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Odonata
Odonata is an order of flying insects that includes the dragonflies and damselflies. Members of the group first appeared during the Triassic, though members of their total group, Odonatoptera, first appeared in Late Carboniferous. The two common groups are distinguished with dragonflies, placed in the suborder Epiprocta, usually being larger, with eyes together and wings up or out at rest, while damselflies, suborder Zygoptera, are usually smaller with eyes placed apart and wings along body at rest. All Odonata have aquatic larvae called naiads (nymphs), and all of them, larvae and adults, are carnivorous. The adults can land, but rarely walk. Their legs are specialised for catching prey. They are almost entirely insectivorous. Etymology and terminology Fabricius coined the term ''Odonata'' in 1793 from the Ancient Greek ( Ionic form of ) 'tooth'. One hypothesis is that it was because their maxillae are notably toothed. Most insects also have toothed mandibles. The ...
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Damselfly
Damselflies are flying insects of the suborder Zygoptera in the order Odonata. They are similar to dragonflies, which constitute the other odonatan suborder, Anisoptera, but are smaller and have slimmer bodies. Most species fold the wings along the body when at rest, unlike dragonflies which hold the wings flat and away from the body. An ancient group, damselflies have existed since at least the Lower Permian, and are found on every continent except Antarctica. All damselflies are predatory insects; both nymphs and adults actively hunt and eat other insects. The nymphs are aquatic, with different species living in a variety of freshwater habitats including acidic bogs, ponds, lakes and rivers. The nymphs moult repeatedly, at the last moult climbing out of the water to undergo metamorphosis. The skin splits down the back, they emerge and inflate their wings and abdomen to gain their adult form. Their presence on a body of water indicates that it is relatively unpolluted, but t ...
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Dragonflies
A dragonfly is a flying insect belonging to the infraorder Anisoptera below the order Odonata. About 3,000 extant species of true dragonfly are known. Most are tropical, with fewer species in temperate regions. Loss of wetland habitat threatens dragonfly populations around the world. Adult dragonflies are characterized by a pair of large, multifaceted compound eyes, two pairs of strong, transparent wings, sometimes with coloured patches, and an elongated body. Many dragonflies have brilliant iridescent or metallic colours produced by structural colouration, making them conspicuous in flight. An adult dragonfly's compound eyes have nearly 24,000 ommatidia each. Dragonflies can be mistaken for the closely related damselflies, which make up the other odonatan infraorder ( Zygoptera) and are similar in body plan though usually lighter in build; however, the wings of most dragonflies are held flat and away from the body, while damselflies hold their wings folded at rest, along ...
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Anisoptera
A dragonfly is a flying insect belonging to the infraorder Anisoptera below the order Odonata. About 3,000 extant species of true dragonfly are known. Most are tropical, with fewer species in temperate regions. Loss of wetland habitat threatens dragonfly populations around the world. Adult dragonflies are characterized by a pair of large, multifaceted compound eyes, two pairs of strong, transparent wings, sometimes with coloured patches, and an elongated body. Many dragonflies have brilliant iridescent or metallic colours produced by structural colouration, making them conspicuous in flight. An adult dragonfly's compound eyes have nearly 24,000 ommatidia each. Dragonflies can be mistaken for the closely related damselflies, which make up the other odonatan infraorder ( Zygoptera) and are similar in body plan though usually lighter in build; however, the wings of most dragonflies are held flat and away from the body, while damselflies hold their wings folded at rest, along ...
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Epiprocta
Epiprocta is one of the two extant suborders of the Odonata (the order to which dragonflies and damselflies belong). It was proposed relatively recently, having been created to accommodate the inclusion of the Anisozygoptera. The latter has been shown to be not a natural suborder, but rather a paraphyletic In taxonomy, a group is paraphyletic if it consists of the group's last common ancestor and most of its descendants, excluding a few monophyletic subgroups. The group is said to be paraphyletic ''with respect to'' the excluded subgroups. In ... collection of lineages, so it has been combined with the previous suborder Anisoptera, the well-known dragonflies, into the Epiprocta. The old suborder Anisoptera is proposed to become an infraorder within the Epiprocta, whereas the "anisozygopterans" included here form the infraorder Epiophlebioptera. References External links * * Insect suborders {{insect-stub ...
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Odonatoptera
The Odonatoptera are a superorder (sometimes treated as an order) of ancient winged insects, placed in the probably paraphyletic group Palaeoptera. The dragonflies and damselflies are the only living members of this group, which was far more diverse in the late Paleozoic and contained gigantic species, including the griffinflies (colloquially called "giant dragonflies", although they were not dragonflies in the strict sense) of the order Meganisoptera (formerly Protodonata). This lineage dates back at least to the Bashkirian, not quite 320 million years ago.  Systematics and taxonomy There is little consensus about the relationships of the Odonatoptera. What is certain is that they are a clade of winged insects that stands outside the Neoptera. But various authors' analyses have yielded any one of three mutually exclusive phylogenies, or some variant thereof: The least problematic (in a taxonomic sense) view is that the Odonatoptera are the sister taxon of the Ephemeropter ...
