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''Uniola paniculata'', also known as sea oats, seaside oats, araña, and arroz de costa, is a tall
subtropical The subtropical zones or subtropics are geographical zone, geographical and Köppen climate classification, climate zones to the Northern Hemisphere, north and Southern Hemisphere, south of the tropics. Geographically part of the Geographical z ...
grass Poaceae () or Gramineae () is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos and the grasses of natural grassland and species cultivated in lawns and ...
that is an important component of coastal
sand dune A dune is a landform composed of wind- or water-driven sand. It typically takes the form of a mound, ridge, or hill. An area with dunes is called a dune system or a dune complex. A large dune complex is called a dune field, while broad, f ...
and
beach A beach is a landform alongside a body of water which consists of loose particles. The particles composing a beach are typically made from rock, such as sand, gravel, shingle, pebbles, etc., or biological sources, such as mollusc shells ...
plant communities in the
southeastern United States The Southeastern United States, also referred to as the American Southeast or simply the Southeast, is a geographical region of the United States. It is located broadly on the eastern portion of the southern United States and the southern ...
, eastern Mexico and some
Caribbean The Caribbean (, ) ( es, El Caribe; french: la Caraïbe; ht, Karayib; nl, De Caraïben) is a region of the Americas that consists of the Caribbean Sea, its islands (some surrounded by the Caribbean Sea and some bordering both the Caribbean S ...
islands. Its large seed heads that turn golden brown in late summer give the plant its common name. Its tall leaves trap wind-blown
sand Sand is a granular material composed of finely divided mineral particles. Sand has various compositions but is defined by its grain size. Sand grains are smaller than gravel and coarser than silt. Sand can also refer to a textural class of ...
and promote sand dune growth, while its deep
root In vascular plants, the roots are the organs of a plant that are modified to provide anchorage for the plant and take in water and nutrients into the plant body, which allows plants to grow taller and faster. They are most often below the sur ...
s and extensive
rhizome In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow h ...
s act to stabilize them, so the plant helps protect beaches and property from damage due to high winds,
storm surge A storm surge, storm flood, tidal surge, or storm tide is a coastal flood or tsunami-like phenomenon of rising water commonly associated with low-pressure weather systems, such as cyclones. It is measured as the rise in water level above the ...
s and
tide Tides are the rise and fall of sea levels caused by the combined effects of the gravitational forces exerted by the Moon (and to a much lesser extent, the Sun) and are also caused by the Earth and Moon orbiting one another. Tide tables can ...
s. It also provides food and habitat for birds, small animals and insects.


Description

''Uniola paniculata'' is a tall, erect
perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...
grass that can grow to in height. Its long, thin leaves reach lengths of and are about in width, tapering to a pointed apex. The plant produces
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed o ...
s of flat spikelets, each of which contains 10 to 12
wind-pollinated Anemophily or wind pollination is a form of pollination whereby pollen is distributed by wind. Almost all gymnosperms are anemophilous, as are many plants in the order Poales, including grasses, sedges, and rushes. Other common anemophilous plan ...
floret This glossary of botanical terms is a list of definitions of terms and concepts relevant to botany and plants in general. Terms of plant morphology are included here as well as at the more specific Glossary of plant morphology and Glossary ...
s. These ripen to golden brown
infructescence Infructescence (fruiting head) is defined as the ensemble of fruits derived from the ovaries of an inflorescence. It usually retains the size and structure of the inflorescence. In some cases, infructescences are similar in appearance to simple fru ...
s or seed heads in late summer. The
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiospe ...
s are dispersed by wind and can be carried long distances by storms and ocean currents, but reproduction commonly occurs
vegetatively Vegetative reproduction (also known as vegetative propagation, vegetative multiplication or cloning) is any form of asexual reproduction occurring in plants in which a new plant grows from a fragment or cutting of the parent plant or speci ...
by forming buds around stem bases. The plant forms dense surface roots and penetrating deep roots that are colonized by beneficial organisms such as
mycorrhizal fungi   A mycorrhiza (from Greek μύκης ', "fungus", and ῥίζα ', "root"; pl. mycorrhizae, mycorrhiza or mycorrhizas) is a symbiotic association between a fungus and a plant. The term mycorrhiza refers to the role of the fungus in the plant ...
. Rhizomes are elongate and produce extensive lateral growth. They root readily when buried in sand. ''Uniola paniculata'' uses a C4 pathway for carbon fixation.


