ecosystem
An ecosystem (or ecological system) consists of all the organisms and the physical environment with which they interact. These biotic and abiotic components are linked together through nutrient cycles and energy flows. Energy enters the syste ...
water
Water (chemical formula ) is an inorganic, transparent, tasteless, odorless, and nearly colorless chemical substance, which is the main constituent of Earth's hydrosphere and the fluids of all known living organisms (in which it acts as a ...
, either permanently (for years or decades) or seasonally (for weeks or months). Flooding results in oxygen-free (
anoxic
The term anoxia means a total depletion in the level of oxygen, an extreme form of hypoxia or "low oxygen". The terms anoxia and hypoxia are used in various contexts:
* Anoxic waters, sea water, fresh water or groundwater that are depleted of diss ...
) processes prevailing, especially in the soils. /ref> The primary factor that distinguishes wetlands from terrestrial land forms or Body of water, water bodies is the characteristic
vegetation
Vegetation is an assemblage of plant species and the ground cover they provide. It is a general term, without specific reference to particular taxa, life forms, structure, spatial extent, or any other specific botanical or geographic character ...
of
aquatic plants
Aquatic plants are plants that have adapted to living in aquatic environments (saltwater or freshwater). They are also referred to as hydrophytes or macrophytes to distinguish them from algae and other microphytes. A macrophyte is a plant that ...
, adapted to the unique anoxic
hydric soil Hydric soil is soil which is permanently or seasonally saturated by water, resulting in anaerobic conditions, as found in wetlands.
Overview
Most soils are aerobic. This is important because plant roots respire (that is, they consume oxygen and ...
s. Wetlands are considered among the most biologically diverse of all ecosystems, serving as home to a wide range of plant and animal species. Methods for assessing wetland functions, wetland
ecological health
Ecological health is a term that has been used in relation to both human health and the condition of the environment.
*In medicine, ecological health has been used to refer to multiple chemical sensitivity, which results from exposure to synthet ...
, and general wetland condition have been developed for many regions of the world. These methods have contributed to
wetland conservation
Wetland conservation is aimed at protecting and preserving areas where water exists at or near the Earth's surface, such as swamps, marshes and bogs. Wetlands cover at least six per cent of the Earth and have become a focal issue for conservation ...
partly by raising public awareness of the functions some wetlands provide.
Wetlands occur naturally on every
continent
A continent is any of several large landmasses. Generally identified by convention rather than any strict criteria, up to seven geographical regions are commonly regarded as continents. Ordered from largest in area to smallest, these seven ...
. The water in wetlands is either
freshwater
Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does include ...
,
brackish
Brackish water, sometimes termed brack water, is water occurring in a natural environment that has more salinity than freshwater, but not as much as seawater. It may result from mixing seawater (salt water) and fresh water together, as in estuari ...
or
saltwater
Saline water (more commonly known as salt water) is water that contains a high concentration of dissolved salts (mainly sodium chloride). On the United States Geological Survey (USGS) salinity scale, saline water is saltier than brackish water, ...
. The main wetland types are classified based on the dominant
plant
Plants are predominantly photosynthetic eukaryotes of the kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all current definitions of Plantae exclud ...
s and/or the source of the water. For example,
marsh
A marsh is a wetland that is dominated by herbaceous rather than woody plant species.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p Marshes can often be found at ...
reed
Reed or Reeds may refer to:
Science, technology, biology, and medicine
* Reed bird (disambiguation)
* Reed pen, writing implement in use since ancient times
* Reed (plant), one of several tall, grass-like wetland plants of the order Poales
* ...
s,
cattail
''Typha'' is a genus of about 30 species of monocotyledonous flowering plants in the family Typhaceae. These plants have a variety of common names, in British English as bulrush or reedmace, in American English as reed, cattail, or punks, in A ...
s and
sedge
The Cyperaceae are a family of graminoid (grass-like), monocotyledonous flowering plants known as sedges. The family is large, with some 5,500 known species described in about 90 genera, the largest being the "true sedges" genus ''Carex'' wit ...
s;
swamp
A swamp is a forested wetland.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p. Swamps are considered to be transition zones because both land and water play a role in ...
s are ones dominated by
woody vegetation
A woody plant is a plant that produces wood as its structural tissue and thus has a hard stem. In cold climates, woody plants further survive winter or dry season above ground, as opposite to herbaceous plants that die back to the ground until sp ...
such as
tree
In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
s and
shrub
A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
s (although reed swamps in Europe are dominated by reeds, not trees). Examples of wetlands classified by their sources of water include
tidal wetland
Tidal is the adjectival form of tide.
Tidal may also refer to:
* ''Tidal'' (album), a 1996 album by Fiona Apple
* Tidal (king), a king involved in the Battle of the Vale of Siddim
* TidalCycles, a live coding environment for music
* Tidal (servic ...
s (
ocean
The ocean (also the sea or the world ocean) is the body of salt water that covers approximately 70.8% of the surface of Earth and contains 97% of Earth's water. An ocean can also refer to any of the large bodies of water into which the wo ...
ic
tide
Tides are the rise and fall of sea levels caused by the combined effects of the gravity, gravitational forces exerted by the Moon (and to a much lesser extent, the Sun) and are also caused by the Earth and Moon orbiting one another.
