Zanthoxylum Scandens
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''Zanthoxylum scandens'' () is a woody plant from the family
Rutaceae The Rutaceae is a family, commonly known as the rueRUTACEAE
in BoDD – Botanical Derm ...
.


Description

''Zanthoxylum scandens'' is a woody plant, observed as a shrub or woody climber. It has been found in lowland forests, open forests, and thickets from an elevation at near sea level to . Geographically, the species has been spotted in areas within South
Anhui Anhui , (; formerly romanized as Anhwei) is a landlocked province of the People's Republic of China, part of the East China region. Its provincial capital and largest city is Hefei. The province is located across the basins of the Yangtze River ...
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Chongqing Chongqing ( or ; ; Sichuanese dialects, Sichuanese pronunciation: , Standard Mandarin pronunciation: ), Postal Romanization, alternately romanized as Chungking (), is a Direct-administered municipalities of China, municipality in Southwes ...
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Fujian Fujian (; alternately romanized as Fukien or Hokkien) is a province on the southeastern coast of China. Fujian is bordered by Zhejiang to the north, Jiangxi to the west, Guangdong to the south, and the Taiwan Strait to the east. Its capi ...
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Guangdong Guangdong (, ), alternatively romanized as Canton or Kwangtung, is a coastal province in South China on the north shore of the South China Sea. The capital of the province is Guangzhou. With a population of 126.01 million (as of 2020) ...
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Guangxi Guangxi (; ; Chinese postal romanization, alternately romanized as Kwanghsi; ; za, Gvangjsih, italics=yes), officially the Guangxi Zhuang Autonomous Region (GZAR), is an Autonomous regions of China, autonomous region of the People's Republic ...
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Guizhou Guizhou (; formerly Kweichow) is a landlocked province in the southwest region of the People's Republic of China. Its capital and largest city is Guiyang, in the center of the province. Guizhou borders the autonomous region of Guangxi to t ...
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Hainan Hainan (, ; ) is the smallest and southernmost province of the People's Republic of China (PRC), consisting of various islands in the South China Sea. , the largest and most populous island in China,The island of Taiwan, which is slightly l ...
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Hubei Hubei (; ; alternately Hupeh) is a landlocked province of the People's Republic of China, and is part of the Central China region. The name of the province means "north of the lake", referring to its position north of Dongting Lake. The prov ...
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Hunan Hunan (, ; ) is a landlocked province of the People's Republic of China, part of the South Central China region. Located in the middle reaches of the Yangtze watershed, it borders the province-level divisions of Hubei to the north, Jiangxi to ...
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Jiangxi Jiangxi (; ; formerly romanized as Kiangsi or Chianghsi) is a landlocked province in the east of the People's Republic of China. Its major cities include Nanchang and Jiujiang. Spanning from the banks of the Yangtze river in the north int ...
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Sichuan Sichuan (; zh, c=, labels=no, ; zh, p=Sìchuān; alternatively romanized as Szechuan or Szechwan; formerly also referred to as "West China" or "Western China" by Protestant missions) is a province in Southwest China occupying most of the ...
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Taiwan Taiwan, officially the Republic of China (ROC), is a country in East Asia, at the junction of the East and South China Seas in the northwestern Pacific Ocean, with the People's Republic of China (PRC) to the northwest, Japan to the nort ...
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Yunnan Yunnan , () is a landlocked Provinces of China, province in Southwest China, the southwest of the People's Republic of China. The province spans approximately and has a population of 48.3 million (as of 2018). The capital of the province is ...
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Zhejiang Zhejiang ( or , ; , also romanized as Chekiang) is an eastern, coastal province of the People's Republic of China. Its capital and largest city is Hangzhou, and other notable cities include Ningbo and Wenzhou. Zhejiang is bordered by Jiang ...
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India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
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Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guine ...
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Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...
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Malaysia Malaysia ( ; ) is a country in Southeast Asia. The federation, federal constitutional monarchy consists of States and federal territories of Malaysia, thirteen states and three federal territories, separated by the South China Sea into two r ...
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Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John C. Wells, Joh ...
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Laos Laos (, ''Lāo'' )), officially the Lao People's Democratic Republic ( Lao: ສາທາລະນະລັດ ປະຊາທິປະໄຕ ປະຊາຊົນລາວ, French: République démocratique populaire lao), is a socialist ...
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Vietnam Vietnam or Viet Nam ( vi, Việt Nam, ), officially the Socialist Republic of Vietnam,., group="n" is a country in Southeast Asia, at the eastern edge of mainland Southeast Asia, with an area of and population of 96 million, making i ...
. The species has a self-supporting growth form, with individual plants growing up to . Prickles can be found on its trunks, branches, branchlets, and leaf rachises. Flowers are tetramerous, with a perianth in two series. It sepals have been found as pale purplish green in color, ovate in shape, and about in size. Male flowers have four stamens, are about three to four millimeters in size, and have a spot at their apex. Female flowers have three carpels with ligulate staminodes. Fruit follicles have been observed as purplish red in color, but grayish brown to black when dry. Seeds are about four to five millimeters in diameter. The species typically flowers from March to May, and fruits from to July to August.


Classification

The species was first published in Carl Ludwig von Blume's ''Bijdragen tot de Flora van Nederlandsch Indie'' in 1825. It has widely been accepted as a species, including in ''Flora of Japan II'' (Iwatsuki, K., Boufford, D.E. & Ohba, H., 1999), ''A Checklist of the Trees, Shrubs, Herbs and Climbers of Myanmar'' (Kress, W.J., DeFilipps, R.A., Farr, E. & Kyi, D.Y.Y., 2003), ''Danh lục các loài thực vật Việt Nam'' (Lê, T.C., 2003), ''Check-List of Flora of Meghalaya'' (Mao, A.A., Sinha, B.K., Verma, D. & Sarma, N., 2016), '' Bokor National Park A picture guide of forest trees in Cambodia'' (Tagane, S. & al., 2017), and ''Flora of China''. (Wu, Z. & Raven, P.H., 2008). 529 occurrences have been officially recorded. Three specimens are kept at the ''
Royal Botanic Gardens, Kew Royal Botanic Gardens, Kew is a non-departmental public body in the United Kingdom sponsored by the Department for Environment, Food and Rural Affairs. An internationally important botanical research and education institution, it employs 1,100 ...
''. Twelve potential synonyms have been identified, including ''Fagara chinensis'', ''Fagara cuspidata'', ''Fagara cyrtorhachia'', ''Fagara laxifoliolata'', ''Fagara leiorhachia'', ''Fagara scandens'', ''Zanthoxylum chinense'', ''Zanthoxylum cuspidatum'', ''Zanthoxylum cyrtorbachium'', ''Zanthoxylum laxifoliolatum'', ''Zanthoxylum leiorhachium'' and ''Zanthoxylum liukiuense''.


References

scandens Flora of China {{Rutaceae-stub