Crassadoma
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''Crassadoma'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of rock scallops, marine
bivalve Bivalvia (), in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class of marine and freshwater molluscs that have laterally compressed bodies enclosed by a shell consisting of two hinged parts. As a group, bival ...
molluscs in the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Pectinidae Scallop () is a common name that encompasses various species of marine bivalve mollusks in the taxonomic family Pectinidae, the scallops. However, the common name "scallop" is also sometimes applied to species in other closely related families ...
. It is
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...
, the only
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
being ''Crassadoma gigantea'', the rock scallop, giant rock scallop or purple-hinge rock scallop. Although the small juveniles are free-swimming, they soon become
sessile Sessility, or sessile, may refer to: * Sessility (motility), organisms which are not able to move about * Sessility (botany), flowers or leaves that grow directly from the stem or peduncle of a plant * Sessility (medicine), tumors and polyps that ...
, and are cemented to the substrate. These scallops occur in the eastern
Pacific Ocean The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the continen ...
.


Description

Adults of ''Crassadoma gigantea'' have one valve (the right valve) fixed to a base, often a vertical rock face. This valve often becomes deformed to fit the contours of the rock. The left valve is roughly circular but irregular in outline, thick and heavy, with deep radial ribs. Every third or fourth rib is clad with blunt spines that overhang each other. These often get abraded and worn smooth as the scallop ages. There are flaps of shell known as auricles on either side of the straight hinge. The general colour is brown with the inside of the shell being glossy white with a purple patch at the hinge and a large, central adductor muscle scar. ''Crassadoma gigantea'' grows to a diameter of in the
intertidal zone The intertidal zone, also known as the foreshore, is the area above water level at low tide and underwater at high tide (in other words, the area within the tidal range). This area can include several types of habitats with various species o ...
and in the subtidal zone. Between the valves, the margin of the orange mantle can be seen, with a row of tiny blue eyes and a sparse fringe of short tentacles.''Crassadoma gigantea (Gray, 1838)''
Intertidal Invertebrates of the Monterey Bay Area, California. Retrieved 2012-01-31.
Subtidal Bivalves, Cephalopods, & Chitons: ''Crassadoma gigantea''
SeaNet. Retrieved 2012-01-31.


Distribution and habitat

''Crassadoma gigantea'' is found on the Pacific Coast of North America, from
British Columbia British Columbia (commonly abbreviated as BC) is the westernmost province of Canada, situated between the Pacific Ocean and the Rocky Mountains. It has a diverse geography, with rugged landscapes that include rocky coastlines, sandy beaches, ...
south to
Baja California Baja California (; 'Lower California'), officially the Free and Sovereign State of Baja California ( es, Estado Libre y Soberano de Baja California), is a state in Mexico. It is the northernmost and westernmost of the 32 federal entities of Mex ...
and
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
. It does not move around as an adult but lives on the sea floor. Its favoured habitats include inside crevices and under boulders, or cemented to rock surfaces, corals or man-made structures such as harbour
pilings A deep foundation is a type of foundation that transfers building loads to the earth farther down from the surface than a shallow foundation does to a subsurface layer or a range of depths. A pile or piling is a vertical structural element ...
. It is found at depths down to about .''Crassadoma gigantea'' (Purple hinged rock scallop)
Beach Watchers. Retrieved 2012-02-01.


