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''Wisteria'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...
s in the legume family, Fabaceae (Leguminosae), that includes ten species of woody twining vines that are native to China, Japan,
Korea Korea ( ko, 한국, or , ) is a peninsular region in East Asia. Since 1945, it has been divided at or near the 38th parallel, with North Korea (Democratic People's Republic of Korea) comprising its northern half and South Korea (Republic o ...
,
Vietnam Vietnam or Viet Nam ( vi, Việt Nam, ), officially the Socialist Republic of Vietnam,., group="n" is a country in Southeast Asia, at the eastern edge of mainland Southeast Asia, with an area of and population of 96 million, making i ...
,
Southern Canada The list of regions of Canada is a summary of geographical areas on a hierarchy that ranges from national (groups of provinces and territories) at the top to local regions and sub-regions of provinces at the bottom. Administrative regions that ran ...
, the
Eastern United States The Eastern United States, commonly referred to as the American East, Eastern America, or simply the East, is the region of the United States to the east of the Mississippi River. In some cases the term may refer to a smaller area or the East C ...
, and north of
Iran Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...
. They were later introduced to France, Germany and various other countries in Europe. Some species are popular ornamental plants. The aquatic flowering plant commonly called wisteria or 'water wisteria' is in fact '' Hygrophila difformis'', in the family
Acanthaceae Acanthaceae is a family (the acanthus family) of dicotyledonous flowering plants containing almost 250 genera and about 2500 species. Most are tropical herbs, shrubs, or twining vines; some are epiphytes. Only a few species are distributed in tem ...
.


Etymology

The botanist
Thomas Nuttall Thomas Nuttall (5 January 1786 – 10 September 1859) was an English botanist and zoologist who lived and worked in America from 1808 until 1841. Nuttall was born in the village of Long Preston, near Settle in the West Riding of Yorkshire and ...
said he named the genus ''Wisteria'' in memory of the American physician and anatomist Caspar Wistar (1761–1818). Both men were living in Philadelphia at the time, where Wistar was a professor in the
School of Medicine A medical school is a tertiary educational institution, or part of such an institution, that teaches medicine, and awards a professional degree for physicians. Such medical degrees include the Bachelor of Medicine, Bachelor of Surgery (MBBS, M ...
at the
University of Pennsylvania The University of Pennsylvania (also known as Penn or UPenn) is a private research university in Philadelphia. It is the fourth-oldest institution of higher education in the United States and is ranked among the highest-regarded universitie ...
. Questioned about the spelling later, Nuttall said it was for "
euphony Phonaesthetics (also spelled phonesthetics in North America) is the study of beauty and pleasantness associated with the sounds of certain words or parts of words. The term was first used in this sense, perhaps by during the mid-20th century and ...
", but his biographer speculated that it may have something to do with Nuttall's friend Charles Jones Wister Sr., of
Grumblethorpe Grumblethorpe, in Germantown, Philadelphia, Pennsylvania, was the home of the Wister family, who lived there for over 160 years. It was built in 1744 as a summer residence, but it became the family's year-round residence in 1793. It is a museum ...
, the grandson of the merchant John Wister. Various sources assert that the naming occurred in
Philadelphia Philadelphia, often called Philly, is the List of municipalities in Pennsylvania#Municipalities, largest city in the Commonwealth (U.S. state), Commonwealth of Pennsylvania, the List of United States cities by population, sixth-largest city i ...
. Another source claims that the person who named ''Wisteria'' after Caspar Wistar was the Portuguese botanist and geologist José Francisco Correia da Serra, who lived in Philadelphia beginning in 1812, four years before his appointment as ambassador of
Portugal Portugal, officially the Portuguese Republic ( pt, República Portuguesa, links=yes ), is a country whose mainland is located on the Iberian Peninsula of Southwestern Europe, and whose territory also includes the Atlantic archipelagos of ...
to the United States. Correia became a close friend of Wistar, "took tea at his home daily, and named the vine 'Wisteria' to commemorate this friendship." As the spelling is apparently deliberate, there is no justification for changing the genus name under the
International Code of Botanical Nomenclature The ''International Code of Nomenclature for algae, fungi, and plants'' (ICN) is the set of rules and recommendations dealing with the formal botanical names that are given to plants, fungi and a few other groups of organisms, all those "trad ...
. However, some spell the plant's common name "wistaria". In some countries in Western and Central Europe, Wisteria is also known by a variant spelling of the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
''
Glycine Glycine (symbol Gly or G; ) is an amino acid that has a single hydrogen atom as its side chain. It is the simplest stable amino acid ( carbamic acid is unstable), with the chemical formula NH2‐ CH2‐ COOH. Glycine is one of the proteinog ...
'', of which some flowers also have lavender-colored petals. Examples include the French ''Glycines'', the German ''Glyzinie'', and the Polish ''Glicynia''.


