cell, hemocyte, or hematocyte, is a cell produced through hematopoiesis and found mainly in the blood. Major types of blood cells include red blood cells (erythrocytes), white blood cells (leukocytes), ...
s were found inside the matrix and vessels. The structures bear resemblance to ostrich blood cells and vessels. However, since an unknown process distinct from normal fossilization seems to have preserved the material, the researchers are being careful not to claim that it is original material from the dinosaur. Paleontologist Thomas Kaye of the University of Washington in Seattle hypothesized that the soft-tissue is
permineralized Permineralization is a process of fossilization of bones and tissues in which mineral deposits form internal casts of organisms. Carried by water, these minerals fill the spaces within organic tissue. Because of the nature of the casts, perminera ...
biofilm created by bacteria while digesting and breaking down the original specimen. He has discovered this to be true in many specimens from the same area. In 2016, it was finally confirmed by Mary Higby Schweitzer and Lindsay Zanno ''et al'' that the soft tissue was medullary bone tissue, like that in modern birds when they are readying to lay eggs. This confirmed the identity of the ''Tyrannosaurus'' MOR 1125 as a female.


"Samson"

A ''T. rex'' specimen was discovered on private land in Harding County, South Dakota, once in 1981 by Michael and Dee Zimmerschied, and again on 4 October 1992 (Alan and Robert Detrich re-discovered Samson after it was originally found and deemed by paleontologists that several bones had washed in and there was nothing left). It was shortly after that when Fred Nuss and Candace Nuss of Nuss Fossils with the Detrich brothers found the most complete and undistorted ''Tyrannosaurus rex'' skull ever discovered. Following the sale of "Sue," another ''Tyrannosaurus rex'' skeleton was, the specimen was put up for auction on eBay in 2000 under the name of "Z-rex", with an asking price of over US$8 million. It failed to sell online but was purchased for an undisclosed price in 2001 by British millionaire Graham Ferguson Lacey, who renamed the skeleton "Samson" after the
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figure of the same name. It was prepared by the Carnegie Museum starting in May 2004."Carnegie Museum digs into controversial, but promising T-rex skull"
by Byron Spice. ''Pittsburgh Post-Gazette''. 12 May 2004. Retrieved 16 July 2006.
After preparation was complete in March 2006, the specimen was returned to Lacey.
by Dan Lagiovane. Carnegie Museum of Natural History
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2 March 2006. Retrieved 30 December 2010.
It, along with some other dinosaur skeletons, was sold again at auction on 3 October 2009. Samson measured , only slightly shorter than Sue.


"Baby Bob"

On 7 July 2013, fossil hunter Robert Detrich of Wichita, Kansas, unearthed the remains of what is believed to be a 4-year-old ''Tyrannosaurus rex''. Detrich unearthed the fossil dubbed "Baby Bob" in a fossil-rich area near the Eastern Montana town of Jordan. Its femur measures about 25 inches, and if all the preliminary data pans out, that would make it among the smallest ''T. rex'' specimens ever found. Baby Bob has been fully excavated, although it will take another year to clean. Detrich said the skull, which is about 75 percent complete, and most of the major skeletal elements were found strewn across a flood plain, although very few vertebra and ribs were found.


