The wolverine (), (''Gulo gulo''; ''Gulo'' is
Latin
Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
for "
glutton"), also referred to as the glutton, carcajou, or quickhatch (from
East Cree
East Cree, also known as (Eastern) James Bay Cree, and East Main Cree, is a group of Cree dialects spoken in Quebec, Canada on the east coast of lower Hudson Bay and James Bay, and inland southeastward from James Bay. Cree is one of the most spok ...
, ''kwiihkwahaacheew''), is the largest land-dwelling
species
In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of the family
Mustelidae
The Mustelidae (; from Latin ''mustela'', weasel) are a family of carnivorous mammals, including weasels, badgers, otters, ferrets, martens, minks and wolverines, among others. Mustelids () are a diverse group and form the largest family in ...
. It is a muscular
carnivore
A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose food and energy requirements derive from animal tissues (mainly muscle, fat and other sof ...
and a
solitary animal
Sociality is the degree to which individuals in an animal population tend to associate in social groups (gregariousness) and form cooperative societies.
Sociality is a survival response to evolutionary pressures. For example, when a mother wasp ...
.
The wolverine has a reputation for ferocity and strength out of proportion to its size, with the documented ability to kill prey many times larger than itself.
The wolverine is found primarily in remote reaches of the Northern
boreal forests
Taiga (; rus, тайга́, p=tɐjˈɡa; relates to Mongolic and Turkic languages), generally referred to in North America as a boreal forest or snow forest, is a biome characterized by coniferous forests consisting mostly of pines, spruces ...
and
subarctic
The subarctic zone is a region in the Northern Hemisphere immediately south of the true Arctic, north of humid continental regions and covering much of Alaska, Canada, Iceland, the north of Scandinavia, Siberia, and the Cairngorms. Generally, ...
and
alpine tundra
Alpine tundra is a type of natural region or biome that does not contain trees because it is at high elevation, with an associated alpine climate, harsh climate. As the latitude of a location approaches the poles, the threshold elevation for alp ...
of the Northern Hemisphere, with the greatest numbers in
Northern Canada
Northern Canada, colloquially the North or the Territories, is the vast northernmost region of Canada variously defined by geography and politics. Politically, the term refers to the three Provinces_and_territories_of_Canada#Territories, territor ...
, the
U.S. state
In the United States, a state is a constituent political entity, of which there are 50. Bound together in a political union, each state holds governmental jurisdiction over a separate and defined geographic territory where it shares its sover ...
of
Alaska
Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S., ...
, the mainland
Nordic countries
The Nordic countries (also known as the Nordics or ''Norden''; literal translation, lit. 'the North') are a geographical and cultural region in Northern Europe and the Atlantic Ocean, North Atlantic. It includes the sovereign states of Denmar ...
of Europe, and throughout western Russia and
Siberia
Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part of ...
. Its population has steadily declined since the 19th century owing to
trapping
Animal trapping, or simply trapping or gin, is the use of a device to remotely catch an animal. Animals may be trapped for a variety of purposes, including food, the fur trade, hunting, pest control, and wildlife management.
History
Neolithic ...
, range reduction and
habitat fragmentation
Habitat fragmentation describes the emergence of discontinuities (fragmentation) in an organism's preferred environment (habitat), causing population fragmentation and ecosystem decay. Causes of habitat fragmentation include geological processes ...
. The wolverine is now essentially absent from the southern end of its range in both Europe and North America.
Taxonomy
![Gulo gulo](https://upload.wikimedia.org/wikipedia/commons/8/89/Gulo_gulo.jpg)
Genetic evidence suggests that the wolverine is most closely related to the
tayra
The tayra (''Eira barbara'') is an omnivorous animal from the weasel family, native to the Americas. It is the only species in the genus ''Eira''.
Tayras are also known as the ''tolomuco'' or ''perico ligero'' in Central America, ''motete'' in ...
and
martens
A marten is a weasel-like mammal in the genus ''Martes'' within the subfamily Guloninae, in the family Mustelidae. They have bushy tails and large paws with partially retractile claws. The fur varies from yellowish to dark brown, depending on t ...
, all of which shared a Eurasian ancestor.
There are two
subspecies
In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
: the
Old World
The "Old World" is a term for Afro-Eurasia that originated in Europe , after Europeans became aware of the existence of the Americas. It is used to contrast the continents of Africa, Europe, and Asia, which were previously thought of by the ...
form, ''Gulo gulo gulo'', and the
New World
The term ''New World'' is often used to mean the majority of Earth's Western Hemisphere, specifically the Americas."America." ''The Oxford Companion to the English Language'' (). McArthur, Tom, ed., 1992. New York: Oxford University Press, p. 3 ...
form, ''G. g. luscus''. Some authors had described as many as four additional North American subspecies, including ones limited to
Vancouver Island
Vancouver Island is an island in the northeastern Pacific Ocean and part of the Canadian Provinces and territories of Canada, province of British Columbia. The island is in length, in width at its widest point, and in total area, while are o ...
(''G. g. vancouverensis'') and the
Kenai Peninsula
The Kenai Peninsula ( Dena'ina: ''Yaghenen'') is a large peninsula jutting from the coast of Southcentral Alaska. The name Kenai (, ) is derived from the word "Kenaitze" or "Kenaitze Indian Tribe", the name of the Native Athabascan Alaskan tribe ...
in Alaska (''G. g. katschemakensis''). However, the most currently accepted taxonomy recognizes either the two continental
subspecies
In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
or ''G. gulo'' as a single
Holarctic
The Holarctic realm is a biogeographic realm that comprises the majority of habitats found throughout the continents in the Northern Hemisphere. It corresponds to the floristic Boreal Kingdom. It includes both the Nearctic zoogeographical region ...
taxon.
Recently compiled genetic evidence suggests most of North America's wolverines are descended from a single source, likely originating from
Beringia
Beringia is defined today as the land and maritime area bounded on the west by the Lena River in Russia; on the east by the Mackenzie River in Canada; on the north by 72 degrees north latitude in the Chukchi Sea; and on the south by the tip ...
during the last glaciation and rapidly expanding thereafter, though considerable uncertainty to this conclusion is due to the difficulty of collecting samples in the extremely depleted southern extent of the range.
