Dromedary
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The dromedary (''Camelus dromedarius'' or ;), also known as the dromedary camel, Arabian camel, or one-humped camel, is a large
even-toed ungulate The even-toed ungulates (Artiodactyla , ) are ungulates—hoofed animals—which bear weight equally on two (an even number) of their five toes: the third and fourth. The other three toes are either present, absent, vestigial, or pointing poster ...
, of the genus ''
Camelus A camel (from: la, camelus and grc-gre, κάμηλος (''kamēlos'') from Hebrew or Phoenician: גָמָל ''gāmāl''.) is an even-toed ungulate in the genus ''Camelus'' that bears distinctive fatty deposits known as "humps" on its back. C ...
'', with one hump on its back. It is the tallest of the three species of
camel A camel (from: la, camelus and grc-gre, κάμηλος (''kamēlos'') from Hebrew or Phoenician: גָמָל ''gāmāl''.) is an even-toed ungulate in the genus ''Camelus'' that bears distinctive fatty deposits known as "humps" on its back. C ...
; adult males stand at the shoulder, while females are tall. Males typically weigh between , and females weigh between . The species' distinctive features include its long, curved neck, narrow chest, a single hump (compared with two on the
Bactrian camel The Bactrian camel (''Camelus bactrianus''), also known as the Mongolian camel or domestic Bactrian camel, is a large even-toed ungulate native to the steppes of Central Asia. It has two humps on its back, in contrast to the single-humped drom ...
and
wild Bactrian camel The wild Bactrian camel (''Camelus ferus'') is a critically endangered species of camel living in parts of northwestern China and southwestern Mongolia. It is closely related to the Bactrian camel (''Camelus bactrianus''). Both are large, doub ...
), and long hairs on the throat, shoulders and hump. The coat is generally a shade of brown. The hump, tall or more, is made of fat bound together by
fibrous tissue Fiber or fibre (from la, fibra, links=no) is a natural or artificial substance that is significantly longer than it is wide. Fibers are often used in the manufacture of other materials. The strongest engineering materials often incorporate ...
. Dromedaries are mainly active during daylight hours. They form herds of about 20 individuals, which are led by a dominant male. They feed on foliage and desert vegetation; several adaptations, such as the ability to tolerate losing more than 30% of its total water content, allow it to thrive in its desert habitat. Mating occurs annually and peaks in the rainy season; females bear a single calf after a
gestation Gestation is the period of development during the carrying of an embryo, and later fetus, inside viviparous animals (the embryo develops within the parent). It is typical for mammals, but also occurs for some non-mammals. Mammals during pregna ...
of 15 months. The dromedary has not occurred naturally in the wild for nearly 2,000 years. It was probably first
domesticated Domestication is a sustained multi-generational relationship in which humans assume a significant degree of control over the reproduction and care of another group of organisms to secure a more predictable supply of resources from that group. A ...
in the
Arabian Peninsula The Arabian Peninsula, (; ar, شِبْهُ الْجَزِيرَةِ الْعَرَبِيَّة, , "Arabian Peninsula" or , , "Island of the Arabs") or Arabia, is a peninsula of Western Asia, situated northeast of Africa on the Arabian Plate ...
about 4,000 years ago, or in
Somalia Somalia, , Osmanya script: 𐒈𐒝𐒑𐒛𐒐𐒘𐒕𐒖; ar, الصومال, aṣ-Ṣūmāl officially the Federal Republic of SomaliaThe ''Federal Republic of Somalia'' is the country's name per Article 1 of thProvisional Constituti ...
where there are paintings in
Laas Geel Laas Geel ( so, Laas Geel), also spelled Laas Gaal, are cave formations on the rural outskirts of Hargeisa, Somaliland, situated in the Maroodi Jeex region of the country. They contain some of the earliest known cave paintings of domesticated Afr ...
that figure it from 5,000 to 9,000 years ago. In the wild, the dromedary inhabited arid regions, including the
Sahara Desert , photo = Sahara real color.jpg , photo_caption = The Sahara taken by Apollo 17 astronauts, 1972 , map = , map_image = , location = , country = , country1 = , ...
. The domesticated dromedary is generally found in the semi-arid to arid regions of the Old World, mainly in Africa and the Arabian Peninsula, and a significant feral population occurs in Australia. Products of the dromedary, including its meat and milk, support several North African tribes; it is also commonly used for riding and as a pack animal.


Etymology

The common name "dromedary" comes from the
Old French Old French (, , ; Modern French: ) was the language spoken in most of the northern half of France from approximately the 8th to the 14th centuries. Rather than a unified language, Old French was a linkage of Romance dialects, mutually intelligib ...
''dromedaire'' or the
Late Latin Late Latin ( la, Latinitas serior) is the scholarly name for the form of Literary Latin of late antiquity.Roberts (1996), p. 537. English dictionary definitions of Late Latin date this period from the , and continuing into the 7th century in t ...
''dromedarius''. These originated from the
Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...
word ''dromas'', δρομάς (ο, η) ( GEN (γενική) ''dromados'', δρομάδος), meaning "running" or "runner",. used in Greek in the combination (''dromas kamelos''), literally "running camel", to refer to the dromedary. The first recorded use in English of the name "dromedary" occurred in the 14th century. The dromedary possibly originated in Arabia or Somalia and is therefore sometimes referred to as the Arabian or East African camel. The word "camel" generally refers either to the dromedary or the
congeneric Congener may refer to: * A thing or person of the same kind as another, or of the same group. * Congener (biology), organisms within the same genus. * Congener (chemistry), related chemicals, e.g., elements in the same group of the periodic table. ...
Bactrian; the word came into English via
Old Norman Old Norman, also called Old Northern French or Old Norman French ( fro, Ancien Normant, nrf, Ancien Normaund), was one of many varieties of the '' langues d'oïl'' native to northern France. It was spoken throughout the region of what is now call ...
, from the
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
word ''camēlus'', from
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic peri ...
(''kámēlos''), ultimately from a Semitic source akin to
Hebrew Hebrew (; ; ) is a Northwest Semitic language of the Afroasiatic language family. Historically, it is one of the spoken languages of the Israelites and their longest-surviving descendants, the Jews and Samaritans. It was largely preserved ...
(''gamál'') and
Arabic Arabic (, ' ; , ' or ) is a Semitic languages, Semitic language spoken primarily across the Arab world.Semitic languages: an international handbook / edited by Stefan Weninger; in collaboration with Geoffrey Khan, Michael P. Streck, Janet C ...
(''jamal'').