Taxonomy

Genetic analysis shows ''
Callerya ''Callerya'' is a genus of flowering plants in the legume family, Fabaceae. It belongs to the subfamily Faboideae. Genetic analysis shows ''Callerya'' and ''Wisteria'' to be each other's closest relatives and quite distinct from other members of ...
'', '' Afgekia'' and ''Wisteria'' to be each other's closest relatives and quite distinct from other members of the tribe Millettieae. Both have eight chromosomes.


Species

In East Asia and North America there are about nine species of wisteria. In North America, '' W. floribunda'' (Japanese wisteria) and '' W. sinensis'' (Chinese wisteria) are far more popular than other species for their abundance of flowers, clusters of large flowers, variety of flower colors, and fragrance. ''W. sinensis'' was brought to the United States for horticultural purposes in 1816, while ''W. floribunda'' was introduced around 1830. Because of their hardiness and tendency to escape cultivation, these non-native wisterias are considered invasive species in many parts of the
U.S. The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territori ...
, especially the
Southeast The points of the compass are a set of horizontal, radially arrayed compass directions (or azimuths) used in navigation and cartography. A compass rose is primarily composed of four cardinal directions—north, east, south, and west—each sepa ...
, due to their ability to overtake and choke out other native plant species. The following is a list of accepted ''Wisteria'' species:


Description

Wisterias climb by twining their stems around any available support. '' W. floribunda'' (Japanese wisteria) twines clockwise when viewed from above, while '' W. sinensis'' (Chinese wisteria) twines counterclockwise. This is an aid in identifying the two most common species of wisteria. They can climb as high as above the ground and spread out laterally. The world's largest known wisteria is in Sierra Madre, California, measuring more than in size and weighing 250 tons. Planted in 1894, it is of the 'Chinese lavender' variety. The leaves are alternate, 15 to 35 cm long,
pinnate Pinnation (also called pennation) is the arrangement of feather-like or multi-divided features arising from both sides of a common axis. Pinnation occurs in biological morphology, in crystals, such as some forms of ice or metal crystals, and in ...
, with 9 to 19 leaflets. The
flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechani ...
s have drooping racemes that vary in length from species to species. '' W. frutescens'' (American wisteria) has the shortest racemes, . ''W. floribunda'' (Japanese wisteria) has the longest racemes, in some
varieties Variety may refer to: Arts and entertainment Entertainment formats * Variety (radio) * Variety show, in theater and television Films * ''Variety'' (1925 film), a German silent film directed by Ewald Andre Dupont * ''Variety'' (1935 film), ...
and or in some cultivars. The flowers come in a variety of colors, including white, lilac, purple, and pink, and some '' W. brachybotrys'' (Silky wisteria) and ''W. floribunda'' cultivars have particularly remarkable colors. The flowers are fragrant, and especially cultivars of ''W. brachybotrys'', ''W. floribunda'', and ''W. sinensis'' are noted for their sweet and musky scents. Flowering is in spring (just before or as the leaves open) in some Asian species, and in mid to late summer in the American species. ''Wisteria'' species are used as food plants by the
larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. ...
e of some Lepidoptera species including
brown-tail moth The brown-tail moth (''Euproctis chrysorrhoea'') is a moth of the family Erebidae. It is native to Europe, neighboring countries in Asia, and the north coast of Africa. Descriptions of outbreaks, i.e., large population increases of several year ...
. The
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiospe ...
s are produced in pods similar to those of ''Laburnum'', and, like the seeds of that genus, are poisonous. All parts of the plant contain a saponin called wisterin, which is
toxic Toxicity is the degree to which a chemical substance or a particular mixture of substances can damage an organism. Toxicity can refer to the effect on a whole organism, such as an animal, bacterium, or plant, as well as the effect on a subs ...
if ingested, and may cause
dizziness Dizziness is an imprecise term that can refer to a sense of disorientation in space, vertigo, or lightheadedness. It can also refer to disequilibrium or a non-specific feeling, such as giddiness or foolishness. Dizziness is a common medical c ...
, confusion, speech problems, nausea,
vomiting Vomiting (also known as emesis and throwing up) is the involuntary, forceful expulsion of the contents of one's stomach through the mouth and sometimes the nose. Vomiting can be the result of ailments like food poisoning, gastroenteri ...
, stomach pains,
diarrhea Diarrhea, also spelled diarrhoea, is the condition of having at least three loose, liquid, or watery bowel movements each day. It often lasts for a few days and can result in dehydration due to fluid loss. Signs of dehydration often begin w ...
and collapse. There is debate over whether the concentration outside of the seeds is sufficient to cause poisoning. Wisteria seeds have caused poisoning in children and pets of many countries, producing mild to severe gastroenteritis and other effects.


