Archaeovenator hamiltonensis
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''Archaeovenator'' is an extinct
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of
Late Carboniferous Late may refer to: * LATE, an acronym which could stand for: ** Limbic-predominant age-related TDP-43 encephalopathy, a proposed form of dementia ** Local-authority trading enterprise, a New Zealand business law ** Local average treatment effect, ...
varanopid Varanopidae is an extinct family of amniotes that resembled monitor lizards and may have filled a similar niche, hence the name. Typically, they are considered synapsids that evolved from an '' Archaeothyris''-like synapsid in the Late Carbonifer ...
synapsid Synapsids + (, 'arch') > () "having a fused arch"; synonymous with ''theropsids'' (Greek, "beast-face") are one of the two major groups of animals that evolved from basal amniotes, the other being the sauropsids, the group that includes reptil ...
s known from
Greenwood County Greenwood County is the name of two counties in the United States: * Greenwood County, Kansas Greenwood County (county code GW) is a county located in the southeast portion of the U.S. state of Kansas. As of the 2020 census, the county populat ...
,
Kansas Kansas () is a state in the Midwestern United States. Its capital is Topeka, and its largest city is Wichita. Kansas is a landlocked state bordered by Nebraska to the north; Missouri to the east; Oklahoma to the south; and Colorado to th ...
of the
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territori ...
. It was first named by Robert R. Reisz and
David W. Dilkes David (; , "beloved one") (traditional spelling), , ''Dāwūd''; grc-koi, Δαυΐδ, Dauíd; la, Davidus, David; gez , ዳዊት, ''Dawit''; xcl, Դաւիթ, ''Dawitʿ''; cu, Давíдъ, ''Davidŭ''; possibly meaning "beloved one". w ...
in 2003 and the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specime ...
is ''Archaeovenator hamiltonensis''. ''Archaeovenator hamiltonensis'' is known from the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of sever ...
KUVP The University of Kansas Natural History Museum is part of the University of Kansas Biodiversity Institute, a KU designated research center dedicated to the study of the life of the planet. The museum's galleries are in Dyche Hall on the unive ...
 12483, a three-dimensionally preserved, nearly complete and articulated skeleton, including the
skull The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, th ...
, with
limb Limb may refer to: Science and technology *Limb (anatomy), an appendage of a human or animal *Limb, a large or main branch of a tree *Limb, in astronomy, the curved edge of the apparent disk of a celestial body, e.g. lunar limb *Limb, in botany, ...
s and girdles slightly separated from
postcrania Postcrania (postcranium, adjective: postcranial) in zoology and vertebrate paleontology is all or part of the skeleton apart from the skull. Frequently, fossil remains, e.g. of dinosaurs or other extinct tetrapods, consist of partial or isolated sk ...
l skeleton. It was collected in the Hamilton Quarry, from the Calhouns Shale Formation of the Shawnee Group, dating to the Virgilian stage (or alternatively late
Kasimovian The Kasimovian is a geochronologic age or chronostratigraphic stage in the ICS geologic timescale. It is the third stage in the Pennsylvanian (late Carboniferous), lasting from to Ma.; 2004: ''A Geologic Time Scale 2004'', Cambridge Univer ...
to early
Gzhelian The Gzhelian ( ) is an age in the ICS geologic timescale or a stage in the stratigraphic column. It is the youngest stage of the Pennsylvanian, the youngest subsystem of the Carboniferous. The Gzhelian lasted from to Ma. It follows the Ka ...
stage) of the Late Pennsylvanian
Series Series may refer to: People with the name * Caroline Series (born 1951), English mathematician, daughter of George Series * George Series (1920–1995), English physicist Arts, entertainment, and media Music * Series, the ordered sets used in ...
, about 300
million years ago The abbreviation Myr, "million years", is a unit of a quantity of (i.e. ) years, or 31.556926 teraseconds. Usage Myr (million years) is in common use in fields such as Earth science and cosmology. Myr is also used with Mya (million years ago) ...
. The generic name is derived from the
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
''Archaeo'' and ''venator'', meaning "ancient hunter". The specific name is named after its finding place
Hamilton Quarry Hamilton Quarry is a Late Carboniferous lagerstätte near Hamilton, Kansas, United States. It has a diverse assemblage of unusually well-preserved marine, euryhaline, freshwater, flying, and terrestrial fossils (invertebrates, vertebrates, and pl ...
. ''Archaeovenator'' is the oldest and the basalmost known varanopid, as it is the sister taxon to all other known varanopsids.


References

Varanopids Prehistoric synapsid genera Carboniferous synapsids Fossil taxa described in 2003 Carboniferous synapsids of North America Taxa named by Robert R. Reisz {{Paleo-synapsid-stub