Physical characteristics
Anatomically, the wolverine is an elongated animal that is low to the ground. With strong limbs, broad and rounded head, small eyes and short rounded ears, it most closely resembles a large
fisher
Fisher is an archaic term for a fisherman, revived as gender-neutral.
Fisher, Fishers or The Fisher may also refer to:
Places
Australia
*Division of Fisher, an electoral district in the Australian House of Representatives, in Queensland
*Elect ...
. Though its legs are short, its large, five-toed paws with crampon-like claws and
plantigrade
151px, Portion of a human skeleton, showing plantigrade habit
In terrestrial animals, plantigrade locomotion means walking with the toes and metatarsals flat on the ground. It is one of three forms of locomotion adopted by terrestrial mammals. T ...
posture enable it to climb up and over steep cliffs, trees and snow-covered peaks with relative ease.
The adult wolverine is about the size of a medium dog, with a body length ranging from ; standing at the shoulder; and a tail length of . Males range in weight from , and females . Exceptionally large males of as much as are referenced in Russian literature, though such weights are deemed in ''Mammals of the Soviet Union'' to be improbable.
[Holbrow, W. C. (1976). ''The biology, mythology, distribution, and management of the wolverine (Gulo gulo) in western Canada''. The University of Manitoba (Masters Thesis).] The males are often 10–15% larger than the females in linear measurements and can be 30–40% greater in weight. According to some sources, Eurasian wolverines are claimed to be larger and heavier than North American, with average weights in excess of . However, this may refer more specifically to areas such as
Siberia
Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part of ...
, as data from
Fennoscandian wolverines shows they are typically around the same size as their American counterparts.
[ It is the largest of terrestrial mustelids; only the marine-dwelling ]sea otter
The sea otter (''Enhydra lutris'') is a marine mammal native to the coasts of the northern and eastern North Pacific Ocean. Adult sea otters typically weigh between , making them the heaviest members of the weasel family, but among the small ...
, the giant otter
The giant otter or giant river otter (''Pteronura brasiliensis'') is a South American carnivorous mammal. It is the longest member of the weasel family, Mustelidae, a globally successful group of predators, reaching up to . Atypical of musteli ...
of the Amazon basin and the semi-aquatic African clawless otter
The African clawless otter (''Aonyx capensis''), also known as the Cape clawless otter or groot otter, is the second-largest freshwater otter species. It inhabits permanent water bodies in savannah and lowland forest areas through most of sub- ...
are larger—while the European badger
The European badger (''Meles meles''), also known as the Eurasian badger, is a badger species in the family Mustelidae native to almost all of Europe. It is classified as least concern on the IUCN Red List as it has a wide range and a large stabl ...
may reach a similar body mass, especially in autumn.
Wolverines have thick, dark, oily fur which is highly hydrophobic
In chemistry, hydrophobicity is the physical property of a molecule that is seemingly repelled from a mass of water (known as a hydrophobe). In contrast, hydrophiles are attracted to water.
Hydrophobic molecules tend to be nonpolar and, th ...
, making it resistant to frost. This has led to its traditional popularity among hunters and trappers as a lining in jackets and parkas
Parkas was a Canadian pop music band based in London, Ontario and later Toronto.["P ...]
in Arctic conditions. A light-silvery facial mask is distinct in some individuals, and a pale buff stripe runs laterally from the shoulders along the side and crossing the rump just above a bushy tail. Some individuals display prominent white hair patches on their throats or chests.
Like many other mustelids, it has potent anal scent glands
Anal may refer to:
Related to the anus
*Related to the anus of animals:
** Anal fin, in fish anatomy
** Anal vein, in insect anatomy
** Anal scale, in reptile anatomy
*Related to the human anus:
** Anal sex, a type of sexual activity involving s ...
used for marking territory
In ethology, territory is the sociographical area that an animal consistently defends against conspecific competition (or, occasionally, against animals of other species) using agonistic behaviors or (less commonly) real physical aggression. ...
and sexual signaling. The pungent odor has given rise to the nicknames "skunk bear" and "nasty cat." Wolverines, like other mustelids
The Mustelidae (; from Latin ''mustela'', weasel) are a family of carnivorous mammals, including weasels, badgers, otters, ferrets, martens, minks and wolverines, among others. Mustelids () are a diverse group and form the largest famil ...
, possess a special upper molar in the back of the mouth that is rotated 90 degrees, towards the inside of the mouth. This special characteristic allows wolverines to tear off meat from prey or carrion that has been frozen solid.
Behavior
Diet and hunting
Wolverines are considered to be primarily scavenger
Scavengers are animals that consume dead organisms that have died from causes other than predation or have been killed by other predators. While scavenging generally refers to carnivores feeding on carrion, it is also a herbivorous feeding b ...
s. A majority of the wolverine's sustenance is derived from carrion
Carrion () is the decaying flesh of dead animals, including human flesh.
Overview
Carrion is an important food source for large carnivores and omnivores in most ecosystems. Examples of carrion-eaters (or scavengers) include crows, vultures, c ...
, on which it depends almost exclusively in winter and early spring. Wolverines may find carrion themselves, feed on it after the predator (often, a pack of wolves
The wolf (''Canis lupus''; plural, : wolves), also known as the gray wolf or grey wolf, is a large Canis, canine native to Eurasia and North America. More than thirty subspecies of Canis lupus, subspecies of ''Canis lupus'' have been reco ...
) has finished, or simply take it from another predator. Wolverines are also known to follow wolf and lynx trails, purportedly with the intent of scavenging the remains of their kills. Whether eating live prey or carrion, the wolverine's feeding style appears voracious, leading to the nickname of "glutton" (also the basis of the scientific name). However, this feeding style is believed to be an adaptation to food scarcity, especially in winter.
