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Walden's hornbill (''Rhabdotorrhinus waldeni'') locally called dulungan, also known as the Visayan wrinkled hornbill, rufous-headed hornbill or writhed-billed hornbill, is a critically endangered species of
hornbill Hornbills (Bucerotidae) are a family (biology), family of bird found in tropical and subtropical Africa, Asia and Melanesia. They are characterized by a long, down-curved bill which is frequently brightly coloured and sometimes has a Casque (an ...
living in the rainforests on the islands of
Negros Negros is the fourth largest and third most populous island in the Philippines, with a total land area of . Negros is one of the many islands of the Visayas, in the central part of the country. The predominant inhabitants of the island region a ...
and
Panay Panay is the sixth-largest and fourth-most populous island in the Philippines, with a total land area of and has a total population of 4,542,926 as of 2020 census. Panay comprises 4.4 percent of the entire population of the country. The City o ...
in the
Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...
. It is closely related to the
writhed hornbill The writhed hornbill (''Rhabdotorrhinus leucocephalus''), also known as the Mindanao wrinkled hornbill, is a species of hornbill in the family Bucerotidae. It is endemic to humid forests on the Philippine islands of Mindanao, Dinagat and Camig ...
, but can be recognized by the yellow throat and ocular skin in the male, and the blue throat and ocular skin in the female (both throat and ocular skin are deep orange or red in both sexes of the writhed hornbill). Its
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
commemorates the Scottish ornithologist Viscount Walden. It is considered one of the
Western Visayas Western Visayas ( hil, Kabisay-an Nakatundan; tl, Kanlurang Kabisayaan or ''Kanlurang Visayas'') is an administrative region in the Philippines, numerically designated as Region VI. It consists of six provinces (Aklan, Antique, Capiz, Guimara ...
Big 5 which includes the
Negros bleeding-heart pigeon The Negros bleeding-heart pigeon (''Gallicolumba keayi'') is endemic to the Philippines where it is found on the islands of Negros and Panay. It is critically endangered; continuing rates of forest loss on the two islands where it occurs sugges ...
,
Visayan spotted deer The Visayan spotted deer (''Rusa alfredi''), also known as the Visayan deer, the Philippine spotted deer or Prince Alfred's deer, is a nocturnal and endangered species of deer located primarily in the rainforests of the Visayan islands of Panay ...
,
Visayan hornbill The Visayan hornbill (''Penelopides panini'') is a hornbill found in rainforests on the islands of Panay Island, Panay, Negros Island, Negros, Masbate, and Guimaras Island, Guimaras, and formerly Ticao Island, Ticao, in the Philippines. It former ...
and the
Visayan warty pig The Visayan warty pig (''Sus cebifrons'') is a critically endangered species in the pig genus ('' Sus''). It is endemic to six of the Visayan Islands (Cebu, Negros, Panay, Masbate, Guimaras, and Siquijor) in the central Philippines. It is known ...
.


Description

EBird eBird is an online database of bird observations providing scientists, researchers and amateur naturalists with real-time data about bird distribution and abundance. Originally restricted to sightings from the Western Hemisphere, the project e ...
describes the bird as "A large rare bird of lowland and foothill forest on Panay and Negros, although almost extinct on the latter. Mostly black with a cream-colored tail with black terminal band, an orange
casque Casque is a French word for helmet. It can refer to: * Casque (anatomy), an enlargement on the beaks of some species of birds, including many hornbills *Hornbill ivory, the casque of the helmeted hornbill, collected as a decorative material * S. C ...
, and a wrinkled lower bill. Male has a rufous head, a yellow pouch, and yellow around the eye. Female has a black neck and head, a smaller bill and
casque Casque is a French word for helmet. It can refer to: * Casque (anatomy), an enlargement on the beaks of some species of birds, including many hornbills *Hornbill ivory, the casque of the helmeted hornbill, collected as a decorative material * S. C ...
, and blue and yellow around the eye and on the chin. The only other hornbill in range is the
Visayan hornbill The Visayan hornbill (''Penelopides panini'') is a hornbill found in rainforests on the islands of Panay Island, Panay, Negros Island, Negros, Masbate, and Guimaras Island, Guimaras, and formerly Ticao Island, Ticao, in the Philippines. It former ...
but Writhe-billed is larger, with a pale bill. Voice includes a three-noted throaty chuckle.." They exhibit sexual dimorphism with the males having a rufous head and yellow facial skin while the females having a black head and blue facial skin and a smaller bill.