Biology

Like other scallops, ''Crassadoma gigantea'' is a
filter feeder Filter feeders are a sub-group of suspension feeding animals that feed by straining suspended matter and food particles from water, typically by passing the water over a specialized filtering structure. Some animals that use this method of feedin ...
, filtering
phytoplankton Phytoplankton () are the autotrophic (self-feeding) components of the plankton community and a key part of ocean and freshwater ecosystems. The name comes from the Greek words (), meaning 'plant', and (), meaning 'wanderer' or 'drifter'. Ph ...
from water as it passes over the gills. The particles are moved by
cilia The cilium, plural cilia (), is a membrane-bound organelle found on most types of eukaryotic cell, and certain microorganisms known as ciliates. Cilia are absent in bacteria and archaea. The cilium has the shape of a slender threadlike projecti ...
along grooves to the mouth where edible matter is separated from sediment particles. The waste is incorporated into
mucous Mucus ( ) is a slippery aqueous secretion produced by, and covering, mucous membranes. It is typically produced from cells found in mucous glands, although it may also originate from mixed glands, which contain both serous and mucous cells. It is ...
balls which are removed from the mantle cavity periodically by a clapping of the valves.Learn about scallops
A snail's odyssey. Retrieved 2012-02-01.
''Crassadoma gigantea'' is cryptic and difficult to detect because of the
sponge Sponges, the members of the phylum Porifera (; meaning 'pore bearer'), are a basal animal clade as a sister of the diploblasts. They are multicellular organisms that have bodies full of pores and channels allowing water to circulate through t ...
s,
sea anemone Sea anemones are a group of predation, predatory marine invertebrates of the order (biology), order Actiniaria. Because of their colourful appearance, they are named after the ''Anemone'', a terrestrial flowering plant. Sea anemones are classifi ...
s,
hydroids Hydroids are a life stage for most animals of the class Hydrozoa, small predators related to jellyfish. Some hydroids such as the freshwater '' Hydra'' are solitary, with the polyp attached directly to the substrate. When these produce buds ...
,
barnacle A barnacle is a type of arthropod constituting the subclass Cirripedia in the subphylum Crustacea, and is hence related to crabs and lobsters. Barnacles are exclusively marine, and tend to live in shallow and tidal waters, typically in eros ...
s, bryozoans, worms and
algae Algae (; singular alga ) is an informal term for a large and diverse group of photosynthetic eukaryotic organisms. It is a polyphyletic grouping that includes species from multiple distinct clades. Included organisms range from unicellular mic ...
which tend to grow on the shell. The
parasitic Parasitism is a close relationship between species, where one organism, the parasite, lives on or inside another organism, the host, causing it some harm, and is adapted structurally to this way of life. The entomologist E. O. Wilson has c ...
boring sponge ''
Cliona celata ''Cliona celata'', occasionally called the boring sponge, is a species of demosponge belonging the family Clionaidae.Hansson, H.G. (Comp.), 1999. NEAT (North East Atlantic Taxa): Scandinavian marine Porifera (Spongiaria) Check- List. Internet p ...
'' damages the shell by making hollows in it. Additional shell material is laid down by the mantle tissue to compensate for this. Other sponges, such as the yellow boring sponge (''Cliona californiana'') that grow on the shell may be considered mutualistic, granting some protection against
predators Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill th ...
such as the sunflower starfish (''Pycnopodia helianthoides'') and purple sea star (''Pisaster ochraceus''). It has been found experimentally that rock scallops grow at a faster rate when living in deep rather than shallow water, perhaps because they are then less hindered by these fouling organisms. The sexes are separate in ''Crassadoma gigantea'' and
spawning Spawn is the eggs and sperm released or deposited into water by aquatic animals. As a verb, ''to spawn'' refers to the process of releasing the eggs and sperm, and the act of both sexes is called spawning. Most aquatic animals, except for aquati ...
takes place in the summer. The
veliger A veliger is the planktonic larva of many kinds of sea snails and freshwater snails, as well as most bivalve molluscs (clams) and tusk shells. Description The veliger is the characteristic larva of the gastropod, bivalve and scaphopod ...
larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. The ...
e that develop from the
eggs Humans and human ancestors have scavenged and eaten animal eggs for millions of years. Humans in Southeast Asia had domesticated chickens and harvested their eggs for food by 1,500 BCE. The most widely consumed eggs are those of fowl, especial ...
form part of the
zooplankton Zooplankton are the animal component of the planktonic community ("zoo" comes from the Greek word for ''animal''). Plankton are aquatic organisms that are unable to swim effectively against currents, and consequently drift or are carried along by ...
for about 40 days before settling, undergoing
metamorphosis Metamorphosis is a biological process by which an animal physically develops including birth or hatching, involving a conspicuous and relatively abrupt change in the animal's body structure through cell growth and differentiation. Some inse ...
and beginning to form a shell. Juveniles are free living, are able to swim and can attach themselves temporarily to the substrate by
byssus A byssus () is a bundle of filaments secreted by many species of bivalve mollusc that function to attach the mollusc to a solid surface. Species from several families of clams have a byssus, including pen shells (Pinnidae), true mussels (Mytilid ...
threads. By the time they have grown to in diameter they have usually cemented themselves to a hard surface and become
sessile Sessility, or sessile, may refer to: * Sessility (motility), organisms which are not able to move about * Sessility (botany), flowers or leaves that grow directly from the stem or peduncle of a plant * Sessility (medicine), tumors and polyps that ...
. Rock scallops can live for at least 20 years but by about half way through their adult life, growth slows, and energy is concentrated into
gamete A gamete (; , ultimately ) is a haploid cell that fuses with another haploid cell during fertilization in organisms that reproduce sexually. Gametes are an organism's reproductive cells, also referred to as sex cells. In species that produce t ...
production.


References

{{Taxonbar, from1=Q19953521, from2=Q3013967 Pectinidae Bivalve genera Monotypic mollusc genera