Tide t ...
s),
estuaries
An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environment ...
(mixed tidal and river waters),
floodplain
A floodplain or flood plain or bottomlands is an area of land adjacent to a river which stretches from the banks of its channel to the base of the enclosing valley walls, and which experiences flooding during periods of high discharge.Goudi ...
s (excess water from overflowed rivers or lakes),
spring
Spring(s) may refer to:
Common uses
* Spring (season), a season of the year
* Spring (device), a mechanical device that stores energy
* Spring (hydrology), a natural source of water
* Spring (mathematics), a geometric surface in the shape of a ...
s,
seep
A seep or flush is a moist or wet place where water, usually groundwater, reaches the earth's surface from an underground aquifer.
Description
Seeps are usually not of sufficient volume to be flowing beyond their immediate above-ground location. ...
s and
fen
A fen is a type of peat-accumulating wetland fed by mineral-rich ground or surface water. It is one of the main types of wetlands along with marshes, swamps, and bogs. Bogs and fens, both peat-forming ecosystems, are also known as mires. T ...
s (
groundwater
Groundwater is the water present beneath Earth's surface in rock and soil pore spaces and in the fractures of rock formations. About 30 percent of all readily available freshwater in the world is groundwater. A unit of rock or an unconsolidate ...
vernal pond
Vernal pools, also called vernal ponds or ephemeral pools, are seasonal pools of water that provide habitat for distinctive plants and animals. They are considered to be a distinctive type of wetland usually devoid of fish, and thus allow the safe ...
s (
rainfall
Rain is water droplets that have condensed from atmospheric water vapor and then fall under gravity. Rain is a major component of the water cycle and is responsible for depositing most of the fresh water on the Earth. It provides water f ...
or
meltwater
Meltwater is water released by the melting of snow or ice, including glacial ice, tabular icebergs and ice shelves over oceans. Meltwater is often found in the ablation zone of glaciers, where the rate of snow cover is reducing. Meltwater can be ...
). Some wetlands have multiple types of plants and are fed by multiple sources of water, making them difficult to classify. The world's largest wetlands include the
Amazon River basin
The Amazon basin is the part of South America drained by the Amazon River and its tributaries. The Amazon drainage basin covers an area of about , or about 35.5 percent of the South American continent. It is located in the countries of Bolivi ...
, the
West Siberian Plain
The West Siberian Plain (russian: За́падно-Сиби́рская равни́на ''Zapadno-Sibirskaya ravnina'') is a large plain that occupies the western portion of Siberia, between the Ural Mountains in the west and the Yenisei River ...
, the
Pantanal
The Pantanal () is a natural region encompassing the world's largest tropical wetland area, and the world's largest flooded grasslands. It is located mostly within the Brazilian state of Mato Grosso do Sul, but it extends into Mato Grosso and p ...
in South America, and the
Sundarbans
Sundarbans (pronounced ) is a mangrove area in the delta formed by the confluence of the Padma, Brahmaputra and Meghna Rivers in the Bay of Bengal. It spans the area from the Baleswar River in Bangladesh's division of Khulna to the Hooghly R ...
in the
Ganges
The Ganges ( ) (in India: Ganga ( ); in Bangladesh: Padma ( )). "The Ganges Basin, known in India as the Ganga and in Bangladesh as the Padma, is an international river to which India, Bangladesh, Nepal and China are the riparian states." is ...
-
Brahmaputra
The Brahmaputra is a trans-boundary river which flows through Tibet, northeast India, and Bangladesh. It is also known as the Yarlung Tsangpo in Tibetan, the Siang/Dihang River in Arunachali, Luit in Assamese, and Jamuna River in Bangla. It ...
delta.
Wetlands contribute a number of functions that benefit people. These are called
ecosystem service
Ecosystem services are the many and varied benefits to humans provided by the natural environment and healthy ecosystems. Such ecosystems include, for example, agroecosystems, forest ecosystem, grassland ecosystems, and aquatic ecosystems. Th ...
s and include
water purification
Water purification is the process of removing undesirable chemicals, biological contaminants, suspended solids, and gases from water. The goal is to produce water that is fit for specific purposes. Most water is purified and disinfected for hu ...
,
groundwater replenishment
Groundwater recharge or deep drainage or deep percolation is a hydrologic process, where water moves downward from surface water to groundwater. Recharge is the primary method through which water enters an aquifer. This process usually occurs in ...
, stabilization of shorelines and storm protection, water storage and
flood control
Flood control methods are used to reduce or prevent the detrimental effects of flood waters."Flood Control", MSN Encarta, 2008 (see below: Further reading). Flood relief methods are used to reduce the effects of flood waters or high water level ...
, processing of carbon (
carbon fixation
Biological carbon fixation or сarbon assimilation is the process by which inorganic carbon (particularly in the form of carbon dioxide) is converted to organic compounds by living organisms. The compounds are then used to store energy and as ...
