''Yurgovuchia'' (meaning "coyote") is a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

dromaeosaurid Dromaeosauridae () is a family of feathered theropod dinosaurs. They were generally small to medium-sized feathered carnivores that flourished in the Cretaceous Period. The name Dromaeosauridae means 'running lizards', from Greek ('), meaning ...

theropod Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally c ...

dinosaurs Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...

that lived in

North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...

during the

Early Cretaceous The Early Cretaceous ( geochronological name) or the Lower Cretaceous (chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145 Ma to 100.5 Ma. Geology Pro ...

period in what is now the

Cedar Mountain Formation The Cedar Mountain Formation is the name given to a distinctive sedimentary geologic formation in eastern Utah, spanning most of the early and mid-Cretaceous. The formation was named for Cedar Mountain (Utah), Cedar Mountain in northern Emery Coun ...

. It contains a single species, ''Yurgovuchia doellingi''. The remains were discovered in

Utah Utah ( , ) is a state in the Mountain West subregion of the Western United States. Utah is a landlocked U.S. state bordered to its east by Colorado, to its northeast by Wyoming, to its north by Idaho, to its south by Arizona, and to it ...

United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...

Discovery and naming

The holotype was collected by Donald D. DeBlieux in 2005, from Don's Place, part of the Doelling's Bowl bone bed in Grand County, Utah. This bone bed is in the lower

Yellow Cat Member The Cedar Mountain Formation is the name given to a distinctive sedimentary geologic formation in eastern Utah, spanning most of the early and mid-Cretaceous. The formation was named for Cedar Mountain in northern Emery County, Utah, where Willia ...

of the

, dating probably to the

Valanginian In the geologic timescale, the Valanginian is an age or stage of the Early or Lower Cretaceous. It spans between 139.8 ± 3.0 Ma and 132.9 ± 2.0 Ma (million years ago). The Valanginian Stage succeeds the Berriasian Stage of the Lower Cretaceou ...

stage of the

period Period may refer to: Common uses * Era, a length or span of time * Full stop (or period), a punctuation mark Arts, entertainment, and media * Period (music), a concept in musical composition * Periodic sentence (or rhetorical period), a concept ...

, about 139–134.6

million years ago The abbreviation Myr, "million years", is a unit of a quantity of (i.e. ) years, or 31.556926 teraseconds. Usage Myr (million years) is in common use in fields such as Earth science and cosmology. Myr is also used with Mya (million years ago). ...

. ''Yurgovuchia'' was first described and named by Phil Senter,

James I. Kirkland James Ian Kirkland (born August 24, 1954) is an American Palaeontologist, paleontologist and geologist. He has worked with dinosaur remains from the south west United States of America and Mexico and has been responsible for discovering new and i ...

, Donald D. DeBlieux, Scott Madsen and Natalie Toth in

2012 File:2012 Events Collage V3.png, From left, clockwise: The passenger cruise ship Costa Concordia lies capsized after the Costa Concordia disaster; Damage to Casino Pier in Seaside Heights, New Jersey as a result of Hurricane Sandy; People gather ...

and the

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...

is ''Yurgovuchia doellingi''. The generic name is derived from the

Ute Ute or UTE may refer to: * Ute (band), an Australian jazz group * Ute (given name) * ''Ute'' (sponge), a sponge genus * Ute (vehicle), an Australian and New Zealand term for certain utility vehicles * Ute, Iowa, a city in Monona County along ...

word ''yurgovuch'', meaning

coyote The coyote (''Canis latrans'') is a species of canis, canine native to North America. It is smaller than its close relative, the wolf, and slightly smaller than the closely related eastern wolf and red wolf. It fills much of the same ecologica ...

, a predator of similar size to ''Y. doellingi'' which currently inhabits the same region. The

specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...

, ''doellingi'', honors the geologist Helmut Doelling, for his 50-plus years of geological research and mapping of Utah for the

Utah Geological Survey The Utah Geological Survey is based in Salt Lake City, Utah, United States. It also has an office in Cedar City, Utah. It is a division of the

and for causing the discovery of the Doelling's Bowl dinosaur sites, in which ''Y. doellingi'' was collected.