The wolverine is also a powerful and versatile predator. Its prey mainly consists of small to medium-sized mammals, but the wolverine has been recorded killing prey such as adult deer that are many times larger than itself. Prey species include porcupine
Porcupines are large rodents with coats of sharp spines, or quills, that protect them against predation. The term covers two families of animals: the Old World porcupines of family Hystricidae, and the New World porcupines of family, Erethizont ...
s, squirrel
Squirrels are members of the family Sciuridae, a family that includes small or medium-size rodents. The squirrel family includes tree squirrels, ground squirrels (including chipmunks and prairie dogs, among others), and flying squirrels. Squ ...
s, chipmunk
Chipmunks are small, striped rodents of the family Sciuridae. Chipmunks are found in North America, with the exception of the Siberian chipmunk which is found primarily in Asia.
Taxonomy and systematics
Chipmunks may be classified either as ...
s, beaver
Beavers are large, semiaquatic rodents in the genus ''Castor'' native to the temperate Northern Hemisphere. There are two extant species: the North American beaver (''Castor canadensis'') and the Eurasian beaver (''C. fiber''). Beavers ar ...
s, marmot
Marmots are large ground squirrels in the genus ''Marmota'', with 15 species living in Asia, Europe, and North America. These herbivores are active during the summer, when they can often be found in groups, but are not seen during the winter, w ...
s, moles Moles can refer to:
* Moles de Xert, a mountain range in the Baix Maestrat comarca, Valencian Community, Spain
* The Moles (Australian band)
*The Moles, alter ego of Scottish band Simon Dupree and the Big Sound
People
*Abraham Moles, French engin ...
, gopher
Pocket gophers, commonly referred to simply as gophers, are burrowing rodents of the family Geomyidae. The roughly 41 speciesSearch results for "Geomyidae" on thASM Mammal Diversity Database are all endemic to North and Central America. They are ...
s, rabbit
Rabbits, also known as bunnies or bunny rabbits, are small mammals in the family Leporidae (which also contains the hares) of the order Lagomorpha (which also contains the pikas). ''Oryctolagus cuniculus'' includes the European rabbit speci ...
s, vole
Voles are small rodents that are relatives of lemmings and hamsters, but with a stouter body; a longer, hairy tail; a slightly rounder head; smaller eyes and ears; and differently formed molars (high-crowned with angular cusps instead of low-c ...
s, mice, rats, shrew
Shrews (family Soricidae) are small mole-like mammals classified in the order Eulipotyphla. True shrews are not to be confused with treeshrews, otter shrews, elephant shrews, West Indies shrews, or marsupial shrews, which belong to different fa ...
s, lemming
A lemming is a small rodent, usually found in or near the Arctic in tundra biomes. Lemmings form the subfamily Arvicolinae (also known as Microtinae) together with voles and muskrats, which form part of the superfamily Muroidea, which also include ...
s, caribou
Reindeer (in North American English, known as caribou if wild and ''reindeer'' if domesticated) are deer in the genus ''Rangifer''. For the last few decades, reindeer were assigned to one species, ''Rangifer tarandus'', with about 10 subspe ...
, roe deer
The roe deer (''Capreolus capreolus''), also known as the roe, western roe deer, or European roe, is a species of deer. The male of the species is sometimes referred to as a roebuck. The roe is a small deer, reddish and grey-brown, and well-adapt ...
, white-tailed deer
The white-tailed deer (''Odocoileus virginianus''), also known as the whitetail or Virginia deer, is a medium-sized deer native to North America, Central America, and South America as far south as Peru and Bolivia. It has also been introduced t ...
, mule deer
The mule deer (''Odocoileus hemionus'') is a deer indigenous to western North America; it is named for its ears, which are large like those of the mule. Two subspecies of mule deer are grouped into the black-tailed deer.
Unlike the related whit ...
, sheep, goats, cattle, bison
Bison are large bovines in the genus ''Bison'' (Greek: "wild ox" (bison)) within the tribe Bovini. Two extant and numerous extinct species are recognised.
Of the two surviving species, the American bison, ''B. bison'', found only in North Ame ...
, moose
The moose (in North America) or elk (in Eurasia) (''Alces alces'') is a member of the New World deer subfamily and is the only species in the genus ''Alces''. It is the largest and heaviest extant species in the deer family. Most adult mal ...
, and elk
The elk (''Cervus canadensis''), also known as the wapiti, is one of the largest species within the deer family, Cervidae, and one of the largest terrestrial mammals in its native range of North America and Central and East Asia. The common ...
. Smaller predators are occasionally preyed on, including marten
A marten is a weasel-like mammal in the genus ''Martes'' within the subfamily Guloninae, in the family Mustelidae. They have bushy tails and large paws with partially retractile claws. The fur varies from yellowish to dark brown, depending on t ...
s, mink
Mink are dark-colored, semiaquatic, carnivorous mammals of the genera ''Neogale'' and '' Mustela'' and part of the family Mustelidae, which also includes weasels, otters, and ferrets. There are two extant species referred to as "mink": the A ...
, foxes, Eurasian lynx
The Eurasian lynx (''Lynx lynx'') is a medium-sized wild cat widely distributed from Northern, Central and Eastern Europe to Central Asia and Siberia, the Tibetan Plateau and the Himalayas. It inhabits temperate and boreal forests up to an eleva ...
,[Heptner, V.G. and Sludskii, A.A. (1992). Humans are apparently exempt. ''Mammals of the Soviet Union. Volume II Part 2 Carnivora: Hyenas and Cats''. New Delhi: Amerind Publishing, p. 625] weasel
Weasels are mammals of the genus ''Mustela'' of the family Mustelidae. The genus ''Mustela'' includes the least weasels, polecats, stoats, ferrets and European mink. Members of this genus are small, active predators, with long and slender bo ...
s, coyote
The coyote (''Canis latrans'') is a species of canis, canine native to North America. It is smaller than its close relative, the wolf, and slightly smaller than the closely related eastern wolf and red wolf. It fills much of the same ecologica ...
