turgor regulation
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Turgor pressure is the force within the cell that pushes the plasma membrane against the cell wall. It is also called ''hydrostatic pressure'', and is defined as the pressure in a fluid measured at a certain point within itself when at equilibrium. Generally, turgor pressure is caused by the
osmotic Osmosis (, ) is the spontaneous net movement or diffusion of solvent molecules through a selectively-permeable membrane from a region of high water potential (region of lower solute concentration) to a region of low water potential (region o ...
flow of water and occurs in
plant Plants are predominantly photosynthetic eukaryotes of the kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all current definitions of Plantae exclu ...
s,
fungi A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from ...
, and
bacteria Bacteria (; singular: bacterium) are ubiquitous, mostly free-living organisms often consisting of one Cell (biology), biological cell. They constitute a large domain (biology), domain of prokaryotic microorganisms. Typically a few micrometr ...
. The phenomenon is also observed in
protist A protist () is any eukaryotic organism (that is, an organism whose cells contain a cell nucleus) that is not an animal, plant, or fungus. While it is likely that protists share a common ancestor (the last eukaryotic common ancestor), the exc ...
s that have cell walls. This system is not seen in animal cells, as the absence of a cell wall would cause the cell to lyse when under too much pressure. The pressure exerted by the osmotic flow of water is called turgidity. It is caused by the osmotic flow of water through a selectively permeable membrane. Movement of water through a semipermeable membrane from a volume with a low solute concentration to one with a higher solute concentration is called osmotic flow. In plants, this entails the water moving from the low concentration solute outside the cell into the cell's
vacuole A vacuole () is a membrane-bound organelle which is present in plant and fungal cells and some protist, animal, and bacterial cells. Vacuoles are essentially enclosed compartments which are filled with water containing inorganic and organic m ...
.


Mechanism

Osmosis is the process in which water flows from a volume with a low
solute In chemistry, a solution is a special type of homogeneous mixture composed of two or more substances. In such a mixture, a solute is a substance dissolved in another substance, known as a solvent. If the attractive forces between the solvent ...
concentration (osmolarity), to an adjacent region with a higher solute concentration until equilibrium between the two areas is reached. It is usually accompanied by a favorable increase in the
entropy Entropy is a scientific concept, as well as a measurable physical property, that is most commonly associated with a state of disorder, randomness, or uncertainty. The term and the concept are used in diverse fields, from classical thermodynam ...
of the solvent. All cells are surrounded by a lipid bi-layer cell membrane which permits the flow of water into and out of the cell while limiting the flow of solutes. When the cell is in a
hypertonic In chemical biology, tonicity is a measure of the effective osmotic pressure gradient; the water potential of two solutions separated by a partially-permeable cell membrane. Tonicity depends on the relative concentration of selective membrane- ...
solution, water flows out of the cell, which decreases the cell's volume. When in a
hypotonic In chemical biology, tonicity is a measure of the effective osmotic pressure gradient; the water potential of two solutions separated by a partially-permeable cell membrane. Tonicity depends on the relative concentration of selective membrane-imp ...
solution, water flows into the membrane and increases the cell's volume, while in an isotonic solution, water flows in and out of the cell at an equal rate. Turgidity is the point at which the cell's membrane pushes against the cell wall, which is when turgor pressure is high. When the cell has low turgor pressure, it is flaccid. In plants, this is shown as wilted anatomical structures. This is more specifically known as plasmolysis.The volume and geometry of the cell affects the value of turgor pressure and how it can affect the cell wall's plasticity. Studies have shown that smaller cells experience a stronger elastic change when compared to larger cells. Turgor pressure also plays a key role in plant cell growth when the cell wall undergoes irreversible expansion due to the force of turgor pressure as well as structural changes in the cell wall that alter its extensibility.


Turgor pressure in plants

Turgor pressure within cells is regulated by osmosis and this also causes the cell wall to expand during growth. Along with size, rigidity of the cell is also caused by turgor pressure; a lower pressure results in a wilted cell or plant structure (i.e. leaf, stalk). One mechanism in plants that regulate turgor pressure is the cell's semipermeable membrane, which only allows some solutes to travel in and out of the cell, which can maintain a minimum amount of pressure. Other mechanisms include
transpiration Transpiration is the process of water movement through a plant and its evaporation from aerial parts, such as leaves, stems and flowers. Water is necessary for plants but only a small amount of water taken up by the roots is used for growth ...
, which results in water loss and decreases turgidity in cells. Turgor pressure is also a large factor for nutrient transport throughout the plant. Cells of the same organism can have differing turgor pressures throughout the organism's structure. In
higher plants Vascular plants (), also called tracheophytes () or collectively Tracheophyta (), form a large group of land plants ( accepted known species) that have lignified tissues (the xylem) for conducting water and minerals throughout the plant. They ...
, turgor pressure is responsible for apical growth of features such as root tips and pollen tubes.


Dispersal

Transport protein A transport protein (variously referred to as a transmembrane pump, transporter, escort protein, acid transport protein, cation transport protein, or anion transport protein) is a protein that serves the function of moving other materials within ...
s that pump solutes into the cell can be regulated by cell turgor pressure. Lower values allow for an increase in the pumping of solutes, which in turn increases osmotic pressure. This function is important as a plant response under drought conditions (seeing as turgor pressure is maintained), and for cells which need to accumulate solutes (i.e. developing
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particu ...
s).


Flowering and reproductive organs

It has been recorded that the petals of ''
Gentiana kochiana ''Gentiana acaulis'', the stemless gentian, or trumpet gentian, is a species of flowering plant in the family Gentianaceae, native to central and southern Europe, from Spain east to the Balkans, growing especially in mountainous regions, such as ...
'' and ''
Kalanchoe blossfeldiana ''Kalanchoe blossfeldiana'' is a herbaceous and commonly cultivated house plant of the genus ''Kalanchoe'' native to Madagascar. It is known by the English common names flaming Katy, Christmas kalanchoe, florist kalanchoe and Madagascar widow's ...
'' bloom via volatile turgor pressure of cells on the plant's
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