Taxonomy and classification

The dromedary shares the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''
Camelus A camel (from: la, camelus and grc-gre, κάμηλος (''kamēlos'') from Hebrew or Phoenician: גָמָל ''gāmāl''.) is an even-toed ungulate in the genus ''Camelus'' that bears distinctive fatty deposits known as "humps" on its back. C ...
'' with the Bactrian camel (''C. bactrianus'') and the wild Bactrian camel (''C. ferus''). The dromedary belongs to the family
Camelidae Camelids are members of the biological family Camelidae, the only currently living family in the suborder Tylopoda. The seven extant members of this group are: dromedary camels, Bactrian camels, wild Bactrian camels, llamas, alpacas, vicuñas, ...
. The
ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic peri ...
philosopher
Aristotle Aristotle (; grc-gre, Ἀριστοτέλης ''Aristotélēs'', ; 384–322 BC) was a Greek philosopher and polymath during the Classical period in Ancient Greece. Taught by Plato, he was the founder of the Peripatetic school of phil ...
(4th century BC) was the first to
describe Shneur Hasofer is a Hasidic musician known as DeScribe. Hasofer's musical style has been characterized as "Hasidic hip-hop," "Hasidic rap" and "Hasidic R&B". Background Hasofer was born to a Chabad Hasidic family in Melbourne, Australia. Hasof ...
the species of ''Camelus''. He named two species in his ''
History of Animals ''History of Animals'' ( grc-gre, Τῶν περὶ τὰ ζῷα ἱστοριῶν, ''Ton peri ta zoia historion'', "Inquiries on Animals"; la, Historia Animalium, "History of Animals") is one of the major texts on biology by the ancient Gr ...
''; the one-humped Arabian camel and the two-humped Bactrian camel. The dromedary was given its current binomial name ''Camelus dromedarius'' by Swedish zoologist
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
in his 1758 publication ''
Systema Naturae ' (originally in Latin written ' with the ligature æ) is one of the major works of the Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the system, now known as binomial nomen ...
''. In 1927, British veterinarian Arnold Leese classified dromedaries by their basic habitats; the hill camels are small, muscular animals and efficient beasts of burden; the larger plains camels could be further divided into the desert type that can bear light burdens and are apt for riding, and the riverine type – slow animals that can bear heavy burdens; and those intermediate between these two types. In 2007, Peng Cui of the
Chinese Academy of Sciences The Chinese Academy of Sciences (CAS); ), known by Academia Sinica in English until the 1980s, is the national academy of the People's Republic of China for natural sciences. It has historical origins in the Academia Sinica during the Republ ...
and colleagues carried out a
phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...
study of the evolutionary relationships between the two
tribes The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide usage of the term in English is in the discipline of anthropology. This definition is contested, in part due to conflic ...
of Camelidae;
Camelini Camelini is a tribe of terrestrial herbivores in the family Camelidae, endemic to Asia, North America, and Africa from the Late Eocene to the present. It includes the living genus ''Camelus'' as the type genus. At least one genus, ''Eulamaops'', ...
– consisting of the three ''Camelus'' species (the study considered the wild Bactrian camel as a
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
of the Bactrian camel) – and
Lamini Lamini (members are called ''laminoids'') is a tribe of the subfamily Camelinae. It contains one extant genus with four species, all exclusively from South America: llamas, alpacas, vicuñas, and guanacos. The former two are domesticated specie ...
, which consists of the
alpaca The alpaca (''Lama pacos'') is a species of South American camelid mammal. It is similar to, and often confused with, the llama. However, alpacas are often noticeably smaller than llamas. The two animals are closely related and can successfu ...
(''Vicugna pacos''), the
guanaco The guanaco (; ''Lama guanicoe'') is a camelid native to South America, closely related to the llama. Guanacos are one of two wild South American camelids, the other being the vicuña, which lives at higher elevations. Etymology The guanaco g ...
(''Lama guanicoe''), the
llama The llama (; ) (''Lama glama'') is a domesticated South American camelid, widely used as a List of meat animals, meat and pack animal by Inca empire, Andean cultures since the Pre-Columbian era. Llamas are social animals and live with othe ...
(''L. glama'') and the
vicuña The vicuña (''Lama vicugna'') or vicuna (both , very rarely spelled ''vicugna'', its former genus name) is one of the two wild South American camelids, which live in the high alpine areas of the Andes, the other being the guanaco, which live ...
(''V. vicugna''). The study showed the two tribes had diverged 25 million years ago (early
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
), earlier than previously estimated from North American fossils. The dromedary and the Bactrian camel often interbreed to produce fertile offspring. Where the ranges of the species overlap, such as in northern Punjab, Persia, and Afghanistan, the
phenotypic In genetics, the phenotype () is the set of observable characteristics or traits of an organism. The term covers the organism's morphology or physical form and structure, its developmental processes, its biochemical and physiological proper ...
differences between them tend to decrease as a result of extensive crossbreeding. The fertility of their
hybrid Hybrid may refer to: Science * Hybrid (biology), an offspring resulting from cross-breeding ** Hybrid grape, grape varieties produced by cross-breeding two ''Vitis'' species ** Hybridity, the property of a hybrid plant which is a union of two dif ...
has given rise to speculation that the dromedary and the Bactrian camel should be merged into a single species with two varieties. However, a 1994 analysis of the
mitochondria A mitochondrion (; ) is an organelle found in the Cell (biology), cells of most Eukaryotes, such as animals, plants and Fungus, fungi. Mitochondria have a double lipid bilayer, membrane structure and use aerobic respiration to generate adenosi ...
l
cytochrome b Cytochrome b within both molecular and cell biology, is a protein found in the mitochondria of eukaryotic cells. It functions as part of the electron transport chain and is the main subunit of transmembrane cytochrome bc1 and b6f complexes. F ...
gene In biology, the word gene (from , ; "...Wilhelm Johannsen coined the word gene to describe the Mendelian units of heredity..." meaning ''generation'' or ''birth'' or ''gender'') can have several different meanings. The Mendelian gene is a ba ...
showed the species display 10.3% divergence in their
sequences In mathematics, a sequence is an enumerated collection of objects in which repetitions are allowed and order matters. Like a set, it contains members (also called ''elements'', or ''terms''). The number of elements (possibly infinite) is called t ...
.


