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Agaricaceae
The Agaricaceae are a family of basidiomycete fungi and include the genus ''Agaricus'', as well as basidiomycetes previously classified in the families Tulostomataceae, Lepiotaceae, and Lycoperdaceae. Taxonomy The family Agaricaceae was published by French botanist François Fulgis Chevallier in 1826. It is named after the type genus ''Agaricus'', originally circumscribed by Carl Linnaeus in his 1753 work ''Species Plantarum''. In his authoritative 1986 classification of the Agaricales, Rolf Singer divided the Agaricaceae into four tribes distinguished largely by spore color: ''Leucocoprineae'', ''Agariceae'', ''Lepioteae'', and ''Cystodermateae''. Genera once classified in the families Tulostomataceae, Battarreaceae, Lycoperdaceae, and Mycenastraceae have since been moved to the Agaricaceae based on molecular phylogenetics studies. According to a standard reference text, the Agaricaceae contains 85 genera and 1340 species. Description Agaricaceae species use a wide variety of ...
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Agaricus
''Agaricus'' is a genus of mushrooms containing both edible and poisonous species, with over 400 members worldwide and possibly again as many disputed or newly-discovered species. The genus includes the common ("button") mushroom ('' Agaricus bisporus'') and the field mushroom ('' A. campestris''), the dominant cultivated mushrooms of the West. Members of ''Agaricus'' are characterized by having a fleshy cap or pileus, from the underside of which grow a number of radiating plates or gills, on which are produced the naked spores. They are distinguished from other members of their family, Agaricaceae, by their chocolate-brown spores. Members of ''Agaricus'' also have a stem or stipe, which elevates it above the object on which the mushroom grows, or substrate, and a partial veil, which protects the developing gills and later forms a ring or annulus on the stalk. The genus contains the most widely consumed and best-known mushroom today, '' A. bisporus'', with '' A. arvensis'' ...
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Agaricales
The fungal order Agaricales, also known as gilled mushrooms (for their distinctive gills) or euagarics, contains some of the most familiar types of mushrooms. The order has 33 extant families, 413 genera, and over 13,000 described species, along with six extinct genera known only from the fossil record. They range from the ubiquitous common mushroom to the deadly destroying angel and the hallucinogenic fly agaric to the bioluminescent jack-o-lantern mushroom. History, classification and phylogeny In his three volumes of ''Systema Mycologicum'' published between 1821 and 1832, Elias Fries put almost all of the fleshy, gill-forming mushrooms in the genus ''Agaricus''. He organized the large genus into "tribes", the names of many of which still exist as common genera of today. Fries later elevated several of these tribes to generic level, but later authors—including Gillet, Karsten, Kummer, Quélet, and Staude—made most of the changes. Fries based his classification on ...
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Secotioid
Secotioid fungi are an intermediate growth form between mushroom-like hymenomycetes and closed bag-shaped gasteromycetes, where an evolutionary process of gasteromycetation has started but not run to completion. Secotioid fungi may or may not have opening caps, but in any case they often lack the vertical geotropic orientation of the hymenophore needed to allow the spores to be dispersed by wind, and the basidiospores are not forcibly discharged or otherwise prevented from being dispersed (e.g. gills completely inclosed and never exposed as in the secotioid form of ''Lentinus tigrinus'')—note—some mycologists do not consider a species to be secotioid unless it has lost ballistospory. Explanation of secotioid development and gasteromycetation Historically agarics and boletes (which bear their spores on a hymenium of gills or tubes respectively) were classified quite separately from the gasteroid fungi, such as puff-balls and truffles, of which the spores are formed in ...
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Rolf Singer
Rolf Singer (June 23, 1906 – January 18, 1994) was a German-born mycologist and one of the most important taxonomists of gilled mushrooms ( agarics) in the 20th century. After receiving his Ph.D. at the University of Vienna in 1931 he worked in Munich. By 1933, however, Singer left Germany for Vienna due to the political deterioration in Germany. There he met his wife, Martha Singer. From Vienna, Singer and his wife went to Barcelona, Spain, where Singer was appointed assistant professor at the Autonomous University of Barcelona. Persecution by the Spanish authorities on behalf of the German government forced Singer to leave Spain for France in 1934. After a fellowship at the Museum d'Histoire Naturelle in Paris, Singer again moved, this time to Leningrad, where he was Senior Scientific Expert at the Botanical Garden of the Academy of Sciences of the USSR. During his time at the Academy, Singer made many expeditions to Siberia, the Altai Mountains, and Karelia. In 1941, Sin ...
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Agaricus Campestris
''Agaricus campestris'' is a widely eaten gilled mushroom closely related to the cultivated button mushroom ''Agaricus bisporus''. It is commonly known as the field mushroom or, in North America, meadow mushroom. Taxonomy This species was originally noted and named in 1753 by Carl Linnaeus as ''Agaricus campestris''. It was placed in the genus ''Psalliota'' by Lucien Quelet in 1872. Some variants have been isolated over the years, a few of which now have species status, for example, '' Agaricus bernardii'' Quel. (1878), ''Agaricus bisporus'' (J.E. Lange) Imbach (1946), ''Agaricus bitorquis'' (Quel.) Sacc. (1887), ''Agaricus cappellianus'' Hlavacek (1987), and ''Agaricus silvicola'' (Vittad.) Peck (1872). Some were so similar they did not warrant even varietal status, while others have retained it. ''Agaricus campestris'' var. ''equestris'' (F.H.Moller) Pilat (1951) is still valid. ''A. campestris'' var. ''isabellinus'' (F.H.Moller) Pilat (1951), and ''A. campestris' ...