, and wolf
The wolf (''Canis lupus''; : wolves), also known as the gray wolf or grey wolf, is a large canine native to Eurasia and North America. More than thirty subspecies of ''Canis lupus'' have been recognized, and gray wolves, as popularly u ...
pups. Wolverines have also been known to kill Canada lynx
The Canada lynx (''Lynx canadensis''), or Canadian lynx, is a medium-sized North American lynx that ranges across Alaska, Canada, and northern areas of the contiguous United States. It is characterized by its long, dense fur, triangular ears ...
in Yukon
Yukon (; ; formerly called Yukon Territory and also referred to as the Yukon) is the smallest and westernmost of Canada's three territories. It also is the second-least populated province or territory in Canada, with a population of 43,964 as ...
, Canada. Wolverines often pursue live prey that are relatively easy to obtain, including animals caught in traps, newborn mammals, and deer (including adult moose and elk) when they are weakened by winter or immobilized by heavy snow. Their diets are sometimes supplemented by birds' eggs, birds (especially geese
A goose (plural, : geese) is a bird of any of several waterfowl species in the family (biology), family Anatidae. This group comprises the genera ''Anser (bird), Anser'' (the grey geese and white geese) and ''Branta'' (the black geese). Some o ...
), root
In vascular plants, the roots are the organs of a plant that are modified to provide anchorage for the plant and take in water and nutrients into the plant body, which allows plants to grow taller and faster. They are most often below the sur ...
s, seed
A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiospe ...
s, insect larvae, and berries
A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone or pit, although many pips or seeds may be present. Common examples are strawberries, raspb ...
.
Wolverines inhabiting the Old World
The "Old World" is a term for Afro-Eurasia that originated in Europe , after Europeans became aware of the existence of the Americas. It is used to contrast the continents of Africa, Europe, and Asia, which were previously thought of by the ...
(specifically, Fennoscandia
__NOTOC__
Fennoscandia (Finnish language, Finnish, Swedish language, Swedish and no, Fennoskandia, nocat=1; russian: Фенноскандия, Fennoskandiya) or the Fennoscandian Peninsula is the geographical peninsula in Europe, which includes ...
) hunt more actively than their North American relatives. This may be because competing predator populations in Eurasia are not as dense, making it more practical for the wolverine to hunt for itself than to wait for another animal to make a kill and then try to snatch it. They often feed on carrion left by wolves, so changes in wolf populations may affect the population of wolverines. They are also known on occasion to eat plant material.
Wolverines frequently cache
Cache, caching, or caché may refer to:
Places United States
* Cache, Idaho, an unincorporated community
* Cache, Illinois, an unincorporated community
* Cache, Oklahoma, a city in Comanche County
* Cache, Utah, Cache County, Utah
* Cache Count ...
their food during times of plenty. This is of particular importance to lactating females in the winter and early spring, a time when food is scarce.
Natural enemies
Wolves
The wolf (''Canis lupus''; plural, : wolves), also known as the gray wolf or grey wolf, is a large Canis, canine native to Eurasia and North America. More than thirty subspecies of Canis lupus, subspecies of ''Canis lupus'' have been reco ...
, American black bear
The American black bear (''Ursus americanus''), also called simply a black bear or sometimes a baribal, is a medium-sized bear endemic to North America. It is the continent's smallest and most widely distributed bear species. American black bear ...
s, brown bear
The brown bear (''Ursus arctos'') is a large bear species found across Eurasia and North America. In North America, the populations of brown bears are called grizzly bears, while the subspecies that inhabits the Kodiak Islands of Alaska is kno ...
s, cougar
The cougar (''Puma concolor'') is a large Felidae, cat native to the Americas. Its Species distribution, range spans from the Canadian Yukon to the southern Andes in South America and is the most widespread of any large wild terrestrial mamm ...
s, and golden eagle
The golden eagle (''Aquila chrysaetos'') is a bird of prey living in the Northern Hemisphere. It is the most widely distributed species of eagle. Like all eagles, it belongs to the family Accipitridae. They are one of the best-known bird of p ...
s are capable of killing wolverines, particularly young and inexperienced individuals. Wolves are thought to be the wolverine's most important natural predator, with the arrival of wolves to a wolverine's territory presumably leading the latter to abandon the area. Armed with powerful jaws, sharp claws, and a thick hide, wolverines, like most mustelids, are remarkably strong for their size. They may defend against larger or more numerous predator
Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill th ...
s such as wolves or bears. By far, their most serious predator is the grey wolf
The wolf (''Canis lupus''; : wolves), also known as the gray wolf or grey wolf, is a large canine native to Eurasia and North America. More than thirty subspecies of ''Canis lupus'' have been recognized, and gray wolves, as popularly ...
, with an extensive record of wolverine fatalities attributed to wolves in both North America and Eurasia. In North America, another predator (less frequent) is the cougar
The cougar (''Puma concolor'') is a large Felidae, cat native to the Americas. Its Species distribution, range spans from the Canadian Yukon to the southern Andes in South America and is the most widespread of any large wild terrestrial mamm ...
. At least one account reported a wolverine's apparent attempt to steal a kill from a black bear
Black bear or Blackbear may refer to:
Animals
* American black bear (''Ursus americanus''), a North American bear species
* Asian black bear (''Ursus thibetanus''), an Asian bear species
Music
* Black Bear (band), a Canadian First Nations group ...
, although the bear won what was ultimately a fatal contest for the wolverine. There are a few accounts of brown bear
The brown bear (''Ursus arctos'') is a large bear species found across Eurasia and North America. In North America, the populations of brown bears are called grizzly bears, while the subspecies that inhabits the Kodiak Islands of Alaska is kno ...
s killing and consuming wolverines as well and, although also reported at times to be chased off prey, in some areas such as Denali National Park
Denali National Park and Preserve, formerly known as Mount McKinley National Park, is an American national park and preserve located in Interior Alaska, centered on Denali, the highest mountain in North America. The park and contiguous preserve e ...
, wolverines seemed to try to actively avoid encounters with grizzly bears as they have been reported in areas where wolves start hunting them.[Murie, A. (2012). ''The grizzlies of Mount McKinley''. University of Washington Press.]