Description

''Yurgovuchia'' is known only from a single individual represented by an associated partial

postcranial Postcrania (postcranium, adjective: postcranial) in zoology and vertebrate paleontology is all or part of the skeleton apart from the skull. Frequently, fossil remains, e.g. of dinosaurs or other extinct tetrapods, consist of partial or isolated sk ...

skeleton A skeleton is the structural frame that supports the body of an animal. There are several types of skeletons, including the exoskeleton, which is the stable outer shell of an organism, the endoskeleton, which forms the support structure inside ...

. The

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...

and the only known specimen, UMNH VP 20211, includes

cervical In anatomy, cervical is an adjective that has two meanings: # of or pertaining to any neck. # of or pertaining to the female cervix: i.e., the ''neck'' of the uterus. *Commonly used medical phrases involving the neck are **cervical collar **cervic ...

dorsal Dorsal (from Latin ''dorsum'' ‘back’) may refer to: * Dorsal (anatomy), an anatomical term of location referring to the back or upper side of an organism or parts of an organism * Dorsal, positioned on top of an aircraft's fuselage * Dorsal co ...

, and

caudal vertebrae The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic ...

as well as the proximal end of a left pubis. The preserved vertebrae are fused, indicating that the specimen was an adult at the time of death and not a juvenile specimen of ''

Utahraptor ''Utahraptor'' (meaning "Utah's thief") is a genus of large dromaeosaurid dinosaur that lived in North America during the Early Cretaceous period. It was a heavy-built, ground-dwelling, bipedal carnivore. It contains a single species, ''Utahrap ...

''. Instead of this, it represents a smaller and different species of

, with an estimed size of .

According to its describers in 2012, ''Yurgovuchia'' can be recognized from other dromaeosaurid taxa by the following characteristics: each side of the axial centrum have a single pneumatopore, third cervical vertebra with cranial end of the centrum not beveled, cervical

prezygapophyses The articular processes or zygapophyses (Greek language, Greek ζυγον = "yoke" (because it links two vertebrae) + απο = "away" + φυσις = "-physis, process") of a vertebra are projections of the vertebra that serve the purpose of fitting ...

is flexed, cervical vertebrae epipophyses is above the postzygapophyseal facets, the pubis lacks pubic

tubercle In anatomy, a tubercle (literally 'small tuber', Latin for 'lump') is any round nodule, small eminence, or warty outgrowth found on external or internal organs of a plant or an animal. In plants A tubercle is generally a wart-like projection ...

, the cranial faces of the centrum of caudal vertebrae are round, the cervical and dorsal vertebrae preserve hypapophyses without pneumatopores, the caudal prezygapophyses is distally elongated to the transition point, but not surpassing the length of a

centrum (Latin for ''center'') may refer to: Places In Greenland * Nuuk Centrum, a district of Nuuk, Greenland * Centrum Lake, Greenland In the Netherlands * Amsterdam-Centrum, the inner-most borough of Amsterdam, Netherlands * Rotterdam Centrum, a borou ...

Classification

According to the

phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...

analysis performed by its describers in 2012, ''Yurgovuchia'' represents advanced

dromaeosaurine Dromaeosaurinae is a subfamily of the theropod group Dromaeosauridae. The earliest dromaeosaurine is ''Utahraptor'', dating back to the Early Cretaceous period in North America, however, some isolated teeth seems to represent an indeterminate spe ...

s, closely related to ''

Achillobator ''Achillobator'' ( ; meaning "Achilles hero") is a genus of large dromaeosaurid dinosaurs that lived in Asia during the Late Cretaceous period about 96 million to 89 million years ago in what is now the Bayan Shireh Formation. The genus is curren ...

'', ''

Dromaeosaurus ''Dromaeosaurus'' (, "running lizard") is a genus of dromaeosaurid theropod dinosaur which lived during the Late Cretaceous period (middle late Campanian and Maastrichtian), sometime between 80 and 69 million years ago, in Alberta, Canada and the ...