,
decomposition
Decomposition or rot is the process by which dead organic substances are broken down into simpler organic or inorganic matter such as carbon dioxide, water, simple sugars and mineral salts. The process is a part of the nutrient cycle and is e ...
pollutants
A pollutant or novel entity is a substance or energy introduced into the environment that has undesired effects, or adversely affects the usefulness of a resource. These can be both naturally forming (i.e. minerals or extracted compounds like oi ...
, and support of plants and animals. Wetlands are reservoirs of
biodiversity
Biodiversity or biological diversity is the variety and variability of life on Earth. Biodiversity is a measure of variation at the genetic (''genetic variability''), species (''species diversity''), and ecosystem (''ecosystem diversity'') l ...
and provide wetland products. According to the UN
Millennium Ecosystem Assessment
The Millennium Ecosystem Assessment (MA) is a major assessment of the human impact on the environment, called for by the United Nations Secretary-General Kofi Annan in 2000, launched in 2001 and published in 2005 with more than $14 million of gr ...
, wetlands are more affected by
environmental degradation
Environmental degradation is the deterioration of the environment (biophysical), environment through depletion of resources such as quality of air, water and soil; the destruction of ecosystems; habitat destruction; the extinction of wildlife; an ...
than any other ecosystem on Earth. Wetlands can be important sources and sinks of carbon, depending on the specific wetland, and thus will play an important role in
climate change
In common usage, climate change describes global warming—the ongoing increase in global average temperature—and its effects on Earth's climate system. Climate change in a broader sense also includes previous long-term changes to E ...
nitrous oxide
Nitrous oxide (dinitrogen oxide or dinitrogen monoxide), commonly known as laughing gas, nitrous, or nos, is a chemical compound, an oxide of nitrogen with the formula . At room temperature, it is a colourless non-flammable gas, and has a ...
.
Constructed wetland
A constructed wetland is an artificial wetland to treat sewage, greywater, stormwater runoff or industrial wastewater. It may also be designed for land reclamation after mining, or as a mitigation step for natural areas lost to land development. ...
s are designed and built to treat municipal and industrial
wastewater
Wastewater is water generated after the use of freshwater, raw water, drinking water or saline water in a variety of deliberate applications or processes. Another definition of wastewater is "Used water from any combination of domestic, industr ...
as well as to divert
stormwater
Stormwater, also spelled storm water, is water that originates from precipitation (storm), including heavy rain and meltwater from hail and snow. Stormwater can soak into the soil ( infiltrate) and become groundwater, be stored on depressed la ...
runoff. Constructed wetlands may also play a role in
water-sensitive urban design
Water-sensitive urban design (WSUD) is a land planning and engineering design approach which integrates the urban water cycle, including stormwater, groundwater, and wastewater management and water supply, into urban design to minimise environme ...
.
Definitions and terminology
Technical definitions
A simplified definition of wetland is "an area of land that is usually saturated with water". More precisely, wetlands are areas where "water covers the soil, or is present either at or near the surface of the soil all year or for varying periods of time during the year, including during the growing season". A patch of land that develops pools of water after a
rain storm
Rain is water droplets that have condensed from atmospheric water vapor and then fall under gravity. Rain is a major component of the water cycle and is responsible for depositing most of the fresh water on the Earth. It provides water fo ...
would not necessarily be considered a "wetland", even though the land is wet. Wetlands have unique characteristics: they are generally distinguished from other
water bodies
A body of water or waterbody (often spelled water body) is any significant accumulation of water on the surface of Earth or another planet. The term most often refers to oceans, seas, and lakes, but it includes smaller pools of water such as p ...
or
landform
A landform is a natural or anthropogenic land feature on the solid surface of the Earth or other planetary body. Landforms together make up a given terrain, and their arrangement in the landscape is known as topography. Landforms include hills, ...
s based on their
water level
Water level, also known as gauge height or stage, is the elevation of the free surface of a sea, stream, lake or reservoir relative to a specified vertical datum.ISO 772: 1996. Hydrometric determinations – Vocabulary and symbols.
See also
* ...
and on the types of
plant
Plants are predominantly photosynthetic eukaryotes of the kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all current definitions of Plantae exclud ...
s that live within them. Specifically, wetlands are characterized as having a
water table
The water table is the upper surface of the zone of saturation. The zone of saturation is where the pores and fractures of the ground are saturated with water. It can also be simply explained as the depth below which the ground is saturated.
T ...
that stands at or near the
land surface
Terrain or relief (also topographical relief) involves the vertical and horizontal dimensions of land surface. The term bathymetry is used to describe underwater relief, while hypsometry studies terrain relative to sea level. The Latin wo ...
for a long enough period each year to support
aquatic plants
Aquatic plants are plants that have adapted to living in aquatic environments (saltwater or freshwater). They are also referred to as hydrophytes or macrophytes to distinguish them from algae and other microphytes. A macrophyte is a plant that ...
.
A more concise definition is a community composed of
hydric soil Hydric soil is soil which is permanently or seasonally saturated by water, resulting in anaerobic conditions, as found in wetlands.