Habitat and behaviour

It inhabits closed-canopy forests, also frequenting logged areas and occasionally isolated trees in clearings at an altitudinal range of 400 to 1,200 m
Panay Panay is the sixth-largest and fourth-most populous island in the Philippines, with a total land area of and has a total population of 4,542,926 as of 2020 census. Panay comprises 4.4 percent of the entire population of the country. The City o ...
and 350 to 950 m in
Negros Negros is the fourth largest and third most populous island in the Philippines, with a total land area of . Negros is one of the many islands of the Visayas, in the central part of the country. The predominant inhabitants of the island region a ...
. It prefers areas abundant fruit-bearing trees for feeding and large trees for nesting. They are known to live in small and noisy
flocks Flocking is the behaviour exhibited when a group of birds, called a flock, are foraging or in flight. Computer simulations and mathematical models that have been developed to emulate the flocking behaviours of birds can also generally be applie ...
. The Walden's hornbills are cavity nesters and use natural or carved-out hollows in tree trunks for its nest. As other hornbills, they reproduce very slowly. As lack of appropriate nesting sites is a problem,
nest box A nest is a structure built for certain animals to hold eggs or young. Although nests are most closely associated with birds, members of all classes of vertebrates and some invertebrates construct nests. They may be composed of organic materia ...
es are being attached to tree trunks in certain reserves. It is omnivorous, taking some animal matter to its nests and feeding in the canopy on figs and other fruits. It may make local nomadic movements in response to food availability. It nests in large trees.