Genetics and hybrids

The dromedary has 74
diploid Ploidy () is the number of complete sets of chromosomes in a cell, and hence the number of possible alleles for autosomal and pseudoautosomal genes. Sets of chromosomes refer to the number of maternal and paternal chromosome copies, respectively ...
chromosomes, the same as other camelids. The
autosome An autosome is any chromosome that is not a sex chromosome. The members of an autosome pair in a diploid cell have the same morphology, unlike those in allosome, allosomal (sex chromosome) pairs, which may have different structures. The DNA in au ...
s consist of five pairs of small to medium-sized metacentrics and submetacentrics. The
X chromosome The X chromosome is one of the two sex-determining chromosomes (allosomes) in many organisms, including mammals (the other is the Y chromosome), and is found in both males and females. It is a part of the XY sex-determination system and XO sex ...
is the largest in the metacentric and submetacentric group. There are 31 pairs of
acrocentric The centromere links a pair of sister chromatids together during cell division. This constricted region of chromosome connects the sister chromatids, creating a short arm (p) and a long arm (q) on the chromatids. During mitosis, spindle fibers ...
s. The dromedary's
karyotype A karyotype is the general appearance of the complete set of metaphase chromosomes in the cells of a species or in an individual organism, mainly including their sizes, numbers, and shapes. Karyotyping is the process by which a karyotype is disce ...
is similar to that of the Bactrian camel. Camel hybridization began in the first millennium BC. For about a thousand years, Bactrian camels and dromedaries have been successfully bred in regions where they are
sympatric In biology, two related species or populations are considered sympatric when they exist in the same geographic area and thus frequently encounter one another. An initially interbreeding population that splits into two or more distinct species sh ...
to form hybrids with either a long, slightly lopsided hump or two humps – one small and one large. These hybrids are larger and stronger than their parents – they can bear greater loads. A cross between a first generation female hybrid and a male Bactrian camel can also produce a hybrid. Hybrids from other combinations tend to be bad-tempered or
runt In a group of animals (usually a litter of animals born in multiple births), a runt is a member which is significantly smaller or weaker than the others. Owing to its small size, a runt in a litter faces obvious disadvantage, including difficulti ...
s.


Evolution

The extinct ''
Protylopus ''Protylopus'' is an extinct genus of camel that lived during middle to late Eocene some 50-40 million years ago in North America. Along with being the oldest camel known, it was also the smallest, reaching a length of , and probably weighing a ...
'', which occurred in North America during the upper
Eocene The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene' ...
, is the oldest and the smallest-known camel. During the transition from
Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fina ...
, several mammals faced extinction. This period marked the successful radiation of the ''Camelus'' species, which migrated over the Bering Strait and dispersed widely into Asia, eastern Europe and Africa. By the Pleistocene, ancestors of the dromedary occurred in the Middle East and northern Africa. The modern dromedary probably evolved in the hotter, arid regions of western Asia from the Bactrian camel, which in turn was closely related to the earliest Old World camels. This hypothesis is supported by the fact that the dromedary
foetus A fetus or foetus (; plural fetuses, feti, foetuses, or foeti) is the unborn offspring that develops from an animal embryo. Following embryonic development the fetal stage of development takes place. In human prenatal development, fetal develo ...
has two humps, while in the adult male an anterior
vestigial Vestigiality is the retention, during the process of evolution, of genetically determined structures or attributes that have lost some or all of the ancestral function in a given species. Assessment of the vestigiality must generally rely on co ...
hump is present. A jawbone of a dromedary that dated from 8,200 BC was found in Saudi Arabia on the southern coast of the
Red Sea The Red Sea ( ar, البحر الأحمر - بحر القلزم, translit=Modern: al-Baḥr al-ʾAḥmar, Medieval: Baḥr al-Qulzum; or ; Coptic: ⲫⲓⲟⲙ ⲛ̀ϩⲁϩ ''Phiom Enhah'' or ⲫⲓⲟⲙ ⲛ̀ϣⲁⲣⲓ ''Phiom ǹšari''; T ...
. In 1975,
Richard Bulliet Richard W. Bulliet (born 1940) is a professor of history at Columbia University who specializes in the history of Islamic society and institutions, the history of technology, and the history of the role of animals in human society. Early life ...
of
Columbia University Columbia University (also known as Columbia, and officially as Columbia University in the City of New York) is a private research university in New York City. Established in 1754 as King's College on the grounds of Trinity Church in Manhatt ...
wrote that the dromedary exists in large numbers in areas from which the Bactrian camel has disappeared; the converse is also true to a great extent. He said this substitution could have taken place because of the heavy dependence on the milk, meat and wool of the dromedary by Syrian and Arabian nomads, while the Asiatic people domesticated the Bactrian camel but did not have to depend upon its products.


Characteristics

The dromedary is the tallest of the three camel species. Adult males range in height between at the shoulder;
Guinness World Records ''Guinness World Records'', known from its inception in 1955 until 1999 as ''The Guinness Book of Records'' and in previous United States editions as ''The Guinness Book of World Records'', is a reference book published annually, listing world ...

Largest species of camel
/ref> females range between . Males typically weigh between ; females range between . The distinctive features are its long, curved neck, narrow chest and single hump (the Bactrian camel has two), thick, double-layered eyelashes and bushy eyebrows. They have sharp vision and a good sense of smell. The male has a soft
palate The palate () is the roof of the mouth in humans and other mammals. It separates the oral cavity from the nasal cavity. A similar structure is found in crocodilians, but in most other tetrapods, the oral and nasal cavities are not truly sepa ...
( in Arabic) nearly long, which he inflates to produce a deep pink sac. The palate, which is often mistaken for the tongue, dangles from one side of the mouth and is used to attract females during the mating season. The coat is generally brown but can range from black to nearly white. Leese reported
piebald A piebald or pied animal is one that has a pattern of unpigmented spots (white) on a pigmented background of hair, feathers or scales. Thus a piebald black and white dog is a black dog with white spots. The animal's skin under the white backgro ...
dromedaries in
Kordofan Kordofan ( ar, كردفان ') is a former province of central Sudan. In 1994 it was divided into three new federal states: North Kordofan, South Kordofan and West Kordofan. In August 2005, West Kordofan State was abolished and its territory ...
and Darfur in Sudan. Piebald coloration in some camels is thought to be caused by the ''KITW1'' allele of the KIT gene, though there is likely at least one other mutation that also causes white spotting. The hair is long and concentrated on the throat, shoulders and the hump. The large eyes are protected by prominent
supraorbital ridge The brow ridge, or supraorbital ridge known as superciliary arch in medicine, is a bony ridge located above the eye sockets of all primates. In humans, the eyebrows are located on their lower margin. Structure The brow ridge is a nodule or crest ...
s; the ears are small and rounded. The hump is at least high. The dromedary has long, powerful legs with two toes on each foot. The feet resemble flat, leathery pads. Like the
giraffe The giraffe is a large African hoofed mammal belonging to the genus ''Giraffa''. It is the tallest living terrestrial animal and the largest ruminant on Earth. Traditionally, giraffes were thought to be one species, ''Giraffa camelopardalis ...
, dromedaries moves both legs on one side of the body at the same time. Compared with the Bactrian camel, the dromedary has a lighter build, longer limbs, shorter hairs, a harder palate and an insignificant or absent ethmoidal
fissure A fissure is a long, narrow crack opening along the surface of Earth. The term is derived from the Latin word , which means 'cleft' or 'crack'. Fissures emerge in Earth's crust, on ice sheets and glaciers, and on volcanoes. Ground fissure ...
. Unlike the camelids of the genus ''Lama'', the dromedary has a hump, and in comparison has a longer tail, smaller ears, squarer feet, and a greater height at the shoulder. The dromedary has four teats instead of the two in the ''Lama'' species.