Mating and reproduction
Successful males will form lifetime relationships with two or three females, which they will visit occasionally, while other males are left without a mate. Mating season is in the summer, but the actual implantation of the embryo (blastocyst) in the uterus
The uterus (from Latin ''uterus'', plural ''uteri'') or womb () is the organ in the reproductive system of most female mammals, including humans that accommodates the embryonic and fetal development of one or more embryos until birth. The uter ...
is stayed until early winter, delaying the development of the fetus
A fetus or foetus (; plural fetuses, feti, foetuses, or foeti) is the unborn offspring that develops from an animal embryo. Following embryonic development the fetal stage of development takes place. In human prenatal development, fetal deve ...
. Females will often not produce young if food is scarce. The gestation period is 30–50 days, and litters of typically two or three young ("kits") are born in the spring. Kits develop rapidly, reaching adult size within the first year. The typical longevity of a wolverine in captivity is around 15 to 17 years, but in the wild the average lifespan is more likely between 8 and 10 years. Fathers make visits to their offspring until they are weaned at 10 weeks of age; also, once the young are about six months old, some reconnect with their fathers and travel together for a time.
Distribution
Wolverines live primarily in isolated arctic
The Arctic ( or ) is a polar regions of Earth, polar region located at the northernmost part of Earth. The Arctic consists of the Arctic Ocean, adjacent seas, and parts of Canada (Yukon, Northwest Territories, Nunavut), Danish Realm (Greenla ...
, boreal, and alpine
Alpine may refer to any mountainous region. It may also refer to:
Places Europe
* Alps, a European mountain range
** Alpine states, which overlap with the European range
Australia
* Alpine, New South Wales, a Northern Village
* Alpine National Pa ...
regions of northern Canada, Alaska
Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S., ...
, Siberia
Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part of ...
, and Fennoscandia
__NOTOC__
Fennoscandia (Finnish language, Finnish, Swedish language, Swedish and no, Fennoskandia, nocat=1; russian: Фенноскандия, Fennoskandiya) or the Fennoscandian Peninsula is the geographical peninsula in Europe, which includes ...
; they are also native to European Russia
European Russia (russian: Европейская Россия, russian: европейская часть России, label=none) is the western and most populated part of Russia. It is geographically situated in Europe, as opposed to the cou ...
, the Baltic
Baltic may refer to:
Peoples and languages
* Baltic languages, a subfamily of Indo-European languages, including Lithuanian, Latvian and extinct Old Prussian
*Balts (or Baltic peoples), ethnic groups speaking the Baltic languages and/or originati ...
countries, the Russian Far East
The Russian Far East (russian: Дальний Восток России, r=Dal'niy Vostok Rossii, p=ˈdalʲnʲɪj vɐˈstok rɐˈsʲiɪ) is a region in Northeast Asia. It is the easternmost part of Russia and the Asian continent; and is admini ...
, northeast China and Mongolia. In the Sierra Nevada
The Sierra Nevada () is a mountain range in the Western United States, between the Central Valley of California and the Great Basin. The vast majority of the range lies in the state of California, although the Carson Range spur lies primarily ...
, wolverines were sighted near Winnemucca Lake in spring 1995 and at Toe Jam Lake north of the Yosemite border in 1996; and later photographed by baited cameras, including in 2008 and 2009, near Lake Tahoe
Lake Tahoe (; was, Dáʔaw, meaning "the lake") is a Fresh water, freshwater lake in the Sierra Nevada (U.S.), Sierra Nevada of the United States. Lying at , it straddles the state line between California and Nevada, west of Carson City, Nevad ...
. According to a 2014 U.S. Fish and Wildlife Service
The United States Fish and Wildlife Service (USFWS or FWS) is an agency within the United States Department of the Interior dedicated to the management of fish, wildlife, and natural habitats. The mission of the agency is "working with oth ...
publication, "wolverines are found in the North Cascades
The North Cascades are a section of the Cascade Range of western North America. They span the border between the Canadian province of British Columbia and the U.S. state of Washington and are officially named in the U.S. and Canada as the Cascad ...
in Washington and the Northern Rocky Mountains in Idaho, Montana, Oregon (Wallowa Range
The Wallowa Mountains () are a mountain range located in the Columbia Plateau of northeastern Oregon in the United States. The range runs approximately northwest to southeast in southwestern Wallowa County, Oregon, Wallowa County and eastern Uni ...
), and Wyoming. Individual wolverines have also moved into historic range in the Sierra Nevada Mountains of California and the Southern Rocky Mountains of Colorado, but have not established breeding populations in these areas." In 2022, Colorado Parks and Wildlife
Colorado Parks and Wildlife manages the state parks system and the wildlife of the U.S. state of Colorado. , the division managed the 42 state parks and 307 wildlife areas of Colorado.
, the Colorado Natural Areas Program had 93 designated si ...
considered plans to reintroduce the wolverine to the state.
Wolverines are also found in Utah but are very rarely seen, with only 6 confirmed sightings since the first confirmed sighting in 1979. 3 of these 6 confirmed Utah sightings have been caught on video. A wolverine, a male, was finally captured and tagged in Utah in 2022 before being released back into the wild to better understand the animal's range.
In August 2020, the National Park Service
The National Park Service (NPS) is an agency of the United States federal government within the U.S. Department of the Interior that manages all national parks, most national monuments, and other natural, historical, and recreational propertie ...
reported that wolverines had been sighted at Mount Rainier
Mount Rainier (), indigenously known as Tahoma, Tacoma, Tacobet, or təqʷubəʔ, is a large active stratovolcano in the Cascade Range of the Pacific Northwest, located in Mount Rainier National Park about south-southeast of Seattle. With a s ...
, Washington
Washington commonly refers to:
* Washington (state), United States
* Washington, D.C., the capital of the United States
** A metonym for the federal government of the United States
** Washington metropolitan area, the metropolitan area centered on ...
, for the first time in more than a century. The sighting was of a reproductive female and her two offspring.