Cultivation

'' W. floribunda'' (Japanese wisteria), which has the longest racemes of wisteria species, is decorative and has given rise to many
cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture ...
s that have won the prestigious Award of Garden Merit. Wisteria, especially '' W. sinensis'' (Chinese wisteria), is very hardy and fast-growing. It can grow in fairly poor-quality soils, but prefers fertile, moist, well-drained soil. It thrives in full sun. It can be propagated via hardwood cutting, softwood cuttings, or seed. However, specimens grown from seed can take decades to bloom; for this reason, gardeners usually grow plants that have been started from rooted cuttings or grafted
cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture ...
s known to flower well. Another reason for failure to bloom can be excessive fertilizer (particularly
nitrogen Nitrogen is the chemical element with the symbol N and atomic number 7. Nitrogen is a nonmetal and the lightest member of group 15 of the periodic table, often called the pnictogens. It is a common element in the universe, estimated at se ...
). Wisteria has nitrogen fixing capability (provided by Rhizobia bacteria in root nodules), and thus mature plants may benefit from added
potassium Potassium is the chemical element with the symbol K (from Neo-Latin ''kalium'') and atomic number19. Potassium is a silvery-white metal that is soft enough to be cut with a knife with little force. Potassium metal reacts rapidly with atmosph ...
and
phosphate In chemistry, a phosphate is an anion, salt, functional group or ester derived from a phosphoric acid. It most commonly means orthophosphate, a derivative of orthophosphoric acid . The phosphate or orthophosphate ion is derived from phosph ...
, but not nitrogen. Finally, wisteria can be reluctant to bloom before it has reached maturity. Maturation may require only a few years, as in '' W. macrostachya'' (Kentucky wisteria), or nearly twenty, as in ''W. sinensis''. Maturation can be forced by physically abusing the main trunk, root pruning, or
drought A drought is defined as drier than normal conditions.Douville, H., K. Raghavan, J. Renwick, R.P. Allan, P.A. Arias, M. Barlow, R. Cerezo-Mota, A. Cherchi, T.Y. Gan, J. Gergis, D.  Jiang, A.  Khan, W.  Pokam Mba, D.  Rosenfeld, J. Tierney, an ...
stress. Wisteria can grow into a mound when unsupported, but is at its best when allowed to clamber up a tree, pergola, wall, or other supporting structure. If you want wisteria to grow along the pergola, ''W. floribunda'' (Japanese wisteria) with longer racemes is the best choice. If you want wisteria to grow along the wall, ''W. sinensis'' (Chinese wisteria) with shorter racemes is the best choice. Whatever the case, the support must be very sturdy, because mature wisteria can become immensely strong with heavy wrist-thick trunks and stems. These can collapse latticework, crush thin wooden posts, and even strangle large trees. Wisteria allowed to grow on houses can cause damage to gutters, downspouts, and similar structures. Wisteria flowers develop in buds near the base of the previous year's growth, so
pruning Pruning is a horticultural, arboricultural, and silvicultural practice involving the selective removal of certain parts of a plant, such as branches, buds, or roots. The practice entails the ''targeted'' removal of diseased, damaged, dead, ...
back side shoots to the basal few buds in early spring can enhance the visibility of the flowers. If it is desired to control the size of the plant, the side shoots can be shortened to between 20 and 40 cm long in midsummer, and back to in the fall. Once the plant is a few years old, a relatively compact, free-flowering form can be achieved by pruning off the new tendrils three times during the growing season in the summer months. The flowers of some varieties are edible, and can even be used to make wine. Others are said to be toxic. Careful identification by an expert is strongly recommended before consuming this or any wild plant. In the United Kingdom, the national collection of wisteria is held by Chris Lane at the Witch Hazel Nursery in Newington, near
Sittingbourne Sittingbourne is an industrial town in Kent, south-east England, from Canterbury and from London, beside the Roman Watling Street, an ancient British trackway used by the Romans and the Anglo-Saxons and next to the Swale, a strip of sea separa ...
in
Kent Kent is a county in South East England and one of the home counties. It borders Greater London to the north-west, Surrey to the west and East Sussex to the south-west, and Essex to the north across the estuary of the River Thames; it faces ...
.


Art and symbolism

Wisteria and their racemes have been widely used in Japan throughout the centuries and were a popular symbol in family crests and heraldry. One popular dance in
kabuki is a classical form of Japanese dance- drama. Kabuki theatre is known for its heavily-stylised performances, the often-glamorous costumes worn by performers, and for the elaborate make-up worn by some of its performers. Kabuki is though ...
known as the '' Fuji Musume--'' or "''The Wisteria Maiden''"-- is the sole extant dance of a series of five personifying dances in which a maiden becomes the embodiment of the spirit of wisteria. In the West, both in building materials such as tile, as well as stained glass, wisterias have been used both in realism and stylistically in artistic works and industrial design.


References


External links


Ohio State University fact sheet on wisteria
{{Taxonbar, from=Q261903 Fabaceae genera Vines Poisonous plants