Anatomy

The
cranium The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, the ...
of the dromedary consists of a
postorbital bar The postorbital bar (or postorbital bone) is a bony arched structure that connects the frontal bone of the skull to the zygomatic arch, which runs laterally around the eye socket. It is a trait that only occurs in mammalian taxa, such as most strep ...
, a
tympanic bulla The tympanic part of the temporal bone is a curved plate of bone lying below the squamous part of the temporal bone, in front of the mastoid process, and surrounding the external part of the ear canal. It originates as a separate bone (tympanic ...
filled with spongiosa, a well-defined sagittal crest, a long facial part and an indented nasal bone. Typically, there are eight
sternal The sternum or breastbone is a long flat bone located in the central part of the chest. It connects to the ribs via cartilage and forms the front of the rib cage, thus helping to protect the heart, lungs, and major blood vessels from injury. Sh ...
and four non-sternal pairs of ribs. The spinal cord is nearly long; it terminates in the second and third
sacral vertebra The sacrum (plural: ''sacra'' or ''sacrums''), in human anatomy, is a large, triangular bone at the base of the spine that forms by the fusing of the sacral vertebrae (S1S5) between ages 18 and 30. The sacrum situates at the upper, back part ...
. The
fibula The fibula or calf bone is a leg bone on the lateral side of the tibia, to which it is connected above and below. It is the smaller of the two bones and, in proportion to its length, the most slender of all the long bones. Its upper extremity is ...
is reduced to a malleolar bone. The dromedary is a digitigrade animal; it walks on its toes, which are known as digits. It lacks the second and fifth digits. The front feet are wide and long; they are larger than the hind feet, which measure wide and long. The dromedary has 22
milk teeth Deciduous teeth or primary teeth, also informally known as baby teeth, milk teeth, or temporary teeth,Illustrated Dental Embryology, Histology, and Anatomy, Bath-Balogh and Fehrenbach, Elsevier, 2011, page 255 are the first set of teeth in the ...
, which are eventually replaced by 34
permanent teeth Permanent teeth or adult teeth are the second set of teeth formed in diphyodont mammals. In humans and old world simians, there are thirty-two permanent teeth, consisting of six maxillary and six mandibular molars, four maxillary and four mandibul ...
. The
dental formula Dentition pertains to the development of teeth and their arrangement in the mouth. In particular, it is the characteristic arrangement, kind, and number of teeth in a given species at a given age. That is, the number, type, and morpho-physiolo ...
for permanent dentition is , and for
milk Milk is a white liquid food produced by the mammary glands of mammals. It is the primary source of nutrition for young mammals (including breastfed human infants) before they are able to digestion, digest solid food. Immune factors and immune ...
dentition. In the juvenile, the lower first
molars The molars or molar teeth are large, flat teeth at the back of the mouth. They are more developed in mammals. They are used primarily to grind food during chewing. The name ''molar'' derives from Latin, ''molaris dens'', meaning "millstone to ...
develop by 12 to 15 months and the permanent lower
incisor Incisors (from Latin ''incidere'', "to cut") are the front teeth present in most mammals. They are located in the premaxilla above and on the mandible below. Humans have a total of eight (two on each side, top and bottom). Opossums have 18, whe ...
s appear at 4.5 to 6.5 years of age. All teeth are in use by 8 years. The
lenses A lens is a transmissive optical device which focuses or disperses a light beam by means of refraction. A simple lens consists of a single piece of transparent material, while a compound lens consists of several simple lenses (''elements''), ...
of the eyes contain
crystallin In anatomy, a crystallin is a water-soluble structural protein found in the lens and the cornea of the eye accounting for the transparency of the structure. It has also been identified in other places such as the heart, and in aggressive breast c ...
, which constitutes 8 to 13% of the protein present there. The skin is black; the
epidermis The epidermis is the outermost of the three layers that comprise the skin, the inner layers being the dermis and hypodermis. The epidermis layer provides a barrier to infection from environmental pathogens and regulates the amount of water rele ...
is thick and the dermis is thick. The hump is composed of fat bound together by
fibrous tissue Fiber or fibre (from la, fibra, links=no) is a natural or artificial substance that is significantly longer than it is wide. Fibers are often used in the manufacture of other materials. The strongest engineering materials often incorporate ...
. There are no glands on the face; males have glands that appear to be modified apocrine sweat glands that secrete pungent, coffee-coloured fluid during the rut, located on either side of the neck midline. The glands generally grow heavier during the rut, and range from . Each cover hair is associated with an
arrector pilli The arrector pili muscles, also known as hair erector muscles, are small muscles attached to hair follicles in mammals. Contraction of these muscles causes the hairs to stand on end, known colloquially as goose bumps (piloerection). Structure ...
muscle, a hair follicle, a ring of
sebaceous gland A sebaceous gland is a microscopic exocrine gland in the skin that opens into a hair follicle to secrete an oily or waxy matter, called sebum, which lubricates the hair and skin of mammals. In humans, sebaceous glands occur in the greatest nu ...
s and a sweat gland. Females have cone-shaped, four-chambered
mammary gland A mammary gland is an exocrine gland in humans and other mammals that produces milk to feed young offspring. Mammals get their name from the Latin word ''mamma'', "breast". The mammary glands are arranged in organs such as the breasts in primat ...
s that are long with a base diameter of . These glands can produce milk with up to 90% water content even if the mother is at risk of dehydration. The heart weighs around ; it has two ventricle (heart), ventricles with the tip curving to the left. The pulse rate is 50 beats per minute. The dromedary is the only mammal with oval red blood corpuscles, which facilitates blood flow during dehydration. The pH of the blood varies from 7.1 to 7.6 (slightly alkaline). The individual's state of hydration and sex and the time of year can influence blood values. The lungs lack lobes. A dehydrated camel has a lower breathing rate. Each kidney has a capacity of , and can produce urine with high chloride concentrations. Like the horse, the dromedary has no gall bladder. The grayish violet, crescent-like spleen weighs less than . The triangular, four-chambered liver weighs ; its dimensions are .