"Scotty": RSM P2523.8

" Scotty", cataloged as RSM P2523.8, was discovered in Saskatchewan, Canada in 1991. Since its discovery and extensive subsequent study, "Scotty" has been referred to as the largest ''T. rex'' ever discovered in the world, the largest of any dinosaur discovered in Canada, and as one of the oldest and most complete fossils of its kind at more than 70% bulk. "Scotty" resides at the Royal Saskatchewan Museum's T. rex Discovery Centre in Eastend, SK, Canada. In May 2019, a second mount was erected at the Royal Saskatchewan Museum in Regina, where the exhibit reflects the recent discoveries about the fossil. "Scotty" was discovered by Robert Gebhardt, a high school principal from Eastend, SK who accompanied palaeontologists from the Royal Saskatchewan Museum on a prospective expedition into the Frenchman Formation in southwestern Saskatchewan on 16 August 1991. It wasn't until June 1994 that the Royal Saskatchewan Museum was able to begin the excavation, which was led and overseen by the Museum's Ron Borden, as well as resident paleontologists Tim Tokaryk and John Storer who were with Gebhardt when he uncovered the first fossils. The bones were deeply packed in dense, iron-laden sandstone, which took more than twenty years for the team to fully remove, excavate, and assemble the majority of the skeleton, with additional trips being made to the site to retrieve smaller bones and teeth. The entire process of excavating the skeleton was also slowed down by its considerable size. "Scotty" is reported to be long and weighed an estimated . Despite it not being a complete fossil, paleontologists were able to create the estimation for the weight and length through measurements of important weight bearing bones such as the femur, hip, and shoulder bones that have all been measured to be larger and thicker with "Scotty" than the corresponding bones with "Sue". Going from the latest study "Scotty" exceeds "Sue" in 84.6% of the published measurements. While the reported measurements and weight for "Scotty" are larger than those of "Sue", some scientists posit that the two fossils are too close in size to officially declare "Scotty" the largest. Like other ''T. rex'' fossils, "Scotty" shows signs of trichomoniasis, a parasitic infection in the jaw that left visible holes in the bone and was unique to this specific species of dinosaur. Additionally, a broken and healed rib on its right side, broken tail vertebra, as well as a hole near the eye socket are possibly the result of another ''T. rex'' attack. Other abnormalities, such as impacted teeth, suggest that "Scotty" was not only bitten, but also bit other animals.


"Tristan"

Commercial paleontologist Craig Pfister discovered the specimen in the lower Hell Creek Formation in Carter County, Montana, in 2010. Its excavation and preservation lasted four years. It was later sold to Danish-born investment banker Niels Nielsen, who loaned the specimen to the Museum für Naturkunde in Berlin, Germany, for research and exhibition. It has been on display at Museum für Naturkunde between 2015 and 2020, moving to the
Natural History Museum of Denmark The Natural History Museum of Denmark ( da, Statens Naturhistoriske Museum) is a natural history museum located in Copenhagen, Denmark. It was created as a 1 January 2004 merger of Copenhagen's Zoological Museum, Geological Museum, Botanical Muse ...
for one year, and expected back in 2021.Back in 202
''Back in 2021 - Tristan''
accessed 13.2.2020
Nielsen and his friend Jens Jensen named the specimen Tristan-Otto (short: Tristan) for their sons. The Museum für Naturkunde Berlin lists it under specimen number MB.R.91216. Several European museums have ''Tyrannosaurus'' casts (replicas) or parts, but Tristan is one of only two original skeletons on display in the continent (the other is "Trix" in the Netherlands). The matte-black fossilised skeleton is about long and tall at the hips. Tristan is among the most complete known ''Tyrannosaurus'' skeletons: It was re-assembled from about 300 separate parts, 170 of which are original (including 98% of the skull and all the teeth), the rest reproductions. It is estimated to have died when about 20 years old and it was in poor health, having several bone fractures, bite marks to the skull and signs of disease in the jaw. The disease present in Tristan's jaw was suggested to be a case of tumefactive osteomyelitis.


"Thomas"

From 2003 to 2005, Thomas was excavated by NHMLA paleontologists in southeastern Montana. At 17 years old, long and nearly , it is estimated to be a 70% complete specimen. Thomas is mounted in a "growth series" with the youngest-known ''Tyrannosaurus rex'' fossil, a two-year-old, specimen, and a 13-year-old, juvenile specimen. This fossil is one of the geologically youngest ''T. rex'' specimens known, discovered very near the Cretaceous–Tertiary boundary.


"Victoria"

Victoria is a specimen found near Faith, South Dakota in 2013. Victoria is estimated to be around 12 ft tall and 40 ft long, and she is thought to have died in her thirties. Victoria has also been the subject of a traveling exhibition being displayed in places such as the Arizona Science Center. Her cause of death is unknown; however, she was believed to have been bitten in the lower jaw by another ''Tyrannosaurus''. The bite may have become infected, spreading and leading to sepsis.