In 2004, the first confirmed sighting of a wolverine in Michigan since the early 19th century took place, when a Michigan Department of Natural Resources
The Michigan Department of Natural Resources (DNR) is the agency of the state of Michigan charged with maintaining natural resources such as state parks, state forests, and recreation areas. It is governed by a director appointed by the Governor a ...
wildlife biologist photographed a wolverine in Ubly, Michigan
Ubly is a village in Huron County in the U.S. state of Michigan. The population was 858 at the 2010 census. The village is within Bingham Township.
On February 25, 2004, a wolverine was spotted near Ubly, the first one sighted in Michigan in o ...
. The specimen was found dead at the Minden City State Game Area in Sanilac County, Michigan
Sanilac County ( ) is a county located in the Thumb region of the U.S. state of Michigan. As of the 2020 Census, the population was 40,611. The county seat is Sandusky. The county was created on September 10, 1822, and was fully organized on D ...
in 2010; no further wolverines have been spotted in Michigan.
Most New World
The term ''New World'' is often used to mean the majority of Earth's Western Hemisphere, specifically the Americas."America." ''The Oxford Companion to the English Language'' (). McArthur, Tom, ed., 1992. New York: Oxford University Press, p. 3 ...
wolverines live in Canada and Alaska. However, wolverines were once recorded as also being present in Colorado, areas of the southwestern United States
The Southwestern United States, also known as the American Southwest or simply the Southwest, is a geographic and cultural region of the United States that generally includes Arizona, New Mexico, and adjacent portions of California, Colorado, Ne ...
(Arizona and New Mexico), the Midwest (Indiana, Nebraska, North and South Dakota, Ohio, Minnesota, and Wisconsin), New England
New England is a region comprising six states in the Northeastern United States: Connecticut, Maine, Massachusetts, New Hampshire, Rhode Island, and Vermont. It is bordered by the state of New York to the west and by the Canadian provinces ...
(Maine, New Hampshire, Vermont, and Massachusetts) and in New York and Pennsylvania.
There had been sporadic records of wolverines in Ukraine
Ukraine ( uk, Україна, Ukraïna, ) is a country in Eastern Europe. It is the second-largest European country after Russia, which it borders to the east and northeast. Ukraine covers approximately . Prior to the ongoing Russian inv ...
, but it is rather unclear whether the wolverines would have formed sustainable populations.
Unique records of encounters with wolverines have been noted in Latvia
Latvia ( or ; lv, Latvija ; ltg, Latveja; liv, Leţmō), officially the Republic of Latvia ( lv, Latvijas Republika, links=no, ltg, Latvejas Republika, links=no, liv, Leţmō Vabāmō, links=no), is a country in the Baltic region of ...
as well, the most recent one being in late July 2022 (although it can be disputed because of the unclear footage); the population was widespread around 16th—17th centuries, but nowadays it is not native to the area.
Conservation
The world's total wolverine population is not known. The animal exhibits a low population density and requires a very large home range.[Glenn Hurowitz (5 March 2008).]
First wolverine in 30 years spotted in California
", Grist.org
''Grist'' (originally ''Grist Magazine''; also referred to as Grist.org) is an American non-profit online magazine founded in 1999 that publishes environmental news and commentary. ''Grists tagline is "Climate. Justice. Solutions." ''Grist'' ...
; also The wolverine is listed by the IUCN as Least Concern because of its "wide distribution, remaining large populations, and the unlikelihood that it is in decline at a rate fast enough to trigger even Near Threatened".
The range of a male wolverine can be more than 620 km2 (240 mi2), encompassing the ranges of several females which have smaller home ranges of roughly 130–260 km2 (50–100 mi2). Adult wolverines try for the most part to keep nonoverlapping ranges with adults of the same sex. Radio tracking suggests an animal can range hundreds of miles in a few months.
Female wolverines burrow into snow in February to create a den, which is used until weaning in mid-May. Areas inhabited nonseasonally by wolverines are thus restricted to zones with late-spring snowmelt
In hydrology, snowmelt is surface runoff produced from melting snow. It can also be used to describe the period or season during which such runoff is produced. Water produced by snowmelt is an important part of the annual water cycle in many part ...
s. This fact has led to concern that global warming
In common usage, climate change describes global warming—the ongoing increase in global average temperature—and its effects on Earth's climate system. Climate change in a broader sense also includes previous long-term changes to E ...
will shrink the ranges of wolverine populations.
This requirement for large territories brings wolverines into conflict with human development, and hunting and trapping further reduce their numbers, causing them to disappear from large parts of their former range; attempts to have them declared an endangered species have met with little success. In February 2013, the United States Fish and Wildlife Service
The United States Fish and Wildlife Service (USFWS or FWS) is an agency within the United States Department of the Interior dedicated to the management of fish, wildlife, and natural habitats. The mission of the agency is "working with othe ...
proposed giving Endangered Species Act
The Endangered Species Act of 1973 (ESA or "The Act"; 16 U.S.C. § 1531 et seq.) is the primary law in the United States for protecting imperiled species. Designed to protect critically imperiled species from extinction as a "consequence of ec ...
protections to the wolverine due to its winter habitat
In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
in the northern Rockies diminishing. This was as a result of a lawsuit brought by the Center for Biological Diversity
The Center for Biological Diversity is a nonprofit membership organization known for its work protecting endangered species through legal action, scientific petitions, creative media and grassroots activism. It was founded in 1989 by Kieran Suckl ...
and Defenders of Wildlife
Defenders of Wildlife is a 501(c)(3) non-profit conservation organization based in the United States. It works to protect all native animals and plants throughout North America in their natural communities.
Background
Defenders of Wildlife is a n ...
.
The Wildlife Conservation Society
The Wildlife Conservation Society (WCS) is a non-governmental organization headquartered at the Bronx Zoo in New York City, that aims to conserve the world's largest wild places in 14 priority regions. Founded in 1895 as the New York Zoological ...
reported in June 2009 that a wolverine researchers had been tracking for almost three months had crossed into northern Colorado
Colorado (, other variants) is a state in the Mountain West subregion of the Western United States. It encompasses most of the Southern Rocky Mountains, as well as the northeastern portion of the Colorado Plateau and the western edge of t ...