Overview
Most soils are aerobic. This is important because plant roots respire (that is, they consume oxygen and ...
and
hydrophytes
Aquatic plants are plants that have adapted to living in aquatic environments (saltwater or freshwater). They are also referred to as hydrophytes or macrophytes to distinguish them from algae and other microphytes. A macrophyte is a plant that ...
.
Wetlands have also been described as
ecotone
An ecotone is a transition area between two biological communities, where two communities meet and integrate. It may be narrow or wide, and it may be local (the zone between a field and forest) or regional (the transition between forest and gras ...
s, providing a transition between dry land and water bodies. Wetlands exist "...at the interface between truly
terrestrial
Terrestrial refers to things related to land or the planet Earth.
Terrestrial may also refer to:
* Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on or near the ground, as opposed to ...
ecosystems and aquatic systems, making them inherently different from each other, yet highly dependent on both."
In environmental decision-making, there are subsets of definitions that are agreed upon to make regulatory and policy decisions.
Under the Ramsar international wetland conservation treaty, wetlands are defined as follows:
*Article 1.1: "...wetlands are areas of marsh,
fen
A fen is a type of peat-accumulating wetland fed by mineral-rich ground or surface water. It is one of the main types of wetlands along with marshes, swamps, and bogs. Bogs and fens, both peat-forming ecosystems, are also known as mires. T ...
,
peatland
A mire, peatland, or quagmire is a wetland area dominated by living peat-forming plants. Mires arise because of incomplete decomposition of organic matter, usually litter from vegetation, due to water-logging and subsequent anoxia. All types ...
or water, whether natural or artificial, permanent or temporary, with water that is static or flowing,
fresh
Fresh or FRESH may refer to:
People
*DJ Fresh (born 1977), UK-based drum and bass artist
*DJ Fresh (producer), US-based R&B producer born Marqus Brown
Arts, entertainment, and media Films
* ''Fresh'' (1994 film), a crime film
* ''Fresh'' (200 ...
,
brackish
Brackish water, sometimes termed brack water, is water occurring in a natural environment that has more salinity than freshwater, but not as much as seawater. It may result from mixing seawater (salt water) and fresh water together, as in estuari ...
or
salt
Salt is a mineral composed primarily of sodium chloride (NaCl), a chemical compound belonging to the larger class of salts; salt in the form of a natural crystalline mineral is known as rock salt or halite. Salt is present in vast quantitie ...
, including areas of marine water the depth of which at low tide does not exceed six meters."
*Article 2.1: " etlandsmay incorporate
riparian
A riparian zone or riparian area is the interface between land and a river or stream. Riparian is also the proper nomenclature for one of the terrestrial biomes of the Earth. Plant habitats and communities along the river margins and banks ar ...
and coastal zones adjacent to the wetlands, and
island
An island (or isle) is an isolated piece of habitat that is surrounded by a dramatically different habitat, such as water. Very small islands such as emergent land features on atolls can be called islets, skerries, cays or keys. An island ...
s or bodies of marine water deeper than six meters at
low tide
Tides are the rise and fall of sea levels caused by the combined effects of the gravitational forces exerted by the Moon (and to a much lesser extent, the Sun) and are also caused by the Earth and Moon orbiting one another.
Tide tables can ...
lying within the wetlands."
An ecological definition of a wetland is "an ecosystem that arises when inundation by water produces soils dominated by anaerobic and aerobic processes, which, in turn, forces the biota, particularly rooted plants, to adapt to flooding".
Sometimes a precise legal definition of a wetland is required. The definition used for regulation by the United States government is: 'The term "wetlands" means those areas that are inundated or saturated by surface or ground water at a frequency and duration to support, and that under normal circumstances do support, a prevalence of vegetation typically adapted for life in saturated soil conditions. Wetlands generally included swamps, marshes, bogs, and similar areas.'
For each of these definitions and others, regardless of the purpose, hydrology is emphasized (shallow waters, water-logged soils). The soil characteristics and the plants and animals controlled by the wetland hydrology are often additional components of the definitions.
Types
Wetlands can be
tidal
Tidal is the adjectival form of tide.
Tidal may also refer to:
* ''Tidal'' (album), a 1996 album by Fiona Apple
* Tidal (king), a king involved in the Battle of the Vale of Siddim
* TidalCycles, a live coding environment for music
* Tidal (servic ...
(inundated by tides) or non-tidal. The water in wetlands is either
freshwater
Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does include ...
,
brackish
Brackish water, sometimes termed brack water, is water occurring in a natural environment that has more salinity than freshwater, but not as much as seawater. It may result from mixing seawater (salt water) and fresh water together, as in estuari ...
, or
saltwater
Saline water (more commonly known as salt water) is water that contains a high concentration of dissolved salts (mainly sodium chloride). On the United States Geological Survey (USGS) salinity scale, saline water is saltier than brackish water, ...
. There are four main kinds of wetlands –
marsh
A marsh is a wetland that is dominated by herbaceous rather than woody plant species.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p Marshes can often be found at ...