Reproductive system

The Ovary, ovaries are reddish, circular and flattened. They are enclosed in a conical Bursa (anatomy), bursa and have the dimensions during anestrus. The oviducts are long. The uterus is Bicornuate uterus, bicornuate. The vagina is long and has well-developed Bartholin's glands. The vulva is deep and has a small clitoris. The placenta is diffuse and Placentation#Placentation in mammals, epitheliochorial, with a crescent-like chorion. The penis is covered by a triangular penile sheath that opens backwards; it is about long. The scrotum is located high in the perineum with the testicles in separate sacs. Testicles are long, deep and wide. The right testicle is often smaller than the left. The typical mass of either testicle is less than ; during the rut the mass increases from . The Cowper's gland is white, almond-shaped and lacks seminal vesicles; the prostate gland is dark yellow, disc-shaped and divided into two lobes. The camel epididymis interstitium revealed several blood vessels harboring special regulatory devices such as the spiral arteries, spiral veins, and throttle arterioles.


Health and diseases

The dromedary generally suffers from fewer diseases than other domestic livestock such as goats and cattle. Temperature fluctuations occur throughout the day in a healthy dromedary – the temperature falls to its minimum at dawn, rises until sunset and falls during the night. Nervous camels may vomit if they are carelessly handled; this does not always indicate a disorder. Rutting males may develop nausea. The dromedary is prone to trypanosomiasis, a disease caused by a parasite transmitted by the tsetse fly. The main symptoms are recurring fever, anaemia and weakness; the disease is typically fatal for the camel. Brucellosis is another prominent malady. In an observational study, the seroprevalence of this disease was generally low (2 to 5%) in nomadic or moderately free dromedaries, but it was higher (8 to 15%) in denser populations. Brucellosis is caused by different biotypes of ''Brucella abortus'' and ''Brucella melitensis, B. melitensis''. Other internal parasites include ''Fasciola gigantica'' (trematode), two types of cestode (tapeworm) and various nematodes (roundworms). Among external parasites, ''Sarcoptes'' species cause sarcoptic mange. In a 2000 study in Jordan, 83% of the 32 camels studied tested positive for sarcoptic mange. In another study, dromedaries were found to have natural antibody, antibodies against the rinderpest virus, rinderpest and ovine rinderpest viruses. In 2013, a seroepidemiological study (a study investigating the patterns, causes and effects of a disease on a specific population on the basis of serologic tests) in Egypt was the first to show the dromedary might be a host for the Middle East respiratory syndrome coronavirus (MERS-CoV). A 2013–14 study of dromedaries in Saudi Arabia concluded the unusual genetic stability of MERS-CoV coupled with its high seroprevalence in the dromedary makes this camel a highly probable host for the virus. The full genome sequence of MERS-CoV from dromedaries in this study showed a 99.9% match to the genomes of human clade B MERS-CoV. Another study in Saudi Arabia showed the presence of MERS-CoV in 90% of the evaluated dromedaries and suggested that camels could be the animal source of MERS-CoV. Fleas and ticks are common causes of physical irritation. ''Hyalomma dromedarii'' is especially adapted to arid conditions, changing its moulting process to complete more or all of its life cycle on a single host if stressed, and having an unusually wide host range. The larvae are not well understood but larval questing, their questing phase is assumed to occur during the winter, which is also when rain arrives. The nymphs infest the host mostly in January, then the adults May to September. In a study in Egypt, ''H. dromedarii'' was dominant in dromedaries, comprising 95.6% of the adult ticks isolated from the camels. In Israel, the number of ticks per camel ranged from 20 to 105. Nine camels in the date palm plantations in Arava Valley were injected with ivermectin, which is not effective against ''Hyalomma'' tick infestations. Larvae of the camel nasal fly ''Cephalopsis titillator'' can cause possibly fatal brain compression and Neurological disorder, nervous disorders. Illnesses that can affect dromedary productivity are Pus, pyogenic diseases and wound infections caused by ''Corynebacterium'' and ''Streptococcus'', pulmonary disease, pulmonary disorders caused by ''Pasteurella'' such as hemorrhagic septicemia and ''Rickettsia'' species, camelpox, anthrax, and cutaneous necrosis caused by ''Streptothrix'' and deficiency of salt in the diet.


Ecology

The dromedary is diurnality, diurnal (active mainly during daylight); free-ranging herds feed and roam throughout the day, though they rest during the hottest hours around noon. The night is mainly spent resting. Dromedaries form cohesive groups of about 20 individuals, which consist of several females led by a dominant male. Females may also lead in turns. Some males either form bachelor groups or roam alone. Herds may congregate to form associations of hundreds of camels during migrations at the time of natural disasters. The males of the herd prevent female members from interacting with bachelor males by standing or walking between them and sometimes driving the bachelor males away. In Australia, short-term home ranges of feral dromedaries cover ; annual home ranges can spread over several thousand square kilometres. Special behavioral features of the dromedary include snapping at others without biting them and showing displeasure by stamping their feet. They are generally non-aggressive, with the exception of rutting males. They appear to remember their homes; females, in particular, remember the places they first gave birth or suckled their offspring. Males become aggressive in the mating season, and sometimes wrestle. A 1980 study showed androgen levels in males influences their behavior. Between January and April when these levels are high during the rut, they become difficult to manage, blow out the palate from the mouth, vocalize and throw urine over their backs. Camels scratch parts of their bodies with their legs or with their lower incisors. They may also rub against tree bark and roll in the sand. Free-ranging dromedaries face large predators typical of their regional distribution, which includes Subspecies of Canis lupus#Eurasia and Australia, wolves, lions and tigers.