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Pileus (mycology)
The pileus is the technical name for the cap, or cap-like part, of a basidiocarp or ascocarp ( fungal fruiting body) that supports a spore-bearing surface, the hymenium.Moore-Landecker, E: "Fundamentals of the Fungi", page 560. Prentice Hall, 1972. The hymenium (hymenophore) may consist of lamellae, tubes, or teeth, on the underside of the pileus. A pileus is characteristic of agarics, boletes, some polypores, tooth fungi, and some ascomycetes. Classification Pilei can be formed in various shapes, and the shapes can change over the course of the developmental cycle of a fungus. The most familiar pileus shape is hemispherical or ''convex.'' Convex pilei often continue to expand as they mature until they become flat. Many well-known species have a convex pileus, including the button mushroom, various ''Amanita The genus ''Amanita'' contains about 600 species of agarics, including some of the most toxic known mushrooms found worldwide, as well as some well-regarded edible s ...
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Stipe (mycology)
In mycology, a stipe () is the stem or stalk-like feature supporting the cap of a mushroom. Like all tissues of the mushroom other than the hymenium, the stipe is composed of sterile hyphal tissue. In many instances, however, the fertile hymenium extends down the stipe some distance. Fungi that have stipes are said to be stipitate. The evolutionary benefit of a stipe is generally considered to be in mediating spore dispersal. An elevated mushroom will more easily release its spores into wind currents or onto passing animals. Nevertheless, many mushrooms do not have stipes, including cup fungi, puffballs, earthstars, some polypores, jelly fungi, ergots, and smuts. It is often the case that features of the stipe are required to make a positive identification of a mushroom. Such distinguishing characters include: # the texture of the stipe (fibrous, brittle, chalky, leathery, firm, etc.) # whether it has remains of a partial veil (such as an annulus or cortina Cortina ...
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Gasteroid
The gasteroid fungi are a group of fungi in the Basidiomycota. Species were formerly placed in the obsolete class Gasteromycetes Fr. (literally "stomach fungi"), or the equally obsolete order Gasteromycetales Rea, because they produce spores inside their basidiocarps (fruit bodies) rather than on an outer surface. However, the class is polyphyletic, as such species—which include puffballs, earthstars, stinkhorns, and false truffles—are not closely related to each other. Because they are often studied as a group, it has been convenient to retain the informal (non- taxonomic) name of "gasteroid fungi". History Several gasteroid fungi—such as the stinkhorn, ''Phallus impudicus'' L.—were formally described by Carl Linnaeus in his original '' Species Plantarum'' of 1753, but the first critical treatment of the group was by Christiaan Hendrik Persoon in his ''Synopsis methodica fungorum'' of 1801. Until 1981, this book was the starting point for the naming ...
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Lamella (mycology)
In mycology, a lamella, or gill, is a papery hymenophore rib under the cap of some mushroom species, most often agarics. The gills are used by the mushrooms as a means of spore dispersal, and are important for species identification. The attachment of the gills to the stem is classified based on the shape of the gills when viewed from the side, while color, crowding and the shape of individual gills can also be important features. Additionally, gills can have distinctive microscopic or macroscopic features. For instance, ''Lactarius'' species typically seep latex from their gills. It was originally believed that all gilled fungi were Agaricales, but as fungi were studied in more detail, some gilled species were demonstrated not to be. It is now clear that this is a case of convergent evolution (i.e. gill-like structures evolved separately) rather than being an anatomic feature that evolved only once. The apparent reason that various basidiomycetes have evolved gills is that ...
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Basidiocarp
In fungi, a basidiocarp, basidiome, or basidioma () is the sporocarp of a basidiomycete, the multicellular structure on which the spore-producing hymenium is borne. Basidiocarps are characteristic of the hymenomycetes; rusts and smuts do not produce such structures. As with other sporocarps, epigeous (above-ground) basidiocarps that are visible to the naked eye (especially those with a more or less agaricoid morphology) are commonly referred to as mushrooms, while hypogeous (underground) basidiocarps are usually called false truffles. Structure All basidiocarps serve as the structure on which the hymenium is produced. Basidia are found on the surface of the hymenium, and the basidia ultimately produce spores. In its simplest form, a basidiocarp consists of an undifferentiated fruiting structure with a hymenium on the surface; such a structure is characteristic of many simple jelly and club fungi. In more complex basidiocarps, there is differentiation into a stipe, a ...
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Umbo (mycology)
'' Cantharellula umbonata'' has an umbo. The cap of '' Psilocybe makarorae'' is acutely papillate.">papillate.html" ;"title="Psilocybe makarorae'' is acutely papillate">Psilocybe makarorae'' is acutely papillate. An umbo is a raised area in the center of a mushroom cap. pileus (mycology), Caps that possess this feature are called ''umbonate''. Umbos that are sharply pointed are called ''acute'', while those that are more rounded are ''broadly umbonate''. If the umbo is elongated, it is '' cuspidate'', and if the umbo is sharply delineated but not elongated (somewhat resembling the shape of a human areola The human areola (''areola mammae'', or ) is the pigmented area on the breast around the nipple. Areola, more generally, is a small circular area on the body with a different histology from the surrounding tissue, or other small circular ...), it is called '' mammilate'' or ''papillate''. References {{reflist Fungal morphology and anatomy Mycology ...
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