Brachysuchus
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''Brachysuchus'' (meaning "wide crocodile") is an extinct
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of
phytosaur Phytosaurs (Φυτόσαυροι in greek) are an extinct group of large, mostly semiaquatic Late Triassic archosauriform reptiles. Phytosaurs belong to the order Phytosauria. Phytosauria and Phytosauridae are often considered to be equivalent g ...
known from the late
Triassic The Triassic ( ) is a geologic period and system (stratigraphy), system which spans 50.6 million years from the end of the Permian Period 251.902 million years ago (Year#Abbreviations yr and ya, Mya), to the beginning of the Jurassic Period 201.36 ...
period Period may refer to: Common uses * Era, a length or span of time * Full stop (or period), a punctuation mark Arts, entertainment, and media * Period (music), a concept in musical composition * Periodic sentence (or rhetorical period), a concept ...
(
Carnian The Carnian (less commonly, Karnian) is the lowermost stage (stratigraphy), stage of the Upper Triassic series (stratigraphy), Series (or earliest age (geology), age of the Late Triassic Epoch (reference date), Epoch). It lasted from 237 to 227 m ...
stage) of
Dockum Group The Dockum is a Late Triassic (approximately late Carnian through Rhaetian, or 223–200 Ma) geologic group found primarily on the Llano Estacado of western Texas and eastern New Mexico with minor exposures in southwestern Kansas, eastern Colora ...
in
Texas Texas (, ; Spanish: ''Texas'', ''Tejas'') is a state in the South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2020, it is the second-largest U.S. state by ...
, United States. It is known from the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of sever ...
UMMP 10336 is composed of a
skull The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, th ...
, lower
jaw The jaw is any opposable articulated structure at the entrance of the mouth, typically used for grasping and manipulating food. The term ''jaws'' is also broadly applied to the whole of the structures constituting the vault of the mouth and serv ...
s and partial
postcranium Postcrania (postcranium, adjective: postcranial) in zoology and vertebrate paleontology is all or part of the skeleton apart from the skull. Frequently, fossil remains, e.g. of dinosaurs or other extinct tetrapods, consist of partial or isolated s ...
and from the associated paratype UMMP 14366, nearly complete skull, recovered from the 'Pre-Tecovas Horizon' in the
Dockum Group The Dockum is a Late Triassic (approximately late Carnian through Rhaetian, or 223–200 Ma) geologic group found primarily on the Llano Estacado of western Texas and eastern New Mexico with minor exposures in southwestern Kansas, eastern Colora ...
. It was first named by Case in
1929 This year marked the end of a period known in American history as the Roaring Twenties after the Wall Street Crash of 1929 ushered in a worldwide Great Depression. In the Americas, an agreement was brokered to end the Cristero War, a Catholic ...
and the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specime ...
is ''Brachysuchus megalodon''. Its closest relative was '' Angistorhinus''. However, its rostral crest was much smaller than that of ''Angistorhinus,'' and the
rostrum Rostrum may refer to: * Any kind of a platform for a speaker: **dais **pulpit * Rostrum (anatomy), a beak, or anatomical structure resembling a beak, as in the mouthparts of many sucking insects * Rostrum (ship), a form of bow on naval ships * Ros ...
as a whole is shorter and thicker.


Features

The lower jaws of ''Brachysuchus'' are expanded at the tip to form a large bulge, holding the creature's largest tusks. The surface of this part of the
bone A bone is a rigid organ that constitutes part of the skeleton in most vertebrate animals. Bones protect the various other organs of the body, produce red and white blood cells, store minerals, provide structure and support for the body, ...
has a wrinkled look, with many
blood vessel The blood vessels are the components of the circulatory system that transport blood throughout the human body. These vessels transport blood cells, nutrients, and oxygen to the tissues of the body. They also take waste and carbon dioxide away ...
s running through it. In the lower jaw, there are three tusks in each side of this protuberance. Behind this part the jaws have fused together for a little under half their length before diverging after thirty-one of forty-six post-protuberance teeth. While some of the tusks at the tip are missing, most of the rest of the teeth in the jaw are still present. There are new tusks forming in the sockets, indicating that ''Brachysuchus'' regrew teeth all its life. There is a large
foramen In anatomy and osteology, a foramen (;Entry "foramen"
in
between the
dentary In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower teeth in place. The mandible sits beneath the maxilla. It is the only movable bone ...
, the angular and the prearticular that passes right through the jaw (visible on the picture). The teeth themselves are
asymmetrical Asymmetry is the absence of, or a violation of, symmetry (the property of an object being invariant to a transformation, such as reflection). Symmetry is an important property of both physical and abstract systems and it may be displayed in pre ...
in cross-section, with the outer side more
convex Convex or convexity may refer to: Science and technology * Convex lens, in optics Mathematics * Convex set, containing the whole line segment that joins points ** Convex polygon, a polygon which encloses a convex set of points ** Convex polytop ...
than the inner side. This asymmetry increases from front to back. They are long and conical, and those from the ninth to twenty-ninth have flutings on the sides. From the thirtieth to the forty-ninth, they have no flutings but are crenulated and broaden antero-posteriorly. This suggests that these teeth were better for slicing
prey Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill ...
than the previous tusks and conical teeth, which would have impaled and injured it. In parts of the jaws the fused section in the middle rises so high that the teeth could only grip prey and could probably have done little damage. The prearticular is fused with the
articular The articular bone is part of the lower jaw of most vertebrates, including most jawed fish, amphibians, birds and various kinds of reptiles, as well as ancestral mammals. Anatomy In most vertebrates, the articular bone is connected to two oth ...
. The skull is around 125 cm long, with a distinctive hooked tip. It is much shorter and broader than that of many phytosaurs such as ''
Leptosuchus ''Leptosuchus'' is an extinct genus of leptosuchomorph phytosaur with a complex taxonomical history. Fossils have been found from the Dockum Group and lower Chinle Formation outcropping in Texas, New Mexico, and Arizona, USA, and date back to th ...
'' or ''Angistorhinus''. The
eye sockets In anatomy, the orbit is the cavity or socket of the skull in which the eye and its appendages are situated. "Orbit" can refer to the bony socket, or it can also be used to imply the contents. In the adult human, the volume of the orbit is , o ...
are narrow and long, and the fenestrae are smaller and less rounded than ''Angistorhinus.'' It has a comparatively small rostral crest. There are several rough surfaces at the posterior end denoting probable muscle attachments, which show that it had very powerful jaw muscles and a strong bite. Overall ''Brachysuchus'' was approximately eight metres long, much bigger than most contemporary phytosaurs. It had extremely heavy jaws and a relatively short fused region, giving it a wider bite. There is also evidence of heavily developed jaw muscles. This, as well as the cutting teeth mentioned earlier, indicates that ''Brachysuchus'' preyed on different animals to its thinner-jawed relatives. Instead of using its jaws to catch
fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of ...
, it was able to attack and feed off the heavily
armoured Armour (British English) or armor (American English; see spelling differences) is a covering used to protect an object, individual, or vehicle from physical injury or damage, especially direct contact weapons or projectiles during combat, or f ...
stegocephalians that roamed the surrounding area, and would probably have eaten other phytosaurs if food was scarce. The slightly deformed tip of one specimen's lower jaw suggests that it used a lot of force to break through its prey's armour.


References

{{Taxonbar, from=Q4953588 Phytosaurs Prehistoric reptile genera Carnian genera Late Triassic reptiles of North America Triassic geology of Texas Paleontology in Texas Fossil taxa described in 1929 Taxa named by Ermine Cowles Case