"Ivan"

Ivan is a 65% complete ''T. rex'' displayed at the Museum of World Treasures in Wichita, Kansas. The specimen has the most complete tail of any ''T. rex'', only missing around 3 vertebrae. Ivan is around 40 ft long and 12 feet high.


"Trix": RGM 792.000

In 2013, a team of paleontologists from the Naturalis Biodiversity Center (
Leiden, Netherlands Leiden (; in English and archaic Dutch also Leyden) is a city and municipality in the province of South Holland, Netherlands. The municipality of Leiden has a population of 119,713, but the city forms one densely connected agglomeration with ...
) traveled to Montana where they discovered and unearthed a large and remarkably complete ''Tyrannosaurus rex'' specimen that lived 67 million years ago. Black Hills institute collaborated with the team in the excavation. The bones were cleaned and assembled in a mount at Black Hills Institute's installations, with the help of both Chicago's Field Museum of Natural History and the Naturalis Leiden Museum. Chicago's Field museum sent digital models of their famous specimen, FMNH PR 2081 (Sue) to complete the cast and Naturalis museum replicated the bones using 3D-printing technology. The specimen was named ''Trix'' after former Netherlands' Queen Beatrix. At arrival in the Netherlands, it started touring on public display in an itinerant exhibition titled ''T. rex in Town''. The first exhibit spanned from 10 September 2016 to 5 June 2017 and was set at the only room of the Naturalis museum open to public at the time (the 17th-century building known as ''Pesthuis''), due to the fact that the museum was undergoing restoration. When the Netherlands exhibition ended, it continued travelling through other European countries in 2017, 2018 and 2019. As of August 2019, Trix was returned to display at the Naturalis museum where it is installed in a special room that was under construction during Trix's European tour. According to Peter Larson (director of Black Hills Institute), Trix is among the most complete ''Tyrannosaurus'' found. Between 75% and 80% of its skeletal volume was recovered.http://www.thefossilforum.com/uploads/monthly_2016_09/10.thumb.jpg.592a065fd0a51cd2c23fa6d1a2063c29.jpg They are thought to have been at least 30 years old at death


"Titus"

"Titus" is the name given to an obsidian-black skeleton of a ''Tyrannosaurus rex'' discovered in Montana's Hell Creek Formation in 2014 and excavated in 2018. It is 20% complete, and was named after the protagonist in Shakespeare's '' Titus Andronicus''. Exhibited in the Nottingham Natural History Museum for 13 months beginning July 2021, it was during that time only the second specimen of ''Tyrannosaurus'' to be on exhibit in United Kingdom, the other being the type of the junior synonym ''Dynamosaurus imperosus'' which has the jaw on display in the Natural History Museum, London. External bone inspection has revealed injuries to Titus' right tibia (possibly a claw or bite wound); a deformed toe on the right foot; and a bitten and healed tail. The bite wound near the end of the tail indicates a possible attack by another ''Tyrannosaurus rex''. The remains of "Titus" were discovered in September 2014 by commercial paleontologist Craig Pfister near Ekalaka, Carter County, Montana. Excavation of the specimen began in 2018, and took 18 months. The bones of "Titus" were shipped to conservationist Nigel Larkin in the UK, who constructed the mount using a cast of the ''Tyrannosaurus'' specimen
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to supplement the known bones of "Titus", after scanning the bones using
photogrammetry Photogrammetry is the science and technology of obtaining reliable information about physical objects and the environment through the process of recording, measuring and interpreting photographic images and patterns of electromagnetic radiant ima ...
to create digital models that were
3D printed 3D printing or additive manufacturing is the construction of a three-dimensional object from a CAD model or a digital 3D model. It can be done in a variety of processes in which material is deposited, joined or solidified under computer co ...
for use in the exhibition, alongside the display of the real fossil skeleton, and which remained at the museum after the end of the exhibition. For the exhibit at the Nottingham Natural History Museum at Wollaton Hall, Titus was reconstructed "in a walking mode, perhaps searching for prey or returning home after a hunt."