Diet

The dromedary's diet consists mostly of foliage, dry grasses and desert vegetation – mostly thorny plants. A study said the typical diet of the dromedary is dwarf shrubs (47.5%), trees (29.9%), grasses (11.2%), other herbs (0.2%) and vines (11%). The dromedary is primarily a browser; forbs and shrubs comprise 70% of its diet in summer and 90% of its diet in winter. The dromedary may also graze on tall, young, succulent grasses. In the Sahara, 332 plant species have been recorded as food plants of the dromedary. These include ''Aristida pungens'', ''Acacia tortilis'', ''Panicum turgidum'', ''Launaea arborescens'' and ''Balanites aegyptiaca''. The dromedary eats ''Acacia'', ''Atriplex'' and ''Salsola'' when they are available. Feral dromedaries in Australia prefer ''Trichodesma zeylanicum'' and ''Euphorbia tannensis''. In India, dromedaries are fed with forage plants such as ''Vigna aconitifolia'', ''Vigna mungo, V. mungo'', ''Cyamopsis tetragonolaba'', ''Melilotus parviflora'', ''Eruca sativa'', ''Trifolium'' species and ''Brassica campestris''. Dromedaries keep their mouths open while chewing thorny food. They use their lips to grasp the food and chew each bite 40 to 50 times. Its long eyelashes, eyebrows, lockable nostrils, Caudal (anatomical term)#Anterior and posterior, caudal opening of the foreskin, prepuce and a relatively small vulva help the camel avoid injuries, especially while feeding. They graze for 8–12 hours per day and ruminate for an equal amount of time.


Biology


Adaptations

The dromedary is specially adapted to its desert habitat; these adaptations are aimed at conserving water and regulating body temperature. The bushy eyebrows and the double row of eyelashes prevent sand and dust from entering the eyes during strong windstorms, and shield them from the sun's glare. The dromedary is able to close its nostrils voluntarily; this assists in water conservation. The dromedary can conserve water by reducing perspiration by fluctuating the body temperature throughout the day from . The kidneys are specialized to minimize water loss through excretion. Groups of camels avoid excess heat from the environment by pressing against each other. The dromedary can tolerate greater than 30% water loss, which is generally impossible for other mammals. In temperatures between , it needs water every 10 to 15 days. In the hottest temperatures, the dromedary takes water every four to seven days. This camel has a quick rate of rehydration and can drink at per minute. The dromedary has a rete mirabile, a complex of arteries and veins lying very close to each other which uses countercurrent blood flow to cool blood flowing to the brain. This effectively controls the temperature of the brain. The hump stores up to of fat, which the camel can break down into energy to meet its needs when resources are scarce; the hump also helps dissipate body heat. When this tissue is metabolized, through lipolysis, fat metabolization, it releases energy while causing water to evaporate from the lungs during breathing, respiration (as oxygen is required for the metabolic process): overall, there is a net decrease in water. If the hump is small, the animal can show signs of starvation. In a 2005 study, the mean volume of adipose tissues (in the external part of the hump that have cells to store lipids) is related to the dromedary's unique mechanism of food and water storage. In case of starvation, they can even eat fish and bones, and drink brackish and salty water. The hair is longer on the throat, hump and shoulders. Though the padded hooves effectively support the camel's weight on the ground, they are not suitable for walking on slippery and muddy surfaces.


Reproduction

Camels have a slow growth rate and reach sexual maturity slower than sheep or goat. The age of sexual maturity varies geographically and depends on the individual, as does the reproductive period. Both sexes might sexual maturity, mature by three to five years of age, though successful breeding could take longer. Camels are described as atypical seasonal breeders; they exhibit spermatogenesis throughout the whole year with a reduction in spermatogenesis during the nonbreeding season compared to that in the breeding season (Zayed et al., 1995). The breeding season in Egypt is during spring; the spring months. Mating occurs once a year, and peaks in the rainy season. The mating season lasts three to five months, but may last a year for older animals. During the reproductive season, males splash their urine on their tails and nether regions. To attract females they extrude their soft palate – a trait unique to the dromedary. As the male gurgles, copious quantities of saliva turns to foam and covers the mouth. Males threaten each other for Dominance (ethology), dominance over the female by trying to stand taller than the other, making low noises and a series of head movements including lowering, lifting and bending their necks backward. Males try to defeat other males by biting the opponent's legs and taking the head between his jaws. Copulation begins with foreplay; the male smells the female's genitalia and often bites her there or around her hump. The male forces the female to sit, then grasps her with his forelegs. Camelmen often aid the male insert his penis into the female's vulva. The male dromedary's ability to penetrate the female on his own is disputed, though feral populations in Australia reproduce naturally. Copulation takes from 7 to 35 minutes, averaging 11 to 15 minutes. Normally, three to four ejaculations occur. The semen of a Bikaneri dromedary is white and viscous, with a pH of around 7.8. A single calf is born after a Pregnancy (mammals), gestation period of 15 months. Calves move freely by the end of their first day. Nursing and maternal care continue for one to two years. In a study to find whether young could exist on milk substitutes, two male, month-old camels were separated from their mothers and were fed on milk substitutes prepared commercially for lambs, and they grew to normal weights for male calves after 30 days. Lactational yield can vary with species, breed, individual, region, diet, management conditions and lactating stage. The largest quantity of milk is produced during the early period of lactation. The lactation period can vary between nine and eighteen months. Dromedaries are Induced ovulation (animals), induced ovulators. Oestrus may be cued by the nutritional status of the camel and the length of day. If mating does not occur, the Ovarian follicle, follicle, which grows during oestrus, usually regresses within a few days. In one study, 35 complete oestrous cycles were observed in five nonpregnant females over 15 months. The cycles were about 28 days long; follicles matured in six days, maintained their size for 13 days, and returned to their original size in eight days. In another study, ovulation could be best induced when the follicle reaches a size of . In another study, pregnancy in females could be recognized as early as 40 to 45 days of gestation by the swelling of the left uterine horn, where 99.5% of pregnancies were located.