,
swamp
A swamp is a forested wetland.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p. Swamps are considered to be transition zones because both land and water play a role in ...
fen
A fen is a type of peat-accumulating wetland fed by mineral-rich ground or surface water. It is one of the main types of wetlands along with marshes, swamps, and bogs. Bogs and fens, both peat-forming ecosystems, are also known as mires. T ...
(bogs and fens being types of
peatlands
A mire, peatland, or quagmire is a wetland area dominated by living peat-forming plants. Mires arise because of incomplete decomposition of organic matter, usually litter from vegetation, due to water-logging and subsequent anoxia. All types ...
or
mire
A mire, peatland, or quagmire is a wetland area dominated by living peat-forming plants. Mires arise because of incomplete decomposition of organic matter, usually litter from vegetation, due to water-logging and subsequent anoxia. All types ...
s). Some experts also recognize wet meadows and
aquatic ecosystem
An aquatic ecosystem is an ecosystem formed by surrounding a body of water, in contrast to land-based terrestrial ecosystems. Aquatic ecosystems contain communities of organisms that are dependent on each other and on their environment. The tw ...
pocosin
A pocosin is a type of palustrine wetland with deep, acidic, sandy, peat soils. Groundwater saturates the soil except during brief seasonal dry spells and during prolonged droughts. Pocosin soils are nutrient-deficient (oligotrophic), especially ...
s,
floodplain
A floodplain or flood plain or bottomlands is an area of land adjacent to a river which stretches from the banks of its channel to the base of the enclosing valley walls, and which experiences flooding during periods of high discharge.Goudi ...
s,
peatlands
A mire, peatland, or quagmire is a wetland area dominated by living peat-forming plants. Mires arise because of incomplete decomposition of organic matter, usually litter from vegetation, due to water-logging and subsequent anoxia. All types ...
,
vernal pool
Vernal pools, also called vernal ponds or ephemeral pools, are seasonal pools of water that provide habitat for distinctive plants and animals. They are considered to be a distinctive type of wetland usually devoid of fish, and thus allow the safe ...
s, sinks, and many others.
The following three groups are used within
Australia
Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
to classify wetland by type: Marine and coastal zone wetlands, inland wetlands and human-made wetlands. In the US, the best known classifications are the
Cowardin classification system The Cowardin classification system is a system for classifying wetlands, devised by Lewis M. Cowardin ''et al.'' in 1979 for the United States Fish and Wildlife Service. The system includes five main types of wetlands:
# Marine wetlands- which are ...
and the hydrogeomorphic (HGM) classification system. The Cowardin system includes five main types of wetlands: marine (ocean-associated); estuarine (mixed ocean- and river-associated); riverine (within river channels); lacustrine (lake-associated); and palustrine (inland nontidal habitats).
Peatlands
Peatlands
A mire, peatland, or quagmire is a wetland area dominated by living peat-forming plants. Mires arise because of incomplete decomposition of organic matter, usually litter from vegetation, due to water-logging and subsequent anoxia. All types ...
are a unique kind of wetland where lush plant growth and slow decay of dead plants (under anoxic conditions) results in organic peat accumulating; bogs, fens, and mires are different names for peatlands.
Wetland names
Variations of names for wetland systems:
*
Bayou
In usage in the Southern United States, a bayou () is a body of water typically found in a flat, low-lying area. It may refer to an extremely slow-moving stream, river (often with a poorly defined shoreline), marshy lake, wetland, or creek. They ...
*
Flooded grasslands and savannas
Flooded grasslands and savannas is a terrestrial biome of the WWF biogeographical system, consisting of large expanses or complexes of flooded grasslands. These areas support numerous plants and animals adapted to the unique hydrologic regimes ...
*
Marsh
A marsh is a wetland that is dominated by herbaceous rather than woody plant species.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p Marshes can often be found at ...
**
Brackish marsh
Brackish marshes develop from salt marshes where a significant freshwater influx dilutes the seawater to brackish levels of salinity. This commonly happens upstream from salt marshes by estuaries of coastal rivers or near the mouths of coastal r ...
**
Freshwater marsh
A freshwater marsh is a non-tidal, non-forested marsh wetland that contains fresh water, and is continuously or frequently flooded. Freshwater marshes primarily consist of sedges, grasses, and emergent plants. Freshwater marshes are usually found ...
*
Mire
A mire, peatland, or quagmire is a wetland area dominated by living peat-forming plants. Mires arise because of incomplete decomposition of organic matter, usually litter from vegetation, due to water-logging and subsequent anoxia. All types ...
**
Fen
A fen is a type of peat-accumulating wetland fed by mineral-rich ground or surface water. It is one of the main types of wetlands along with marshes, swamps, and bogs. Bogs and fens, both peat-forming ecosystems, are also known as mires. T ...
Riparian zone
A riparian zone or riparian area is the interface between land and a river or stream. Riparian is also the proper nomenclature for one of the terrestrial biomes of the Earth. Plant habitats and communities along the river margins and banks ar ...
*
Swamp
A swamp is a forested wetland.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p. Swamps are considered to be transition zones because both land and water play a role in ...
