A limb (from the

Old English Old English (, ), or Anglo-Saxon, is the earliest recorded form of the English language, spoken in England and southern and eastern Scotland in the early Middle Ages In the history of Europe, the Middle Ages or medieval period la ...

''lim'', meaning "body part") or leg is a jointed,

muscle Skeletal muscles (commonly referred to as muscles) are organs of the vertebrate muscular system and typically are attached by tendons to bones of a skeleton. The muscle cells of skeletal muscles are much longer than in the other types of muscl ...

d appendage that tetrapod

vertebrate Vertebrates () comprise all animal taxon, taxa within the subphylum Vertebrata () (chordates with vertebral column, backbones), including all mammals, birds, reptiles, amphibians, and fish. Vertebrates represent the overwhelming majority of the ...

s use for weight-bearing and terrestrial locomotion such as walking, running and jumping, for paddle-swimming, or for grasping and climbing. The distalmost portion of a limb is known as its extremity. The limbs' bony endoskeleton, known as the appendicular skeleton, is

homologous Homology may refer to: Sciences Biology *Homology (biology), any characteristic of biological organisms that is derived from a common ancestor *Sequence homology, biological homology between DNA, RNA, or protein sequences * Homologous chrom ...

among all tetrapods. All tetrapods have four limbs that are organized into two bilaterally symmetrical pairs, with one pair at each end of the

torso The torso or trunk is an anatomical term for the central part, or the core, of the body of many animals (including humans), from which the head, neck The neck is the part of the body on many vertebrates that connects the head with the tors ...

. The cranial pair are known as the forelimbs or ''front legs'', and the

caudal Caudal may refer to: Anatomy * Caudal (anatomical term) (from Latin ''cauda''; tail), used to describe how close something is to the trailing end of an organism * Caudal artery, the portion of the dorsal aorta of a vertebrate that passes into the ...

pair the hindlimbs or ''back legs''. In animals with more upright posture (mainly hominid primates, particularly

human Humans (''Homo sapiens'') are the most abundant and widespread species of primate, characterized by bipedalism and exceptional cognitive skills due to a large and complex brain. This has enabled the development of advanced tools, culture, ...

s), the forelimbs and hindlimbs are often called

upper Upper may refer to: * Shoe upper or ''vamp'', the part of a shoe on the top of the foot * Stimulant, drugs which induce temporary improvements in either mental or physical function or both * ''Upper'', the original film title for the 2013 found fo ...

and lower limbs, respectively. The fore-/upper limbs are connected to the thoracic cage via the shoulder girdles, and the hind-/lower limbs are connected to the

pelvis The pelvis (plural pelves or pelvises) is the lower part of the trunk, between the abdomen and the thighs (sometimes also called pelvic region), together with its embedded skeleton (sometimes also called bony pelvis, or pelvic skeleton). The ...

via the hip joints. Many animals, especially the arboreal species, have prehensile forelimbs adapted for grasping and climbing, while some (

bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...

s and bats) have expanded forelimbs (and sometimes hindlimbs as well) with specialized feathers or

membrane A membrane is a selective barrier; it allows some things to pass through but stops others. Such things may be molecules, ions, or other small particles. Membranes can be generally classified into synthetic membranes and biological membranes. ...

s to achieve lift and fly. Some animals (mostly primates) can also use hindlimbs for grasping. In the human body, the upper and lower limbs are commonly known as the arms and legs, respectively (although in academic definitions, these latter terms actually refer only to the upper arm and lower leg). The human arms have relatively great ranges of motion and are highly adapted for grasping and for carrying objects. The extremity of each arm, known as the hand, specializes in intrinsic and

digit Digit may refer to: Mathematics and science * Numerical digit, as used in mathematics or computer science ** Hindu-Arabic numerals, the most common modern representation of numerical digits * Digit (anatomy), the most distal part of a limb, such ...

al fine motor skills for precise manipulation of objects. The human legs and their extremities — the feet — are specialized for bipedal locomotion. Compared to most other mammals that walk and run on all four limbs, human limbs are proportionally weaker but very mobile and versatile, and the unique dexterity of the human upper extremities allows them to make sophisticated

tool A tool is an object that can extend an individual's ability to modify features of the surrounding environment or help them accomplish a particular task. Although many animals use simple tools, only human beings, whose use of stone tools dates ba ...

s and

machine A machine is a physical system using Power (physics), power to apply Force, forces and control Motion, movement to perform an action. The term is commonly applied to artificial devices, such as those employing engines or motors, but also to na ...

s that compensate for the lack of physical strength and endurance.

Terminology

The overall patterns of the forelimbs and hindlimbs are so similar ancestrally, and branch out in similar ways; that they are given shared names. Limbs are attached to the pectoral girdle or pelvic girdle. The one bony element of the upper limb is the ''stylopodium'', the two bones of the lower limb are the ''zeugopodium''. The distal portion of the limbs, that is, the hands or feet, are known as ''autopodia''. Hands are technically known as the ''manus'', and feet as the ''pes'', which are both composed of carpals and digits. As

metapodial Metapodials are long bone The long bones are those that are longer than they are wide. They are one of five types of bones: long, short, flat, irregular and sesamoid. Long bones, especially the femur and tibia, are subjected to most of the load d ...

s, the metacarpals and metatarsals are analogous to each other.

Development

Limb development is controlled by Hox genes. All jawed vertebrates surveyed so far organize their developing limb buds in a similar way. Growth occurs from proximal to distal part of the limb. On the distal end, the differentiation of skeletal elements occurs in an apical ectodermal ridge (AER) which expands in rays. A Zone of Polarizing Activity (ZPA) at the rear part of the AER coordinates the differentiation of digits.

References

{{Authority control

Terminology

Development

See also

References