Distribution and habitat

''Uniola paniculata'' is found on beach fronts and
barrier island Barrier islands are coastal landforms and a type of dune system that are exceptionally flat or lumpy areas of sand that form by wave and tidal action parallel to the mainland coast. They usually occur in chains, consisting of anything from a ...
s along the Atlantic Coast from
New Jersey New Jersey is a U.S. state, state in the Mid-Atlantic States, Mid-Atlantic and Northeastern United States, Northeastern regions of the United States. It is bordered on the north and east by the state of New York (state), New York; on the ea ...
to
Florida Florida is a U.S. state, state located in the Southeastern United States, Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia (U.S. state), Geo ...
, and the Gulf Coast from Florida to
Tabasco Tabasco (), officially the Free and Sovereign State of Tabasco ( es, Estado Libre y Soberano de Tabasco), is one of the 32 Federal Entities of Mexico. It is divided into 17 municipalities and its capital city is Villahermosa. It is located in ...
, Mexico. The range of ''Uniola paniculata'' along the US Atlantic coastline is expanding northward. It also occurs in
the Bahamas The Bahamas (), officially the Commonwealth of The Bahamas, is an island country within the Lucayan Archipelago of the West Indies in the North Atlantic. It takes up 97% of the Lucayan Archipelago's land area and is home to 88% of the archi ...
and northwestern
Cuba Cuba ( , ), officially the Republic of Cuba ( es, República de Cuba, links=no ), is an island country comprising the island of Cuba, as well as Isla de la Juventud and several minor archipelagos. Cuba is located where the northern Caribbe ...
. It grows primarily on
foredune A foredune is a dune ridge that runs parallel to the shore of an ocean, lake, bay, or estuary. Foredunes consist of sand deposited by wind on a vegetated part of the shore. Foredunes can be classified generally as ''incipient'' or ''established'' ...
s and dune crests. It is uncommon in swales between dunes where salt spray is limited, and it is rarely found inland.


Growing conditions

Due to the harsh conditions in which it grows, ''U. paniculata'' has little competition from other plants. It is heat tolerant and highly resistant to
drought A drought is defined as drier than normal conditions.Douville, H., K. Raghavan, J. Renwick, R.P. Allan, P.A. Arias, M. Barlow, R. Cerezo-Mota, A. Cherchi, T.Y. Gan, J. Gergis, D.  Jiang, A.  Khan, W.  Pokam Mba, D.  Rosenfeld, J. Tierney, an ...
,
salinity Salinity () is the saltiness or amount of salt dissolved in a body of water, called saline water (see also soil salinity). It is usually measured in g/L or g/kg (grams of salt per liter/kilogram of water; the latter is dimensionless and equal t ...
and brief inundation by
sea water Seawater, or salt water, is water from a sea or ocean. On average, seawater in the world's oceans has a salinity of about 3.5% (35 g/L, 35 ppt, 600 mM). This means that every kilogram (roughly one liter by volume) of seawater has ap ...
. It grows in loose sand rather than finer-grained
silt Silt is granular material of a size between sand and clay and composed mostly of broken grains of quartz. Silt may occur as a soil (often mixed with sand or clay) or as sediment mixed in suspension with water. Silt usually has a floury feel whe ...
y or
clay Clay is a type of fine-grained natural soil material containing clay minerals (hydrous aluminium phyllosilicates, e.g. kaolin, Al2 Si2 O5( OH)4). Clays develop plasticity when wet, due to a molecular film of water surrounding the clay parti ...
-rich
soil Soil, also commonly referred to as earth or dirt, is a mixture of organic matter, minerals, gases, liquids, and organisms that together support life. Some scientific definitions distinguish ''dirt'' from ''soil'' by restricting the former ...
s and does not tolerate water-logging. The plants tend to trap blowing sand, and burial of the plant base by sand stimulates growth and helps the plant spread by tacking down the rhizomes. ''Uniola paniculata'' is adversely affected by urban encroachment. Treated and untreated sewage,
urban runoff Urban runoff is surface runoff of rainwater, landscape irrigation, and car washing created by urbanization. Impervious surfaces (roads, parking lots and sidewalks) are constructed during land development. During rain , storms and other precipita ...
and pollution from
marina A marina (from Spanish , Portuguese and Italian : ''marina'', "coast" or "shore") is a dock or basin with moorings and supplies for yachts and small boats. A marina differs from a port in that a marina does not handle large passenger ships ...
s all impact the plant.
Off-road vehicles An off-road vehicle, sometimes referred to as an overland or adventure vehicle, is considered to be any type of vehicle which is capable of driving on and off paved or gravel surface. It is generally characterized by having large tires with d ...
and foot traffic damage the plants, disrupt their roots and compact the sand. Loss of the plants leads to erosion and loss of protective dunes.