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Protodonata
Meganisoptera is an extinct order of very large to gigantic insects, informally called griffinflies. The order was formerly named Protodonata, the "proto-Odonata", for their similar appearance and supposed relation to modern Odonata (damselflies and dragonflies). They range in Palaeozoic (Late Carboniferous to Late Permian) times. Though most were only slightly larger than modern dragonflies, the order includes the largest known insect species, such as the late Carboniferous '' Meganeura monyi'' and the even larger early Permian ''Meganeuropsis permiana'', with wingspans of up to . The forewings and hindwings are similar in venation (a primitive feature) except for the larger anal (rearwards) area in the hindwing. The forewing is usually slenderer and slightly longer than the hindwing. Unlike the true dragonflies, the Odonata, they had no pterostigmas, and had a somewhat simpler pattern of veins in the wings. Most specimens are known from wing fragments only; with only a few as ...
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Palaeoptera
The name Palaeoptera (from Greek ( 'old') + ( 'wing')) has been traditionally applied to those ancestral groups of winged insects (most of them extinct) that lacked the ability to fold the wings back over the abdomen as characterizes the Neoptera. The Diaphanopterodea, which are palaeopteran insects, had independently and uniquely evolved a different wing-folding mechanism. Both mayflies and dragonflies lack any of the smell centers in their brain found in Neoptera. Disputed status The complexities of the wing-folding mechanism, as well as the mechanical operation of the wings in flight ( indirect flight muscles), are such that it clearly indicates the Neoptera are a monophyletic lineage. The problem is that the plesiomorphic absence of wing-folding does not necessarily mean the Palaeoptera form a natural group – they may simply be an assemblage containing all insects, closely related or not, that "are not Neoptera", an example of a wastebasket taxon. If the extinct l ...
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Insect Flight
Insects are the only group of invertebrates that have evolved wings and flight. Insects first flew in the Carboniferous, some 350 to 400 million years ago, making them the first animals to evolve flight. Wings may have evolved from appendages on the sides of existing limbs, which already had nerves, joints, and muscles used for other purposes. These may initially have been used for sailing on water, or to slow the rate of descent when gliding. Two insect groups, the dragonflies and mayflies, have flight muscles attached directly to the wings. In other winged insects, flight muscles attach to the thorax, which make it oscillate in order to induce the wings to beat. Of these insects, some ( flies and some beetles) achieve very high wingbeat frequencies through the evolution of an "asynchronous" nervous system, in which the thorax oscillates faster than the rate of nerve impulses. Not all insects are capable of flight. A number of apterous insects have secondarily lost thei ...
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Insect
Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs of jointed legs, compound eyes and one pair of antennae. Their blood is not totally contained in vessels; some circulates in an open cavity known as the haemocoel. Insects are the most diverse group of animals; they include more than a million described species and represent more than half of all known living organisms. The total number of extant species is estimated at between six and ten million; In: potentially over 90% of the animal life forms on Earth are insects. Insects may be found in nearly all environments, although only a small number of species reside in the oceans, which are dominated by another arthropod group, crustaceans, which recent research has indicated insects are nested within. Nearly all insects hatch from ...
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Insect Mouthparts
Insects have mouthparts that may vary greatly across insect species, as they are adapted to particular modes of feeding. The earliest insects had chewing mouthparts. Most specialisation of mouthparts are for piercing and sucking, and this mode of feeding has evolved a number of times idependently. For example, mosquitoes and aphids (which are true bugs) both pierce and suck, however female mosquitoes feed on animal blood whereas aphids feed on plant fluids. Evolution Like most external features of arthropods, the mouthparts of Hexapoda are highly derived. Insect mouthparts show a multitude of different functional mechanisms across the wide diversity of insect species. It is common for significant homology to be conserved, with matching structures forming from matching primordia, and having the same evolutionary origin. However, even if structures are almost physically and functionally identical, they may not be homologous; their analogous functions and appearance might be the ...
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Ionic Greek
Ionic Greek ( grc, Ἑλληνικὴ Ἰωνική, Hellēnikē Iōnikē) was a subdialect of the Attic–Ionic or Eastern dialect group of Ancient Greek. History The Ionic dialect appears to have originally spread from the Greek mainland across the Aegean at the time of the Dorian invasions, around the 11th century BC during the early Greek Dark Ages. By the end of Archaic Greece and early Classical Greece in the 5th century BC, the central west coast of Asia Minor, along with the islands of Chios and Samos, formed the heartland of Ionia proper. The Ionic dialect was also spoken on islands across the central Aegean and on the large island of Euboea north of Athens. The dialect was soon spread by Ionian colonization to areas in the northern Aegean, the Black Sea, and the western Mediterranean, including Magna Graecia in Sicily and Italy. The Ionic dialect is generally divided into two major time periods, Old Ionic (or Old Ionian) and New Ionic (or New Ionian). The transitio ...
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Clade
A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, the equivalent Latin term ''cladus'' (plural ''cladi'') is often used in taxonomical literature. The common ancestor may be an individual, a population, or a species (extinct or extant). Clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently. Clades are termed monophyletic (Greek: "one clan") groups. Over the last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming taxa that are not clades; that is, taxa that are not monophyletic. Some of the relationships between org ...
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