Range

Its range included hot, arid regions of northern Africa, Ethiopia, the Near East, and western and central Asia. The dromedary typically thrives in areas with a long dry season and a short wet season. They are sensitive to cold and humidity, though some breeds can thrive in humid conditions. The dromedary may have been first domesticated in Somalia or the Arabian Peninsula about 4,000 years ago. In the ninth or tenth century BC, the dromedary became popular in the Near East. The Persian invasion of Egypt under Cambyses II, Cambyses in 525 BC introduced domesticated camels to the area. The Persian camels were not well-suited to trading or travel over the Sahara; journeys across the desert were made on chariots pulled by horses. The dromedary was introduced into Egypt from south-western Asia (Arabia and Persia). The popularity of dromedaries increased after the Islamic conquest of North Africa. While the invasion was accomplished largely on horseback, new links to the Middle East allowed camels to be imported ''en masse''. These camels were well-suited to long desert journeys and could carry a great deal of cargo, allowing substantial trans-Saharan trade for the first time. In Libya, dromedaries were used for transport and their milk and meat constituted the local diet. Dromedaries were also shipped from south-western Asia to Spain, Italy, Turkey, France, Canary Islands, the Americas and Australia. Dromedaries were introduced into Spain in 1020 AD and to Sicily in 1059 AD. Camels were exported to the Canary Islands in 1405 during the European colonisation of the area, and are still extant there, especially in Lanzarote and to the south of Fuerteventura. Attempts to introduce dromedaries into the Caribbean, Colombia, Peru, Bolivia and Brazil were made between the 17th and 19th centuries; some were imported to the western United States in the 1850s and some to Namibia in the early 1900s, but presently they exist in small numbers or are absent in these areas. In 1840, about six camels were shipped from Tenerife to Adelaide, but only one survived the journey to arrive on 12 October that year. The animal, a male called Harry, was owned by the explorer John Ainsworth Horrocks. Harry was ill-tempered but was included in an expedition the following year because he could carry heavy loads. The next major group of camels were imported into Australia in 1860, and between 1860 and 1907 10 to 12 thousand were imported. These were used mainly for riding and transport.


Current distribution of captive animals

In the early 21st century, the domesticated dromedary is found in the semi-arid to arid regions of the Old World.


Africa

Africa has more than 80% of the world's total dromedary population; it occurs in almost every desert zone in the northern part of the continent. The Sahel marks the southern extreme of its range, where the annual rainfall is around . The Horn of Africa has nearly 35% of the world's dromedaries; most of the region's stock is in
Somalia Somalia, , Osmanya script: 𐒈𐒝𐒑𐒛𐒐𐒘𐒕𐒖; ar, الصومال, aṣ-Ṣūmāl officially the Federal Republic of SomaliaThe ''Federal Republic of Somalia'' is the country's name per Article 1 of thProvisional Constituti ...
, followed by Sudan, Eritrea, and Ethiopia (as of the early 2000s). According to the Yearbook of the Food and Agriculture Organization (FAO) for 1984, eastern Africa had about 10 million dromedaries, the largest population of Africa. Western Africa followed with 2.14 million, while northern Africa had nearly 0.76 million. Populations in Africa increased by 16% from 1994 to 2005.


Asia

In Asia, nearly 70% of the population occurs in India and Pakistan. The combined population of the dromedary and the Bactrian camel decreased by around 21% between 1994 and 2004. The dromedary is sympatric with the Bactrian camel in Afghanistan, Pakistan, and central and southwestern Asia. India has a dromedary population of less than one million, with most (0.67 million) in the state of Rajasthan. Populations in Pakistan decreased from 1.1 million in 1994 to 0.8 million in 2005 – a 29% decline. According to the FAO, the dromedary population in six countries of the Persian Gulf was nearly 0.67 million in 2003. In the Persian Gulf region the dromedary is locally classified into breeds including Al-Majahem, Al-Hamrah, Al-Safrah, Al-Zarkah and Al-Shakha, based on coat colour. The UAE has three prominent breeds: Racing camel, Al-Arabiat and Al-Kazmiat.


Feral population

Feral dromedary populations occur in Australia, where they were introduced in 1840. The total dromedary population in Australia was 500,000 in 2005. Nearly 99% of the populations are feral, and they have annual growth rate of 10%. Most of the Australian feral camels are dromedaries, with only a few Bactrian camels. Most of the dromedaries occur in Western Australia, with smaller populations in the Northern Territory, Western Queensland and northern South Australia.


Relationship with humans

The strength and docility of the dromedary make it popular as a domesticated animal. According to
Richard Bulliet Richard W. Bulliet (born 1940) is a professor of history at Columbia University who specializes in the history of Islamic society and institutions, the history of technology, and the history of the role of animals in human society. Early life ...
, they can be used for a wide variety of purposes: riding, transport, ploughing, and trading and as a source of milk, meat, wool and leather. The main attraction of the dromedary for nomadic desert-dwellers is the wide variety of resources they provide, which are crucial for their survival. It is important for several Bedouin pastoralist tribes of northern Arabia, such as the Ruwallah, the Rashaida, the Bani Sakhr and the Mutayr. Camel urine and camel milk is used for Prophetic medicine, medicinal purposes.


Riding camels

Although the role of the camel is diminishing with the advent of technology and modern means of transport, it is still an efficient mode of communication in remote and less-developed areas. The dromedary has been used in warfare since the 2nd century BC. It is particularly prized for its capability to outrun horses in the deserts. Record of its use during the time of Alexander the Great indicate that the animal could cover up to 50 miles per day for a week and they could go for up to a month without water. An account by Aurelian also cited that, in her escape to Euphrates, Zenobia used a dromedary to outrun her pursuers after she was defeated at Palmyra. The dromedary also remains popular for racing, particularly in the Arab world. Riding camels of Arabia, Egypt and the Sahara are locally known as the Dilool, the Hageen, and the Mehara respectively; several local breeds are included within these groups. The ideal riding camel is strong, slender and long-legged with thin, supple skin. The special adaptations of the dromedary's feet allow it to walk with ease on sandy and rough terrain and on cold surfaces. The camels of the Beja people, Bejas of Sudan and the Hedareb, Bilen people, Bilen, and the Tigre people, Tigre people of Eritrea and the Anafi camel bred in Sudan are common breeds used as riding camels. According to Leese, the dromedary walks with four speeds or gaits: walk, jog, fast run and canter. The first is the typical speed of walking, around . Jog is the most common speed, nearly on level ground. He estimated a speed of during a fast run, by observing northern African and Arabian dromedaries. He gave no speed range to describe the Canter and gallop, canter, but implied it was a type of gallop that if induced could exhaust the camel and the rider. Canter could be used only for short periods of time, for example in races. The ideal age to start training dromedaries for riding is three years, although they may be stubborn and unruly. At first the camel's head is controlled, and it is later trained to respond to sitting and standing commands, and to allow mounting. At this stage a camel will often try to escape when a trainer tries to mount it. The next stage involves training it to respond to reins. The animal must be given loads gradually and not forced to carry heavy loads before the age of six. Riding camels should not be struck on their necks, rather they should be struck behind the right leg of the rider. Leese described two types of saddles generally used in camel riding: the Arabian ''markloofa'' used by single riders and the Indian ''pakra'' used when two riders mount the same camel.


