ENDOPTERYGOTA, also known as HOLOMETABOLA, is a superorder of insects within the infraclass Neoptera that go through distinctive larval , pupal , and adult stages. They undergo a radical metamorphosis , with the larval and adult stages differing considerably in their structure and behaviour. This is called holometabolism , or complete metamorphism.
They are distinguished from the
Exopterygota (or Hemipterodea) by the
way in which their wings develop.
The earliest endopterygote fossils date from the
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Holometabolism stages in Hymenoptera.
ITIS considers any subdivision of the Neoptera beyond the orders invalid, but this is almost universally rejected.
More recently, this has increasingly been rejected and DNA sequence data seem to verify that the hymenopterans are indeed among the most basal endopterygotes, whereas flies and fleas are often considered the most radically advanced insects. This calls the previous subdivision into question, and consequently several new taxa have been proposed, splitting up the Endopterygota. While some groups (such as the "sucking-stinging" fly-flea assemblage or the caddisfly-butterfly group) seem indeed to be good clades , it is not likely that the relationships of the endopterygotes, or the neopteran insects in general, will be resolved in detail soon.
* Hymenoptera (ants , bees , etc.)
Proposed superorder Coleopterida
Proposed superorder Neuropterida /Neuropteroidea
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* ^ Rolf G. Beutel; Hans Pohl (2006). "Endopterygote systematics – where do we stand and what is the goal (Hexapoda, Arthropoda)?". Systematic Entomology . 31 (2): 202–219. doi :10.1111/j.1365-3113.2006.00341.x . * ^ A. Nel; P. Roques; P. Nel; J. Prokop; J. S. Steyer (2007). "The earliest holometabolous insect from the Carboniferous: a "crucial" innovation with delayed success (Insecta Protomeropina Protomeropidae)". Annales de la Société Entomologique de France . 43 (3): 349–355.
* Wd : Q304358 * ADW : Holometabola * EoL : 3016961 * ITIS : 109215 * NCBI : 33392
* v * t * e
* Archaeognatha (jumping bristletails)
D i c o n d y l i a
* Thysanura (Zygentoma) (silverfish, firebrats)
P t e r y g o t a
* Ephemeroptera (mayflies)
* Odonata (dragonflies, damselflies)
N e o p t e r a
* Plecoptera (stoneflies) * Dermaptera (earwigs) * Embioptera (webspinners) * Phasmatodea (stick and leaf insects) * Notoptera (ice-crawlers, gladiators) * Orthoptera (crickets, wetas, grasshoppers, locusts) * Zoraptera (angel insects)
* Blattodea (cockroaches, termites) * Mantodea (mantises)
E u m e t a b o l a
E n d o p t e r y g o t a
* HYMENOPTERA (SAWFLIES, WASPS, ANTS, BEES)
* Trichoptera (caddisflies) * LEPIDOPTERA (MOTHS, BUTTERFLIES)
* Four most speciose orders are marked in BOLD * Italic are paraphyletic groups * Based on Sasaki et al. (2013)
* Extinct incertae sedis families and genera are marked in italic