White Willow
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''Salix alba'', the white willow, is a species of
willow Willows, also called sallows and osiers, from the genus ''Salix'', comprise around 400 speciesMabberley, D.J. 1997. The Plant Book, Cambridge University Press #2: Cambridge. of typically deciduous trees and shrubs, found primarily on moist so ...
native to
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and western and central
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.Meikle, R. D. (1984). ''Willows and Poplars of Great Britain and Ireland''. BSBI Handbook No. 4. .Rushforth, K. (1999). ''Trees of Britain and Europe''. Collins . The name derives from the white tone to the undersides of the leaves. It is a medium-sized to large
deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, ...
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
growing up to 10–30 m tall, with a trunk up to 1 m diameter and an irregular, often-leaning crown. The bark is grey-brown, and deeply fissured in older trees. The shoots in the typical species are grey-brown to green-brown. The leaves are paler than most other willows, due to a covering of very fine, silky white hairs, in particular on the underside; they are 5–10 cm long and 0.5–1.5 cm wide. The
flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechani ...
s are produced in catkins in early spring, and pollinated by
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three ...
s. It is dioecious, with male and female catkins on separate trees; the male catkins are 4–5 cm long, the female catkins 3–4 cm long at pollination, lengthening as the fruit matures. When mature in midsummer, the female catkins comprise numerous small (4 mm) capsules, each containing numerous minute
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiospe ...
s embedded in white down, which aids wind dispersal.Mitchell, A. F. (1974). ''A Field Guide to the Trees of Britain and Northern Europe''. Collins


Ecology

Like all willows, ''Salix alba'' is usually to be found in wet or poorly-drained soil at the edge of pools, lakes or rivers. Its wide-spreading roots take up moisture from a large surrounding area. White willows are fast-growing, but relatively short-lived, being susceptible to several
disease A disease is a particular abnormal condition that negatively affects the structure or function of all or part of an organism, and that is not immediately due to any external injury. Diseases are often known to be medical conditions that a ...
s, including watermark disease caused by the
bacterium Bacteria (; singular: bacterium) are ubiquitous, mostly free-living organisms often consisting of one biological cell. They constitute a large domain of prokaryotic microorganisms. Typically a few micrometres in length, bacteria were amon ...
'' Brenneria salicis'' (named because of the characteristic 'watermark' staining in the
wood Wood is a porous and fibrous structural tissue found in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulose fibers that are strong in tension and embedded in a matrix of lignin ...
; syn. ''Erwinia salicis'') and willow
anthracnose A plant canker is a small area of dead tissue, which grows slowly, often over years. Some cankers are of only minor consequence, but others are ultimately lethal and therefore can have major economic implications for agriculture and horticultur ...
, caused by the
fungus A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from t ...
''Marssonina salicicola''. These diseases can be a serious problem on trees grown for timber or ornament. It readily forms natural hybrids with crack willow '' Salix fragilis'', the hybrid being named ''Salix ''×'' rubens'' Schrank.


Varieties, cultivars and hybrids

A number of
cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture ...
s and
hybrid Hybrid may refer to: Science * Hybrid (biology), an offspring resulting from cross-breeding ** Hybrid grape, grape varieties produced by cross-breeding two ''Vitis'' species ** Hybridity, the property of a hybrid plant which is a union of two dif ...
s have been selected for
forestry Forestry is the science and craft of creating, managing, planting, using, conserving and repairing forests, woodlands, and associated resources for human and environmental benefits. Forestry is practiced in plantations and natural stands. ...
and
horticultural Horticulture is the branch of agriculture that deals with the art, science, technology, and business of plant cultivation. It includes the cultivation of fruits, vegetables, nuts, seeds, herbs, sprouts, mushrooms, algae, flowers, seaweeds and no ...
use: *''Salix alba'' 'Caerulea' (cricket-bat willow; syn. ''Salix alba'' var. ''caerulea'' (Sm.) Sm.; ''Salix caerulea'' Sm.) is grown as a specialist
timber Lumber is wood that has been processed into dimensional lumber, including beams and planks or boards, a stage in the process of wood production. Lumber is mainly used for construction framing, as well as finishing (floors, wall panels, w ...
crop in
Britain Britain most often refers to: * The United Kingdom, a sovereign state in Europe comprising the island of Great Britain, the north-eastern part of the island of Ireland and many smaller islands * Great Britain, the largest island in the United King ...
, mainly for the production of cricket bats, and for other uses where a tough, lightweight
wood Wood is a porous and fibrous structural tissue found in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulose fibers that are strong in tension and embedded in a matrix of lignin ...
that does not splinter easily is required. It is distinguished mainly by its growth form, very fast-growing with a single straight stem, and also by its slightly larger leaves (10–11 cm long, 1.5–2 cm wide) with a more blue-green colour. Its origin is unknown; it may be a hybrid between white willow and crack willow, but this is not confirmed. *''Salix alba'' 'Vitellina' (golden willow; syn. ''Salix alba'' var. ''vitellina'' (L.) Stokes) is a cultivar grown in gardens for its shoots, which are golden-yellow for one to two years before turning brown. It is particularly decorative in winter; the best effect is achieved by
coppicing Coppicing is a traditional method of woodland management which exploits the capacity of many species of trees to put out new shoots from their stump or roots if cut down. In a coppiced wood, which is called a copse, young tree stems are repeate ...
it every two to three years to stimulate the production of longer young shoots with better colour. Other similar cultivars include 'Britzensis', 'Cardinal', and 'Chermesina', selected for even brighter orange-red shoots. *''Salix alba'' 'Vitellina-Tristis' (golden weeping willow, synonym 'Tristis') is a weeping cultivar with yellow branches that become reddish-orange in winter. It is now rare in cultivation and has been largely replaced by ''Salix x sepulcralis'' 'Chrysocoma'. It is, however, still the best choice in very cold parts of the world, such as Canada, the northern US, and Russia. *The golden hybrid weeping willow (''Salix ''×'' sepulcralis'' 'Chrysocoma') is a hybrid between white willow and Peking willow ''
Salix babylonica ''Salix babylonica'' (Babylon willow or weeping willow; ) is a species of willow native to dry areas of northern China, but cultivated for millennia elsewhere in Asia, being traded along the Silk Road to southwest Asia and Europe.Flora of China'' ...
''.