**
Freshwater swamp forest
Freshwater swamp forests, or flooded forests, are forests which are inundated with freshwater, either permanently or seasonally. They normally occur along the lower reaches of rivers and around freshwater lakes. Freshwater swamp forests are found ...
**
Coniferous swamp
Coniferous swamps are forested wetlands in which the dominant trees are lowland conifers such as northern white cedar (''Thuja occidentalis''). The soil in these swamp areas is typically saturated for most of the growing season and is occasional ...
**
Peat swamp forest
Peat swamp forests are tropical moist forests where waterlogged soil prevents dead leaves and wood from fully decomposing. Over time, this creates a thick layer of acidic peat. Large areas of these forests are being logged at high rates.
Peat ...
**
Mangrove swamp
Mangrove forests, also called mangrove swamps, mangrove thickets or mangals, are productive wetlands that occur in coastal intertidal zones. Mangrove forests grow mainly at tropical and subtropical latitudes because mangroves cannot withstand fre ...
*
Vernal pool
Vernal pools, also called vernal ponds or ephemeral pools, are seasonal pools of water that provide habitat for distinctive plants and animals. They are considered to be a distinctive type of wetland usually devoid of fish, and thus allow the safe ...
Some wetlands have localized names unique to a region such as the prairie potholes of North America's northern plain,
pocosin
A pocosin is a type of palustrine wetland with deep, acidic, sandy, peat soils. Groundwater saturates the soil except during brief seasonal dry spells and during prolonged droughts. Pocosin soils are nutrient-deficient (oligotrophic), especially ...
s, Carolina bays and baygallsTexas Parks and Wildlife. Ecological Mapping systems of Texas: West Gulf Coastal Plain Seepage Swamp and Baygall '. Retrieved 7 July 2020 of the Southeastern US, mallines of Argentina, Mediterranean seasonal ponds of Europe and California, turloughs of Ireland,
billabong
Billabong ( ) is an Australian term for an oxbow lake, an isolated pond left behind after a river changes course. Billabongs are usually formed when the path of a creek or river changes, leaving the former branch with a dead end. As a result ...
s of Australia, among many others.
Locations
By temperature zone
Wetlands are found throughout the world in different climates. Temperatures vary greatly depending on the location of the wetland. Many of the world's wetlands are in the
temperate zone
In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout t ...
s, midway between the North or South Poles and the equator. In these zones, summers are warm and winters are cold, but temperatures are not extreme. In subtropical zone wetlands, such as along the
Gulf of Mexico
The Gulf of Mexico ( es, Golfo de México) is an oceanic basin, ocean basin and a marginal sea of the Atlantic Ocean, largely surrounded by the North American continent. It is bounded on the northeast, north and northwest by the Gulf Coast of ...
, average temperatures might be . Wetlands in the
tropics
The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in
the Southern Hemisphere at S. The tropics are also referred to ...
are subjected to much higher temperatures for a large portion of the year. Temperatures for wetlands on the
Arabian Peninsula
The Arabian Peninsula, (; ar, شِبْهُ الْجَزِيرَةِ الْعَرَبِيَّة, , "Arabian Peninsula" or , , "Island of the Arabs") or Arabia, is a peninsula of Western Asia, situated northeast of Africa on the Arabian Plate ...
can exceed and these habitats would therefore be subject to rapid evaporation. In northeastern
Siberia
Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part of ...
, which has a polar climate, wetland temperatures can be as low as .
Peatland
A mire, peatland, or quagmire is a wetland area dominated by living peat-forming plants. Mires arise because of incomplete decomposition of organic matter, usually litter from vegetation, due to water-logging and subsequent anoxia. All types ...
s in arctic and subarctic regions insulate the
permafrost
Permafrost is ground that continuously remains below 0 °C (32 °F) for two or more years, located on land or under the ocean. Most common in the Northern Hemisphere, around 15% of the Northern Hemisphere or 11% of the global surface ...
, thus delaying or preventing its thawing during summer, as well as inducing its formation.
By precipitation amount
The amount of precipitation a wetland receives varies widely according to its area. Wetlands in
Wales
Wales ( cy, Cymru ) is a Countries of the United Kingdom, country that is part of the United Kingdom. It is bordered by England to the Wales–England border, east, the Irish Sea to the north and west, the Celtic Sea to the south west and the ...
,
Scotland
Scotland (, ) is a country that is part of the United Kingdom. Covering the northern third of the island of Great Britain, mainland Scotland has a border with England to the southeast and is otherwise surrounded by the Atlantic Ocean to the ...
, and western
Ireland
Ireland ( ; ga, Éire ; Ulster Scots dialect, Ulster-Scots: ) is an island in the Atlantic Ocean, North Atlantic Ocean, in Northwestern Europe, north-western Europe. It is separated from Great Britain to its east by the North Channel (Grea ...
typically receive about per year. In some places in
Southeast Asia
Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical United Nations geoscheme for Asia#South-eastern Asia, south-eastern region of Asia, consistin ...
