Heliozoa, commonly known as sun-animalcules, are microbial

eukaryotes Eukaryotes () are organisms whose cells have a nucleus. All animals, plants, fungi, and many unicellular organisms, are Eukaryotes. They belong to the group of organisms Eukaryota or Eukarya, which is one of the three domains of life. Bacte ...

(

protists A protist () is any eukaryotic organism (that is, an organism whose cells contain a cell nucleus) that is not an animal, plant, or fungus. While it is likely that protists share a common ancestor (the last eukaryotic common ancestor), the excl ...

) with stiff arms ( axopodia) radiating from their spherical bodies, which are responsible for their common name. The axopodia are microtubule-supported projections from the

amoeboid An amoeba (; less commonly spelled ameba or amœba; plural ''am(o)ebas'' or ''am(o)ebae'' ), often called an amoeboid, is a type of cell or unicellular organism with the ability to alter its shape, primarily by extending and retracting pseudopo ...

cell body, and are variously used for capturing food, sensation, movement, and attachment. They are similar to

Radiolaria The Radiolaria, also called Radiozoa, are protozoa of diameter 0.1–0.2 mm that produce intricate mineral skeletons, typically with a central capsule dividing the cell (biology), cell into the inner and outer portions of endoplasm and Ecto ...

, but they are distinguished from them by lacking central capsules and other complex skeletal elements, although some produce simple scales and spines. They may be found in both

freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does include ...

and marine environments.

Classification

Originally the heliozoa were treated together as a formal taxon Heliozoa or Heliozoea, with the rank of class or phylum, but it has been realised that they are

polyphyletic A polyphyletic group is an assemblage of organisms or other evolving elements that is of mixed evolutionary origin. The term is often applied to groups that share similar features known as homoplasies, which are explained as a result of converg ...

, as the various orders show notable differences and are no longer believed to be descended from a single common ancestor. Instead, "heliozoa" is regarded as a descriptive term applying to various lines of protists. The primary groups include: *

Actinophryida The actinophryids are an order of heliozoa, a polyphyletic array of stramenopiles, having a close relationship with pedinellids and ''Ciliophrys''. They are common in fresh water and occasionally found in marine and soil habitats. Actinophryids a ...

Pedinellid Pedinellales is a group of single-celled algae found in both marine environments and freshwater. These are found in both freshwater and marine environments, and most genera are sessile, attached by posterior stalks. The flagellum is at the ant ...

a and Ciliophryida (currently in

Stramenopiles Stramenopile is a clade of organisms distinguished by the presence of stiff tripartite external hairs. In most species, the hairs are attached to flagella, in some they are attached to other areas of the cellular surface, and in some they have be ...

) *

Centrohelida The centrohelids or centroheliozoa are a large group of heliozoan protists. They include both mobile and sessile forms, found in freshwater and marine environments, especially at some depth. Characteristics Individuals are unicellular and spher ...

(some support for

Hacrobia The cryptomonads-haptophytes assemblage is a proposed monophyletic grouping of unicellular eukaryotes that are not included in the SAR supergroup. Several alternative names have been used for the group, including Hacrobia (derived from "ha-" refe ...

) * Desmothoracida, Heliomonadida/Dimorphida and Gymnosphaerida (currently in

Rhizaria The Rhizaria are an ill-defined but species-rich supergroup of mostly unicellular eukaryotes. Except for the Chlorarachniophytes and three species in the genus Paulinella in the phylum Cercozoa, they are all non-photosynthethic, but many foramini ...

Cercozoa Cercozoa is a phylum of diverse single-celled eukaryotes. They lack shared morphological characteristics at the microscopic level, and are instead defined by molecular phylogenies of rRNA and actin or polyubiquitin. They were the first major eu ...

) *

Taxopodida ''Sticholonche'' is a genus of radiolarians with a single species, ''Sticholonche zanclea'', found in open oceans at depths of 99–510 metres. It is generally considered a heliozoan, placed in its own order, called the Taxopodida. However it ...

/''

Sticholonche ''Sticholonche'' is a genus of radiolarians with a single species, ''Sticholonche zanclea'', found in open oceans at depths of 99–510 metres. It is generally considered a heliozoan, placed in its own order, called the Taxopodida. However it ...

'' (currently in

) * Rotosphaerida (currently in Opisthokonta >

Nucleariida Nucleariida is a group of amoebae with filose pseudopods, known mostly from soils and freshwater. They are distinguished from the superficially similar vampyrellids mainly by having mitochondria with discoid cristae, in the absence of superf ...

and in

) Several

nucleariid Nucleariida is a group of amoebae with filose pseudopods, known mostly from soils and freshwater. They are distinguished from the superficially similar vampyrellids mainly by having mitochondria with discoid cristae, in the absence of superfi ...

s were once considered heliozoa, but they do not have microtubule-supported axopods and so are now considered filose amoeboids instead.

References

External links

* ''Actinosphaerium'' (single-celled) is an example of a heliozoa. For an image see http://www.microscopy-uk.org.uk/index.html?http://www.microscopy-uk.org.uk/ponddip/ * Obsolete eukaryote taxa Eukaryote orders {{Eukaryote-stub