Award of Garden Merit

The following have received the
Royal Horticultural Society The Royal Horticultural Society (RHS), founded in 1804 as the Horticultural Society of London, is the UK's leading gardening charity. The RHS promotes horticulture through its five gardens at Wisley (Surrey), Hyde Hall (Essex), Harlow Carr (Nor ...
's Award of Garden Merit *''Salix alba'' 'Golden Ness' *''Salix alba'' var. ''serica'' (silver willow) *''Salix alba'' var. ''vitellina'' 'Yelverton' *''Salix'' × ''sepulcralis'' 'Erythroflexuosa' *''Salix'' × ''sepulcralis'' var. ''chrysocoma''


Uses

The
wood Wood is a porous and fibrous structural tissue found in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulose fibers that are strong in tension and embedded in a matrix of lignin ...
is tough, strong, and light in weight, but has minimal resistance to decay. The stems (
withies A withy or withe (also willow and osier) is a strong flexible willow stem, typically used in thatching, basketmaking, gardening and for constructing woven wattle hurdles.
) from
coppiced Coppicing is a traditional method of woodland management which exploits the capacity of many species of trees to put out new shoots from their stump or roots if cut down. In a coppiced wood, which is called a copse, young tree stems are repeate ...
and
pollarded Pollarding is a pruning system involving the removal of the upper branches of a tree, which promotes the growth of a dense head of foliage and branches. In ancient Rome, Propertius mentioned pollarding during the 1st century BCE. The practice oc ...
plants are used for basket-making. Charcoal made from the wood was important for
gunpowder Gunpowder, also commonly known as black powder to distinguish it from modern smokeless powder, is the earliest known chemical explosive. It consists of a mixture of sulfur, carbon (in the form of charcoal) and potassium nitrate (saltpeter). Th ...
manufacture. The bark tannin was used in the past for
tanning Tanning may refer to: *Tanning (leather), treating animal skins to produce leather *Sun tanning, using the sun to darken pale skin **Indoor tanning, the use of artificial light in place of the sun **Sunless tanning, application of a stain or dye t ...
leather. The wood is used to make cricket bats. ''S. alba'' wood has a low density and a lower transverse compressive strength. This allows the wood to bend, which is why it can be used to make baskets. Willow bark contains
indole-3-butyric acid Indole-3-butyric acid (1''H''-indole-3-butanoic acid, IBA) is a white to light-yellow crystalline solid, with the molecular formula C12H13NO2. It melts at 125 °C in atmospheric pressure and decomposes before boiling. IBA is a plant hormone ...
, which is a plant hormone stimulating root growth; willow trimmings are sometimes used to clone
rootstock A rootstock is part of a plant, often an underground part, from which new above-ground growth can be produced. It could also be described as a stem with a well developed root system, to which a bud from another plant is grafted. It can refer to a ...
in place of commercially synthesized root stimulator. It is also used for ritual purposes by Jews on the holiday of Sukkot.