'' and ''

''. Below are the results obtained:

Paleoecology

Cedar Mountain Formation Yellow Cat Fauna

Other dinosaurs are also known from Don's Place including the iguanodontians ''

Iguanacolossus ''Iguanacolossus'' (meaning "Iguana Colossus" or "Colossal Iguana") is a genus of iguanodontian ornithopod dinosaur that lived in North America during the Early Cretaceous period. It is known from UMNH VP 20205, the associated holotype with a lar ...

'', polacanthines, and

velociraptorine Velociraptorinae is a subfamily of the theropod group Dromaeosauridae. The earliest velociraptorines are probably ''Nuthetes'' from the United Kingdom, and possibly ''Deinonychus'' from North America. However, several indeterminate velociraptorin ...

dromaeosaurids represented by a pubis (UMNH VP 21752) and a tentative radius (UMNH VP 21751). Many additional theropods have previously been described from the Yellow Cat Member, including the therizinosaur ''

Falcarius ''Falcarius'' (meaning "sickle cutter") is a genus of primitive therizinosaur dinosaur that lived during the Early Cretaceous period in what is now North America. Its remains were first collected in the Cedar Mountain Formation in 1999, with subs ...

'' and the small, predatory troodontids ''

Geminiraptor ''Geminiraptor'' (meaning "twin seizer") is a genus of troodontid theropod dinosaur that lived in North America during the Early Cretaceous period. ''Geminiraptor'' was a small, ground-dwelling bipedal carnivorous paravian. The type species of '' ...

'' from the lower part of the member, in addition, the large dromaeosaurines ''

'', small

ornithomimosaurs Ornithomimosauria ("bird-mimic lizards") are theropod dinosaurs which bore a superficial resemblance to the modern-day ostrich. They were fast, omnivorous or herbivorous dinosaurs from the Cretaceous Period of Laurasia (now Asia, Europe and North ...

Nedcolbertia ''Nedcolbertia'' is a genus of theropod dinosaur from the Early Cretaceous Period of North America. Discovery and naming Three skeletons of a theropod were discovered in 1993 by Christopher Whittle near Cisco in the basal Yellow Cat Member of ...

'', and unnamed

eudromaeosaurs Eudromaeosauria ("true dromaeosaurs") is a subgroup of terrestrial Dromaeosauridae, dromaeosaurid theropod dinosaurs. They were relatively small to medium-sized, feathered hypercarnivores (with diets consisting almost entirely of other terrestr ...

represented by a tail skeleton (UMNH VP 20209) from the upper part of the member. ''Yurgovuchia'' shared its environment and lived alongside other dinosaurs in the Lower Yellow Cat, such as the

theropods Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally c ...

''Falcarius'' and ''Geminiraptor'', the

sauropods Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their bo ...

Mierasaurus ''Mierasaurus'' is an extinct genus of sauropod dinosaur from the Early Cretaceous of Utah, United States. The taxon was first described and named in 2017 by Rafael Royo-Torres and colleagues, from a mostly complete skeleton including a disarti ...

'' and the large

iguanodontian Iguanodontia (the iguanodonts) is a clade of herbivorous dinosaurs that lived from the Middle Jurassic to Late Cretaceous. Some members include ''Camptosaurus'', ''Dryosaurus'', ''Iguanodon'', ''Tenontosaurus'', and the hadrosaurids or "duck-bill ...

s ''Iguanacolossus''. There are also indeterminate

goniopholidid Goniopholididae is an extinct family of moderate-sized semi-aquatic neosuchian crocodyliformes. Their bodyplan and morphology are convergent on living crocodilians. They lived across Laurasia (Asia, Europe and North America) between the Middle Ju ...

crocodiles and the unnamed

velociraptorines Velociraptorinae is a subfamily of the theropod group Dromaeosauridae. The earliest velociraptorines are probably ''Nuthetes'' from the United Kingdom, and possibly ''Deinonychus'' from North America. However, several indeterminate velociraptorin ...

known from the Lower Yellow Cat.

References

{{Taxonbar, from=Q3236057 Eudromaeosaurs Early Cretaceous dinosaurs of North America Fossil taxa described in 2012 Taxa named by James I. Kirkland Paleontology in Utah

Discovery and naming

Description

Classification

Paleoecology

See also

References