Uses

Sea oats are well suited to saline environments, and as such, are important to barrier island
ecology Ecology () is the study of the relationships between living organisms, including humans, and their physical environment. Ecology considers organisms at the individual, population, community, ecosystem, and biosphere level. Ecology overlaps ...
and are often used in sand stabilization projects because their long root structure firmly holds loose sand. For example, in
Fort Lauderdale A fortification is a military construction or building designed for the defense of territories in warfare, and is also used to establish rule in a region during peacetime. The term is derived from Latin ''fortis'' ("strong") and ''facere ...
,
Florida Florida is a U.S. state, state located in the Southeastern United States, Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia (U.S. state), Geo ...
, colonies of sea oats have been planted at several beaches. The oats are a crucial component of the area's
hurricane A tropical cyclone is a rapidly rotating storm system characterized by a low-pressure center, a closed low-level atmospheric circulation, strong winds, and a spiral arrangement of thunderstorms that produce heavy rain and squalls. Depend ...
defense strategy and have helped to stave off damage from
tropical storm A tropical cyclone is a rapidly rotating storm system characterized by a low-pressure center, a closed low-level atmospheric circulation, strong winds, and a spiral arrangement of thunderstorms that produce heavy rain and squalls. Depen ...
s. The sea oat colonies and nascent dune structure they support are expected to flourish.


Legal protection

Sea oats are a protected grass in several states along the southeastern Atlantic coast. Picking or disturbing sea oats is punishable by fine in Georgia, South Carolina, and North Carolina. Florida by law also prohibits interfering with sea oats and protects nurserymen that grow sea oats.


Wildlife habitat

Seeds of ''U. paniculata'' provide food for
red-winged blackbird The red-winged blackbird (''Agelaius phoeniceus'') is a passerine bird of the family Icteridae found in most of North America and much of Central America. It breeds from Alaska and Newfoundland south to Florida, the Gulf of Mexico, Mexico, and ...
s, sparrows and other
songbird A songbird is a bird belonging to the suborder Passeri of the perching birds (Passeriformes). Another name that is sometimes seen as the scientific or vernacular name is Oscines, from Latin ''oscen'', "songbird". The Passeriformes contains 500 ...
s, as well as
marsh rabbit The marsh rabbit (''Sylvilagus palustris'') is a small cottontail rabbit found in marshes and swamps of coastal regions of the Eastern and Southern United States. It is a strong swimmer and found only near regions of water. It is similar in app ...
s and
mice A mouse ( : mice) is a small rodent. Characteristically, mice are known to have a pointed snout, small rounded ears, a body-length scaly tail, and a high breeding rate. The best known mouse species is the common house mouse (''Mus musculus' ...
. Florida
ornithologist Ornithology is a branch of zoology that concerns the "methodological study and consequent knowledge of birds with all that relates to them." Several aspects of ornithology differ from related disciplines, due partly to the high visibility and th ...
s have discovered that the pygmy
burrowing owl The burrowing owl (''Athene cunicularia''), also called the shoco, is a small, long-legged owl found throughout open landscapes of North and South America. Burrowing owls can be found in grasslands, rangelands, agricultural areas, deserts, or any ...
makes its nest within sea oat colonies to conceal its young from natural predators such as the
frigatebird Frigatebirds are a family of seabirds called Fregatidae which are found across all tropical and subtropical oceans. The five extant species are classified in a single genus, ''Fregata''. All have predominantly black plumage, long, deeply forked ...
s.


References

{{Taxonbar, from=Q3549978 paniculata Grasses of the United States Grasses of Mexico Flora of the Eastern United States Flora of the Southeastern United States Flora of Northeastern Mexico Flora of the Caribbean Grasses of Alabama Flora of Florida Halophytes Plants described in 1753 Taxa named by Carl Linnaeus Flora without expected TNC conservation status