"Tufts-Love": UWBM 99000

In 2016 Greg Wilson, David DeMar, and a paleontology team from the Burke Museum of Natural History and Culture, the University of Washington, and the Dig Field school excavated the partial remains of a ''Tyrannosaurus rex'' from Montana. The partial skeleton was found by two Burke Museum volunteers, Jason Love and Luke Tufts, and was named the "Tufts-Love" rex. Paleontologists at the Burke Museum believe that the Tufts-Love rex was around 15 years old when it died. The skull is of average size for an adult ''T. rex''. The specimen was found in Late Cretaceous deposits and it is estimated to be 66.3 million years old. The Tufts-Love rex is undergoing preparation by Michael Holland and his team at the Burke Museum. The skeleton is estimated to be 30% complete, but it includes a complete (all of the bones of the skull and jaws are preserved) and mostly articulated skull. Holland describes the skull as minimally distorted and in an "exquisite" state of preservation.


"Peter": AWMM-IL 2022.9

Peter is the nickname given to a specimen on loan to the Auckland War Memorial Museum by an anonymous owner, currently on display alongside “Barbara” until the end of 2023. The specimen is estimated to be 66.8 million years old and almost of adult size. He was recovered from Niobrara County, Wyoming. “Peter” is one of only four incredibly rare and visually stunning obisidian black colored tyrannosaurus rex. He was likely killed by exocannibalism as entire sections of bone were damaged, and some were split open by huge crushing bite-forces. The nature of the crushing on the femur and tibia, along with size of the bite marks, indicates that these bones were bitten through by another Tyrannosaurus rex. There is also a set of smaller, parallel tooth marks nearby on the shaft that are not attributable to an adult T. rex. Explanations for this behavior range from response to over-crowded populations, limited food supply, sexual dominance, or even play.


"Barbara": AWMM-IL 2022.21

Barbara is the nickname given to a specimen on loan to the Auckland War Memorial Museum by the same anonymous owner as "Peter", currently on display alongside him until the end of 2023.The pair will be the first adult male and female T rex to be displayed together. She is one of a few specimens believed to be pregnant. “Barbara’s” circumstances are particularly rare, more so when you take into account that she suffered and survived long after a debilitating foot injury. While she was no longer able to capture her own prey it is suggested that she got by with the help of a mate or cohort feeding her, as the injury would have rendering her immobile for upwards of 6 months. This is vaguely supported by trackway evidence that has been used to imply tyrannosaur group hunting. It is doubtful that ‘Barbara’ ever successfully hunted again as a predator unless its prey was in close proximity.


"Dueling Dinosaurs" ''T. rex''

This specimen, considered the most complete of any ''Tyrannosaurus'' specimen at more than 98% preservation, was discovered in Montana in 2006. Following years of unsuccessful attempts to sell it to museums or auction it off, the North Carolina Museum of Natural Sciences started negotiations in 2016, which were prolonged due to a legal battle over the rights to the fossil, which was resolved in 2020. The fossils are set to be displayed by the NC Museum in 2023. The Dueling Dinosaurs ''T. rex'' is an adolescent that was preserved entangled with a '' Triceratops''. Given the injuries present on both fossils, it has been theorized that both died while fighting one another. The specimen has not yet been properly studied due to its recent acquisition to science. However, important biological data is likely preserved with the specimen, including body outlines, skin impressions, soft tissues, injuries, stomach contents, and even original proteins.


See also

*
Timeline of tyrannosaur research This timeline of tyrannosaur research is a chronological listing of events in the history of paleontology focused on the tyrannosaurs, a group of predatory Theropoda, theropod dinosaurs that began as small, long-armed bird-like creatures with ela ...


References


External links


Listing of ''Tyrannosaurus rex'' specimens
at The Theropod Database.
AMNH Article on the First Tyrannosaur specimens


at Skeletal Drawing. {{DEFAULTSORT:Specimens Of Tyrannosaurus Collection of the Carnegie Museum of Natural History Tyrannosaurus Tyrannosaurus Collection of the Smithsonian Institution Articles containing video clips Cretaceous fossil record sv:Tyrannosaurus#Berömda skelett