, where heavy rains occur, they can receive up to . In some drier regions, wetlands exist where as little as precipitation occurs each year.
Temporal variation:
*
Perennial
A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...
systems
*
Seasonal
A season is a division of the year based on changes in weather, ecology, and the number of daylight hours in a given region. On Earth, seasons are the result of the axial parallelism of Earth's tilted orbit around the Sun. In temperate and po ...
systems
*Episodic (periodic or intermittent) systems
*
Ephemeral
Ephemerality (from the Greek word , meaning 'lasting only one day') is the concept of things being transitory, existing only briefly. Academically, the term ephemeral constitutionally describes a diverse assortment of things and experiences, fr ...
(short-lived) systems
Surface flow may occur in some segments, with subsurface flow in other segments.
Processes
Wetlands vary widely due to local and regional differences in
topography
Topography is the study of the forms and features of land surfaces. The topography of an area may refer to the land forms and features themselves, or a description or depiction in maps.
Topography is a field of geoscience and planetary sci ...
,
hydrology
Hydrology () is the scientific study of the movement, distribution, and management of water on Earth and other planets, including the water cycle, water resources, and environmental watershed sustainability. A practitioner of hydrology is calle ...
,
vegetation
Vegetation is an assemblage of plant species and the ground cover they provide. It is a general term, without specific reference to particular taxa, life forms, structure, spatial extent, or any other specific botanical or geographic character ...
, and other factors, including human involvement. Other important factors include fertility, natural disturbance, competition,
herbivory
A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpart ...
, burial and salinity. When
peat
Peat (), also known as turf (), is an accumulation of partially decayed vegetation or organic matter. It is unique to natural areas called peatlands, bogs, mires, moors, or muskegs. The peatland ecosystem covers and is the most efficien ...
fen
A fen is a type of peat-accumulating wetland fed by mineral-rich ground or surface water. It is one of the main types of wetlands along with marshes, swamps, and bogs. Bogs and fens, both peat-forming ecosystems, are also known as mires. T ...
s arise.
Hydrology
The most important factor producing wetlands is hydrology, or
flooding
A flood is an overflow of water ( or rarely other fluids) that submerges land that is usually dry. In the sense of "flowing water", the word may also be applied to the inflow of the tide. Floods are an area of study of the discipline hydrolog ...
. The duration of flooding or prolonged soil saturation by
groundwater
Groundwater is the water present beneath Earth's surface in rock and soil pore spaces and in the fractures of rock formations. About 30 percent of all readily available freshwater in the world is groundwater. A unit of rock or an unconsolidate ...
determines whether the resulting wetland has aquatic,
marsh
A marsh is a wetland that is dominated by herbaceous rather than woody plant species.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p Marshes can often be found at ...
or
swamp
A swamp is a forested wetland.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p. Swamps are considered to be transition zones because both land and water play a role in ...
vegetation
Vegetation is an assemblage of plant species and the ground cover they provide. It is a general term, without specific reference to particular taxa, life forms, structure, spatial extent, or any other specific botanical or geographic character ...
. Other important factors include soil fertility, natural disturbance, competition,
herbivory
A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpart ...
, burial, and salinity. When
peat
Peat (), also known as turf (), is an accumulation of partially decayed vegetation or organic matter. It is unique to natural areas called peatlands, bogs, mires, moors, or muskegs. The peatland ecosystem covers and is the most efficien ...
fen
A fen is a type of peat-accumulating wetland fed by mineral-rich ground or surface water. It is one of the main types of wetlands along with marshes, swamps, and bogs. Bogs and fens, both peat-forming ecosystems, are also known as mires. T ...
s develop.
Wetland hydrology is associated with the spatial and temporal dispersion, flow, and physio-chemical attributes of surface and ground waters. Sources of hydrological flows into wetlands are predominantly
precipitation
In meteorology, precipitation is any product of the condensation of atmospheric water vapor that falls under gravitational pull from clouds. The main forms of precipitation include drizzle, rain, sleet, snow, ice pellets, graupel and hail. ...
, surface water (saltwater or freshwater), and
groundwater
Groundwater is the water present beneath Earth's surface in rock and soil pore spaces and in the fractures of rock formations. About 30 percent of all readily available freshwater in the world is groundwater. A unit of rock or an unconsolidate ...
. Water flows out of wetlands by
evapotranspiration
Evapotranspiration (ET) is the combined processes by which water moves from the earth’s surface into the atmosphere. It covers both water evaporation (movement of water to the air directly from soil, canopies, and water bodies) and transpi ...
, surface flows and
tide
Tides are the rise and fall of sea levels caused by the combined effects of the gravity, gravitational forces exerted by the Moon (and to a much lesser extent, the Sun) and are also caused by the Earth and Moon orbiting one another.
Tide t ...
s, and subsurface water outflow.
Hydrodynamics
In physics and engineering, fluid dynamics is a subdiscipline of fluid mechanics that describes the flow of fluids—liquids and gases. It has several subdisciplines, including ''aerodynamics'' (the study of air and other gases in motion) and ...