Conservation status

This is a critically Endangered species, one of the three critically endangered hornbills including the
Sulu hornbill The Sulu hornbill (''Anthracoceros montani''), or Montano's hornbill, is a species of hornbill in the family Bucerotidae. It is endemic to the Sulu archipelago in the Philippines, with the remaining populations in Tawi-Tawi with it believed to ...
and
Helmeted hornbill The helmeted hornbill (''Rhinoplax vigil'') is a very large bird in the hornbill family. It is found on the Malay Peninsula, Sumatra, Borneo, Thailand and Myanmar. The casque (helmetlike structure on the head) accounts for some 11% of its 3  ...
.. It is presumed extinct on
Guimaras Guimaras , officially the Province of Guimaras ( hil, Kapuoran sang Guimaras; tl, Lalawigan ng Guimaras), is an island province in the Philippines located in the Western Visayas region. Its capital is Jordan while its largest local government ...
and now survives only on Negros and Panay The total population has been estimated at less than 160, though recent work from the
Central Panay Mountain Range Central Panay Mountain Range is the longest and largest mountain range in the island of Panay and Western Visayas in the Philippines. With a total length of long north–south and width east–west. Mount Madja-as is the highest point with a ...
suggests 600-700 pairs may remain there. No recent figures are available for Negros, where it may be
functionally extinct Functional extinction is the extinction of a species or other taxon such that: #It disappears from the fossil record, or historic reports of its existence cease; #The reduced population no longer plays a significant role in ecosystem function; or ...
. The
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biol ...
estimates the population to be 1,000 to 2,499 mature individuals with the population continuing to decline. Walden's hornbill reproduce very slowly and thus are unable to survive high hunting pressures coupled with heavy logging of the rainforests. By 2007,
Negros Negros is the fourth largest and third most populous island in the Philippines, with a total land area of . Negros is one of the many islands of the Visayas, in the central part of the country. The predominant inhabitants of the island region a ...
and
Panay Panay is the sixth-largest and fourth-most populous island in the Philippines, with a total land area of and has a total population of 4,542,926 as of 2020 census. Panay comprises 4.4 percent of the entire population of the country. The City o ...
had a 3% and 6% remaining forest cover with most of this being higher elevation forest where this bird does not thrive in. Despite already paltry forest cover, deforestation still continues thanks to both legal and
illegal logging Illegal logging is the harvest, transportation, purchase or sale of timber in violation of laws. The harvesting procedure itself may be illegal, including using corrupt means to gain access to forests; extraction without permission, or from a pro ...
, conversion into farmland,
mining Mining is the extraction of valuable minerals or other geological materials from the Earth, usually from an ore body, lode, vein, seam, reef, or placer deposit. The exploitation of these deposits for raw material is based on the economic via ...
and road development. As for the case of the
Northwest Panay Peninsula Natural Park The Northwest Panay Peninsula Natural Park is located on the island of Panay, in the provinces of Aklan and Antique of the Philippines which was proclaimed a natural park by President Gloria Macapagal Arroyo on 18 April 2002 (Presidential Procla ...
, a road was built splitting it from the
Central Panay Mountain Range Central Panay Mountain Range is the longest and largest mountain range in the island of Panay and Western Visayas in the Philippines. With a total length of long north–south and width east–west. Mount Madja-as is the highest point with a ...
and has severely affected the population in the former as these roads allow for easier access by hunters and loggers. Another huge threat is hunting and capture for the
illegal wildlife trade Wildlife trade refers to the of products that are derived from non-domesticated animals or plants usually extracted from their natural environment or raised under controlled conditions. It can involve the trade of living or dead individuals, ti ...
. According to the Philippine Initiative for Environmental Conservation (Philincon), up to 50% of all nests in
Central Panay Mountain Range Central Panay Mountain Range is the longest and largest mountain range in the island of Panay and Western Visayas in the Philippines. With a total length of long north–south and width east–west. Mount Madja-as is the highest point with a ...
were affected by poachers who would either climb up or cut down the nesting trees to poach the nestlings and the mother. Due to this high poaching rate, Philincon has devised a nest guarding program where they pay locals US$20 a month to guard hornbill nests and offer an incentive of US$11 for each successful fledge. This program has been effective cutting down the poaching rate to just 5%. Philincon now guards more than 1,000 nest holes which is believed to be at least two-thirds of all nests in the
Central Panay Mountain Range Central Panay Mountain Range is the longest and largest mountain range in the island of Panay and Western Visayas in the Philippines. With a total length of long north–south and width east–west. Mount Madja-as is the highest point with a ...
. In areas where their large nesting trees have been logged, Philincon installs nest boxes for these hornbills to breed in. The bird is listed as an
EDGE species Evolutionarily Distinct and Globally Endangered (EDGE) species are animal species which have a high 'EDGE score', a metric combining endangered conservation status with the genetic distinctiveness of the particular taxon. Distinctive species have ...
by the
Zoological Society of London The Zoological Society of London (ZSL) is a charity devoted to the worldwide conservation of animals and their habitats. It was founded in 1826. Since 1828, it has maintained the London Zoo, and since 1931 Whipsnade Park. History On 29 ...
where it is ranked as the 50th which uses the basis of evolutionary distinctness and endangeredness. This bird was bred in captivity for the first time by the Mari-it Wildlife and Conservation Park in 2005. As of 2010, they have bred 15 Walden's hornbills. In 2019, The Talarak Foundation has also successfully bred them in captivity in their
Negros Negros is the fourth largest and third most populous island in the Philippines, with a total land area of . Negros is one of the many islands of the Visayas, in the central part of the country. The predominant inhabitants of the island region a ...
facility after nine years of attempts. Further conservation actions proposed are: more funding should be allocated primarily to in situ protection along the lines of PhilCon's guarding scheme; conduct further surveys, particularly on
Panay Panay is the sixth-largest and fourth-most populous island in the Philippines, with a total land area of and has a total population of 4,542,926 as of 2020 census. Panay comprises 4.4 percent of the entire population of the country. The City o ...
to identify important sites and use this data to decide further actions; continue community awareness programmes to reduce hunting and illegal logging on both
Negros Negros is the fourth largest and third most populous island in the Philippines, with a total land area of . Negros is one of the many islands of the Visayas, in the central part of the country. The predominant inhabitants of the island region a ...
and
Panay Panay is the sixth-largest and fourth-most populous island in the Philippines, with a total land area of and has a total population of 4,542,926 as of 2020 census. Panay comprises 4.4 percent of the entire population of the country. The City o ...
; work in partnership at the government level to strengthen protected area legislation and improve the network in the long term; and support the development of captive breeding and reintroduction programmes.


References

*ARKive
images and movies of the Walden's hornbill ''(Aceros waldeni)''
{{Taxonbar, from=Q27074827 Rhabdotorrhinus Birds of Negros Island Birds of Panay Critically endangered biota of Asia Birds described in 1877 Taxa named by Richard Bowdler Sharpe