Medicinal uses

Willow Willows, also called sallows and osiers, from the genus ''Salix'', comprise around 400 speciesMabberley, D.J. 1997. The Plant Book, Cambridge University Press #2: Cambridge. of typically deciduous trees and shrubs, found primarily on moist so ...
(of unspecified species) has long been used by herbalists for various ailments, although it is a myth that they attribute to it any analgesic effect. One of the first references to White Willow specifically was by Edward Stone, of Chipping Norton, Oxfordshire, England, in 1763. He 'accidentally' tasted the bark and found it had a bitter taste, which reminded him of Peruvian Bark ( Cinchona), which was used to treat
malaria Malaria is a mosquito-borne infectious disease that affects humans and other animals. Malaria causes symptoms that typically include fever, tiredness, vomiting, and headaches. In severe cases, it can cause jaundice, seizures, coma, or death. S ...
. After researching all the 'dispensaries and books on botany,' he found no suggestion of willow ever being used to treat fevers, and decided to experiment with it himself. Over the next seven years he successfully used the dried powder of willow bark to cure 'agues and intermittent fevers' of around fifty people, although it worked better when combined with quinine. Stone appears to have been largely ignored by the medical profession and herbalists alike. There are reports of two pharmacists trialling this remedy, but no evidence that it worked. By the early 20th century, Maud Grieve, one of the most famous herbalists of the era, did not consider White Willow to be a febrifuge. Instead, she describes using the bark and the powdered root for its tonic, antiperiodic and
astringent An astringent (sometimes called adstringent) is a chemical that shrinks or constricts body tissues. The word derives from the Latin ''adstringere'', which means "to bind fast". Calamine lotion, witch hazel, and yerba mansa, a Californian pla ...
qualities and recommended its use in treating
dyspepsia Indigestion, also known as dyspepsia or upset stomach, is a condition of impaired digestion. Symptoms may include upper abdominal fullness, heartburn, nausea, belching, or upper abdominal pain. People may also experience feeling full earlier ...
,
worms Worms may refer to: *Worm, an invertebrate animal with a tube-like body and no limbs Places *Worms, Germany Worms () is a city in Rhineland-Palatinate, Germany, situated on the Upper Rhine about south-southwest of Frankfurt am Main. It had ...
, chronic
diarrhoea Diarrhea, also spelled diarrhoea, is the condition of having at least three loose, liquid, or watery bowel movements each day. It often lasts for a few days and can result in dehydration due to fluid loss. Signs of dehydration often begin wi ...
and
dysentery Dysentery (UK pronunciation: , US: ), historically known as the bloody flux, is a type of gastroenteritis that results in bloody diarrhea. Other symptoms may include fever, abdominal pain, and a feeling of incomplete defecation. Complications ...
. She considered
tannin Tannins (or tannoids) are a class of astringent, polyphenolic biomolecules that bind to and precipitate proteins and various other organic compounds including amino acids and alkaloids. The term ''tannin'' (from Anglo-Norman ''tanner'' ...
to be the active constituent. An active extract of the bark, called
salicin Salicin is an alcoholic β- glucoside. Salicin is produced in (and named after) willow (''Salix'') bark. It is a biosynthetic precursor to salicylaldehyde. Medicinal aspects Salicin is found in the bark of and leaves of willows, poplars and v ...
, after the Latin name ''Salix'', was isolated to its crystalline form in 1828 by Henri Leroux, a French pharmacist, and Raffaele Piria, an Italian chemist, who then succeeded in separating out the acid in its pure state. Salicylic acid is a chemical derivative of salicin, and is widely used in medicine. Acetylsalicylic acid (
Aspirin Aspirin, also known as acetylsalicylic acid (ASA), is a nonsteroidal anti-inflammatory drug (NSAID) used to reduce pain, fever, and/or inflammation, and as an antithrombotic. Specific inflammatory conditions which aspirin is used to treat inc ...
) is, however, a chemical that does not occur in nature and which was originally synthesised from salicylic acid extracted from Meadowsweet, and is not connected to willow except in the similarity of its name.


References


Further reading

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External links

*
''Salix alba''
— information, genetic conservation units and related resources.
European Forest Genetic Resources Programme European Forest Genetic Resources Programme (EUFORGEN) is an international network that supports the conservation and sustainable use of forest genetic resources in Europe. The programme’s tasks include to coordinate and promote '' in situ'' an ...
(EUFORGEN) * * * * * {{Authority control
alba ''Alba'' ( , ) is the Scottish Gaelic name for Scotland. It is also, in English language historiography, used to refer to the polity of Picts and Scots united in the ninth century as the Kingdom of Alba, until it developed into the Kingdom ...
Flora of North Africa Flora of temperate Asia Flora of Europe Medicinal plants Garden plants of Asia Garden plants of Europe Ornamental trees Plants described in 1753 Nonsteroidal anti-inflammatory drugs Taxa named by Carl Linnaeus