. Society officials had tagged the young male wolverine in Wyoming
Wyoming () is a U.S. state, state in the Mountain states, Mountain West subregion of the Western United States. It is bordered by Montana to the north and northwest, South Dakota and Nebraska to the east, Idaho to the west, Utah to the south ...
near Grand Teton National Park
Grand Teton National Park is an American national park in northwestern Wyoming. At approximately , the park includes the major peaks of the Teton Range as well as most of the northern sections of the valley known as Jackson Hole. Grand Teton Na ...
, and it had traveled southward for about . It was the first wolverine seen in Colorado since 1919, and its appearance was also confirmed by the Colorado Division of Wildlife
Colorado Parks and Wildlife manages the state parks system and the wildlife of the U.S. state of Colorado. , the division managed the 42 state parks and 307 wildlife areas of Colorado.
, the Colorado Natural Areas Program had 93 designated s ...
. In May 2016 the same wolverine was killed by a cattle ranch-hand in North Dakota, ending a greater-than-800-mile (1287 km) trip by this lone male wolverine, dubbed M-56. This was the first verified sighting of a Wolverine in North Dakota in 150 years. In February 2014, a wolverine was seen in Utah, the first confirmed sighting in that state in 30 years.
In captivity
Around a hundred wolverines are held in zoos across North America and Europe, and they have been bred in captivity, but only with difficulty and high infant mortality.[
]
Name
The wolverine's questionable reputation as an insatiable glutton (reflected in the Latin genus name ''Gulo'') may be in part due to a false etymology
A false etymology (fake etymology, popular etymology, etymythology, pseudo-etymology, or par(a)etymology) is a popular but false belief about the origin or derivation of a specific word. It is sometimes called a folk etymology, but this is also a ...
. The less common name for the animal in Norwegian
Norwegian, Norwayan, or Norsk may refer to:
*Something of, from, or related to Norway, a country in northwestern Europe
*Norwegians, both a nation and an ethnic group native to Norway
*Demographics of Norway
*The Norwegian language, including the ...
, ''fjellfross'', meaning "mountain cat", is thought to have worked its way into German as ''Vielfraß'', which means "glutton" (literally "devours much"). Its name in other West Germanic languages is similar (e.g. nl, veelvraat).
The Finnish name is ''ahma'', derived from ''ahmatti,'' which is translated as "glutton". Similarly, the Estonian
Estonian may refer to:
* Something of, from, or related to Estonia, a country in the Baltic region in northern Europe
* Estonians, people from Estonia, or of Estonian descent
* Estonian language
* Estonian cuisine
* Estonian culture
See also ...
name is ''ahm'', with the equivalent meaning to the Finnish name. In Lithuanian, it is ''ernis''; in Latvian, ''tinis'' or ''āmrija''.
The Eastern Slavic росомаха (''rosomakha'') and the Polish and Czech
Czech may refer to:
* Anything from or related to the Czech Republic, a country in Europe
** Czech language
** Czechs, the people of the area
** Czech culture
** Czech cuisine
* One of three mythical brothers, Lech, Czech, and Rus'
Places
*Czech, ...
name ''rosomák'' seem to be borrowed from the Finnish ''rasva-maha'' (fat belly). Similarly, the Hungarian name is ''rozsomák'' or ''torkosborz'' which means "gluttonous badger".
In French-speaking parts of Canada, the wolverine is referred to as ''carcajou'', borrowed from the Innu-aimun
Innu-aimun or Montagnais is an Algonquian language spoken by over 10,000 Innu in Labrador and Quebec in Eastern Canada. It is a member of the Cree–Montagnais–Naskapi dialect continuum and is spoken in various dialects depending on the comm ...
or Montagnais ''kuàkuàtsheu''. However, in France, the wolverine's name is ''glouton'' (glutton).
Purported gluttony is reflected neither in the English name ''wolverine'' nor in the names used in North Germanic languages
The North Germanic languages make up one of the three branches of the Germanic languages—a sub-family of the Indo-European languages—along with the West Germanic languages and the extinct East Germanic languages. The language group is also r ...
. The English word wolverine (alteration of the earlier form, wolvering, of uncertain origin) probably implies "a little wolf". The name in Proto-Norse
Proto-Norse (also called Ancient Nordic, Ancient Scandinavian, Ancient Norse, Primitive Norse, Proto-Nordic, Proto-Scandinavian and Proto-North Germanic) was an Indo-European language spoken in Scandinavia that is thought to have evolved as a ...
, '' erafaz'' and Old Norse
Old Norse, Old Nordic, or Old Scandinavian, is a stage of development of North Germanic languages, North Germanic dialects before their final divergence into separate Nordic languages. Old Norse was spoken by inhabitants of Scandinavia and t ...
, ''jarfr'', lives on in the regular Icelandic name ''jarfi'', regular Norwegian name ''jerv'', regular Swedish name ''järv'' and regular Danish name ''jærv''.
In culture
Many North American cities, teams, and organizations use the wolverine as a mascot. For example, the US state of Michigan
Michigan () is a state in the Great Lakes region of the upper Midwestern United States. With a population of nearly 10.12 million and an area of nearly , Michigan is the 10th-largest state by population, the 11th-largest by area, and the ...
is, by tradition, known as "the Wolverine State", and the University of Michigan
, mottoeng = "Arts, Knowledge, Truth"
, former_names = Catholepistemiad, or University of Michigania (1817–1821)
, budget = $10.3 billion (2021)
, endowment = $17 billion (2021)As o ...
takes the animal as its mascot. There have also been professional baseball
Baseball is a bat-and-ball sport played between two teams of nine players each, taking turns batting and fielding. The game occurs over the course of several plays, with each play generally beginning when a player on the fielding tea ...
and football
Football is a family of team sports that involve, to varying degrees, kicking a ball to score a goal. Unqualified, the word ''football'' normally means the form of football that is the most popular where the word is used. Sports commonly c ...
clubs called the "Wolverines". The association is well and long established: for example, many Detroiters volunteered to fight during the American Civil War
The American Civil War (April 12, 1861 – May 26, 1865; also known by other names) was a civil war in the United States. It was fought between the Union ("the North") and the Confederacy ("the South"), the latter formed by states th ...
and George Armstrong Custer
George Armstrong Custer (December 5, 1839 – June 25, 1876) was a United States Army officer and cavalry commander in the American Civil War and the American Indian Wars.