Baggage and draught camels

The baggage camel should be robust and heavy. Studies have recommended the camel should have either a small or a large head with a narrow aquiline nose, prominent eyes and large lips. The neck should be medium to long so the head is held high. The chest should be deep and the hump should be well-developed with sufficient space behind it to accommodate the saddle. The hindlegs should be heavy, muscular and sturdy. The dromedary can be trained to carry baggage from the age of five years, but must not be given heavy loads before the age of six. The ''hawia'' is a typical baggage saddle from Sudan. The methods of training the baggage camels are similar to those for riding camels. Draught camels are used for several purposes including ploughing, processing in oil mills and pulling carts. There is no clear description for the ideal draught camel, though its strength, its ability to survive without water and the flatness of its feet could be indicators. It may be used for ploughing in pairs or in groups with buffaloes or bullocks. The draught camel can plough at around , and should not be used for more than six hours a day – four hours in the morning and two in the afternoon. The camel is not easily exhausted unless diseased or undernourished, and has remarkable endurance and hardiness.


Dairy products

Camel milk is a staple food of nomadic tribes living in deserts. It consists of 11.7% solids, 3% protein, 3.6% fat, 0.8% Ash (analytical chemistry), ash, 4.4% lactose and 0.13% acidity (pH 6.5). The quantities of sodium, potassium, zinc, iron, copper, manganese, Niacin (substance), niacin and vitamin C were relatively higher than the amounts in cow milk. However, the levels of thiamin, riboflavin, folacin, Vitamin B12, vitamin B12, pantothenic acid, vitamin A, lysine, and tryptophan were lower than those in cow milk. The mole (unit), molar percentages of the fatty acids in milk fat were 26.7% for palmitic acid, 25.5% oleic acid, 11.4% myristic acid and 11% palmitoleic acid. Camel milk has higher thermal stability compared with cow milk, but it does not compare favourably with sheep milk. Daily milk yield generally varies from and from 1.3% to 7.8% of the body weight. Milk yield varies geographically and depends upon the animals' diet and living conditions. At the peak of lactation, a healthy female would typically provide milk per day. Leese estimated a lactating female would yield besides the amount ingested by the calf. The Pakistani dromedary, which is considered a better milker and bigger, can yield when well-fed. Dromedaries in Somalia may be milked between two and four times a day, while those in Afar Region, Afar, Ethiopia, may be milked up to seven times a day. The acidity of dromedary milk stored at increases at a slower rate than that of cow milk. Though the preparation of butter from dromedary milk is difficult, it is produced in small amounts by nomads, optimized at 22.5% fat in the cream. In 2001, the ability of dromedary milk to form curd was studied; coagulation did not show curd formation, and had a pH of 4.4. It was much different from curd produced from cow milk, and had a fragile, heterogeneous composition probably composed of casein flakes. Nevertheless, cheese and other dairy products can be made from camel milk. A study found bovine calf rennet could be used to coagulate dromedary milk. A special factory has been set up in Nouakchott to pasteurise and make cheese from camel milk. Mystical beliefs surround the use of camel milk in some places; for example, it may be used as an aphrodisiac in Ethiopia.


Meat

The meat of a five-year-old dromedary has a typical composition of 76% water, 22% protein, 1% fat, and 1% ash. The carcass, weighing for a five-year-old dromedary, is composed of nearly 57% muscle, 26% bone and 17% fat. A seven-to-eight-year-old camel can produce a carcass of . The meat is bright red to a dark brown or maroon, while the fat is white. It has the taste and texture of beef. A study of the meat of Iranian dromedaries showed its high glycogen content, which makes it taste sweet like horse meat. The carcasses of well-fed camels were found to be covered with a thin layer of good quality fat. In a study of the fatty acid composition of raw meat taken from the hind legs of seven one-to-three years old males, 51.5% of the fatty acids were saturated, 29.9% mono-unsaturated, and 18.6% polyunsaturated. The major fatty acids in the meat were palmitic acid (26.0%), oleic acid (18.9%) and linoleic acid (12.1%). In the hump, palmitic acid was dominant (34.4%), followed by oleic acid (28.2%), myristic acid (10.3%) and stearic acid (10%). Dromedary slaughter is more difficult than the slaughter of other domestic livestock such as cattle because of the size of the animal and the significant manual work involved. More males than females are slaughtered. Though less affected by mishandling than other livestock, the pre-slaughter handling of the dromedary plays a crucial role in determining the quality of meat obtained; mishandling can often disfigure the hump. The animal is stunned, seated in a crouching position with the head in a caudal position and slaughtered. The dressing percentage – the percentage of the mass of the animal that forms the carcass – is 55–70%, more than the 45–50% of cattle. Camel meat is often eaten by African camel herders, who use it only during severe food scarcity or for rituals. Camel meat is processed into food items such as burgers, patties, sausages and ''shawarma''. Dromedaries can be slaughtered between four and ten years of age. As the animal ages, the meat grows tougher and deteriorates in taste and quality. In Somalian and Djiboutian culture, the dromedary is a staple food and can be found in many recipes and dishes. A 2005 report issued jointly by the Ministry of Health (Saudi Arabia) and the United States Centers for Disease Control and Prevention details five cases of bubonic plague in humans resulting from the ingestion of raw camel liver. Four of the five patients had severe pharyngitis and submandibular lymphadenitis. ''Yersinia pestis'' was isolated from the camel's bone marrow, from the jird (''Meriones libycus'') and from fleas (''Xenopsylla cheopis'') captured at the camel's corral.


Camel hair, wool and hides

Camels in hot climates generally do not develop long coats. Camel hair is light, and has low thermal conductivity and durability, and is thus suitable for manufacturing warm clothes, blankets, tents, and rugs. Hair of highest quality is typically obtained from juvenile or wild camels. In India, camels are clipped usually in spring and around hair is produced per clipping. In colder regions one clipping can yield as much as . A dromedary can produce wool per year, whereas a Bactrian camel has an annual yield of nearly . Dromedaries under the age of two years have a fine undercoat that tends to fall off and should be cropped by hand. Little information about camel hides has been collected but they are usually of inferior quality and are less preferred for manufacturing leather.


See also

* Camel urine * List of animals with humps


References


External links

*
Could
United Arab Emirates, Emirati camels hold the key to Antivenom, treating venomous snake bites? {{Authority control Camels Mammals of Africa Mammals of Asia Mammals of Pakistan Fauna of Iran Fauna of the Middle East Fauna of Jordan Fauna of Syria Fauna of Iraq Mammals of the Arabian Peninsula Fauna of the Sahara Mammals of Somalia Livestock Pack animals Mammals described in 1758 Mammals of North Africa Taxa named by Carl Linnaeus Symbols of Rajasthan