Custer graduated from West Point in 1861 at the bottom of his class, b ...
, who led the Michigan Brigade The Michigan Brigade, sometimes called the Wolverines, the Michigan Cavalry Brigade or Custer's Brigade, was a brigade of cavalry in the volunteer Union Army during the latter half of the American Civil War. Composed primarily of the 1st Michigan C ...
, called them the "Wolverines". The origins of this association are obscure; it may derive from a busy trade in wolverine furs in Sault Ste. Marie in the 18th century or may recall a disparagement intended to compare early settlers in Michigan with the vicious mammal. Wolverines are, however, extremely rare in Michigan. A sighting in February 2004 near Ubly
Ubly is a village in Huron County, Michigan, Huron County in the U.S. state of Michigan. The population was 858 at the 2010 United States Census, 2010 census. The village is within Bingham Township, Huron County, Michigan, Bingham Township.
On Fe ...
was the first confirmed sighting in Michigan in 200 years. The animal was found dead in 2010.
Marvel Comics
Marvel Comics is an American comic book publishing, publisher and the flagship property of Marvel Entertainment, a divsion of The Walt Disney Company since September 1, 2009. Evolving from Timely Comics in 1939, ''Magazine Management/Atlas Co ...
character James "Logan" Howlett was given the name "Wolverine" because of his short stature, keen animal senses, and ferocity.
The wolverine is prevalent in stories and oral history from various Algonquian tribes and figures prominently in the mythology of the Innu
The Innu / Ilnu ("man", "person") or Innut / Innuat / Ilnuatsh ("people"), formerly called Montagnais from the French colonial period ( French for "mountain people", English pronunciation: ), are the Indigenous inhabitants of territory in the ...
people of eastern Quebec
Quebec ( ; )According to the Canadian government, ''Québec'' (with the acute accent) is the official name in Canadian French and ''Quebec'' (without the accent) is the province's official name in Canadian English is one of the thirtee ...
and Labrador
, nickname = "The Big Land"
, etymology =
, subdivision_type = Country
, subdivision_name = Canada
, subdivision_type1 = Province
, subdivision_name1 ...
. The wolverine is known as Kuekuatsheu, a conniving trickster who created the world. The story of the formation of the Innu world begins long ago when Kuekuatsheu built a big boat similar to Noah's Ark
Noah's Ark ( he, תיבת נח; Biblical Hebrew: ''Tevat Noaḥ'')The word "ark" in modern English comes from Old English ''aerca'', meaning a chest or box. (See Cresswell 2010, p.22) The Hebrew word for the vessel, ''teva'', occurs twice in t ...
and put all the various animal species in it. There was a great deal of rain, and the land was flooded. Kuekuatsheu told a mink
Mink are dark-colored, semiaquatic, carnivorous mammals of the genera ''Neogale'' and '' Mustela'' and part of the family Mustelidae, which also includes weasels, otters, and ferrets. There are two extant species referred to as "mink": the A ...
to dive into the water to retrieve some mud and rocks which he mixed together to create an island, which is the world that is presently inhabited along with all the animals. Many tales of Kuekuatsheu are often humorous and irreverent and include crude references to bodily functions. Some Northeastern tribes, such as the Miꞌkmaq
The Mi'kmaq (also ''Mi'gmaq'', ''Lnu'', ''Miꞌkmaw'' or ''Miꞌgmaw''; ; ) are a First Nations people of the Northeastern Woodlands, indigenous to the areas of Canada's Atlantic Provinces and the Gaspé Peninsula of Quebec as well as the nort ...
and Passamaquoddy
The Passamaquoddy ( Maliseet-Passamaquoddy: ''Peskotomuhkati'') are a Native American/First Nations people who live in northeastern North America. Their traditional homeland, Peskotomuhkatik'','' straddles the Canadian province of New Brunswick ...
, refer to the wolverine as Lox, who usually appears in tales as a trickster and thief (although generally more dangerous than its Innu counterpart) and is often depicted as a companion to the wolf
The wolf (''Canis lupus''; : wolves), also known as the gray wolf or grey wolf, is a large canine native to Eurasia and North America. More than thirty subspecies of ''Canis lupus'' have been recognized, and gray wolves, as popularly u ...
. Similarly, the Dené, a group of the Athabaskan
Athabaskan (also spelled ''Athabascan'', ''Athapaskan'' or ''Athapascan'', and also known as Dene) is a large family of indigenous languages of North America, located in western North America in three areal language groups: Northern, Pacific C ...
-speaking natives of northwestern Canada, have many stories of the wolverine as a trickster and cultural transformer much like the coyote
The coyote (''Canis latrans'') is a species of canis, canine native to North America. It is smaller than its close relative, the wolf, and slightly smaller than the closely related eastern wolf and red wolf. It fills much of the same ecologica ...
in the Navajo
The Navajo (; British English: Navaho; nv, Diné or ') are a Native American people of the Southwestern United States.
With more than 399,494 enrolled tribal members , the Navajo Nation is the largest federally recognized tribe in the United ...
tradition or raven
A raven is any of several larger-bodied bird species of the genus ''Corvus''. These species do not form a single taxonomic group within the genus. There is no consistent distinction between "crows" and "ravens", common names which are assigned t ...
in Northwest Coast traditions.[Moore, Patrick and Angela Wheelock. (1990). ''Wolverine Myths and Visions: Dene Traditions from Northern Alberta''. University of Nebraska Press. ]
Gallery
Brehms Het Leven der Dieren Zoogdieren Orde 4 Veelvraat (Gulo borealis).jpg
Wolverine display at Arctic Interagency Visitor Center at Coldfoot.jpg
Gulo gulo 01.jpg
Wolverin pup in Sweden.jpg
Wolverine in Sweden.jpg
References
External links
Large Carnivore Initiative for Europe: Wolverine
scientific articles about wolverines