(the movement of water through and from a wetland) affects hydro-periods (temporal fluctuations in water levels) by controlling the water balance and water storage within a wetland.
Landscape characteristics control wetland hydrology and water chemistry. The O2 and CO2
concentrations
In chemistry, concentration is the abundance of a constituent divided by the total volume of a mixture. Several types of mathematical description can be distinguished: '' mass concentration'', ''molar concentration'', ''number concentration'', an ...
of water depend on
temperature
Temperature is a physical quantity that expresses quantitatively the perceptions of hotness and coldness. Temperature is measured with a thermometer.
Thermometers are calibrated in various temperature scales that historically have relied o ...
,
atmospheric pressure
Atmospheric pressure, also known as barometric pressure (after the barometer), is the pressure within the atmosphere of Earth. The standard atmosphere (symbol: atm) is a unit of pressure defined as , which is equivalent to 1013.25 millibars, 7 ...
and mixing with the air (from winds or water flows). Water chemistry within wetlands is determined by the pH,
salinity
Salinity () is the saltiness or amount of salt dissolved in a body of water, called saline water (see also soil salinity). It is usually measured in g/L or g/kg (grams of salt per liter/kilogram of water; the latter is dimensionless and equal ...
, nutrients,
conductivity
Conductivity may refer to:
*Electrical conductivity, a measure of a material's ability to conduct an electric current
**Conductivity (electrolytic), the electrical conductivity of an electrolyte in solution
**Ionic conductivity (solid state), elec ...
, soil composition,
hardness
In materials science, hardness (antonym: softness) is a measure of the resistance to localized plastic deformation induced by either mechanical indentation or abrasion. In general, different materials differ in their hardness; for example hard ...
, and the sources of water. Water chemistry varies across landscapes and climatic regions. Wetlands are generally
minerotrophic Minerotrophic refers to environments that receive nutrients primarily through groundwater that flows through mineral-rich soils or rock,Environment Canada (2014). Ontario wetland evaluation system: Northern Manual, 1st edition, version 3.2. Queen’ ...
(waters contain dissolved materials from soils) with the exception of
ombrotrophic
Ombrotrophic ("cloud-fed"), from Ancient Greek ὄμβρος (''ómvros'') meaning "rain" and τροφή (''trofí'') meaning "food"), refers to Soil, soils or vegetation which receive all of their water and nutrients from precipitation, rather ...
bogs that are fed only by water from precipitation.
Because bogs receive most of their water from the
atmosphere
An atmosphere () is a layer of gas or layers of gases that envelop a planet, and is held in place by the gravity of the planetary body. A planet retains an atmosphere when the gravity is great and the temperature of the atmosphere is low. A s ...
, their water usually has low mineral ionic composition. In contrast, wetlands fed by groundwater or tides have a higher concentration of dissolved nutrients and minerals.
Fen peatlands receive water both from precipitation and ground water in varying amounts so their water chemistry ranges from acidic with low levels of dissolved minerals to alkaline with high accumulation of
calcium
Calcium is a chemical element with the symbol Ca and atomic number 20. As an alkaline earth metal, calcium is a reactive metal that forms a dark oxide-nitride layer when exposed to air. Its physical and chemical properties are most similar to ...
and
magnesium
Magnesium is a chemical element with the symbol Mg and atomic number 12. It is a shiny gray metal having a low density, low melting point and high chemical reactivity. Like the other alkaline earth metals (group 2 of the periodic ta ...
.
Role of salinity
Salinity has a strong influence on wetland water chemistry, particularly in coastal wetlands and in arid and semiarid regions with large precipitation deficits. Natural salinity is regulated by interactions between ground and surface water, which may be influenced by human activity.
Soil
Carbon
Carbon () is a chemical element with the symbol C and atomic number 6. It is nonmetallic and tetravalent
In chemistry, the valence (US spelling) or valency (British spelling) of an element is the measure of its combining capacity with o ...
is the major nutrient cycled within wetlands. Most nutrients, such as
sulfur
Sulfur (or sulphur in British English) is a chemical element with the symbol S and atomic number 16. It is abundant, multivalent and nonmetallic. Under normal conditions, sulfur atoms form cyclic octatomic molecules with a chemical formula ...
,
phosphorus
Phosphorus is a chemical element with the symbol P and atomic number 15. Elemental phosphorus exists in two major forms, white phosphorus and red phosphorus, but because it is highly reactive, phosphorus is never found as a free element on Ear ...
,
carbon
Carbon () is a chemical element with the symbol C and atomic number 6. It is nonmetallic and tetravalent
In chemistry, the valence (US spelling) or valency (British spelling) of an element is the measure of its combining capacity with o ...
, and
nitrogen
Nitrogen is the chemical element with the symbol N and atomic number 7. Nitrogen is a nonmetal and the lightest member of group 15 of the periodic table, often called the pnictogens. It is a common element in the universe, estimated at se ...
are found within the soil of wetlands.
Anaerobic
Anaerobic means "living, active, occurring, or existing in the absence of free oxygen", as opposed to aerobic which means "living, active, or occurring only in the presence of oxygen." Anaerobic may also refer to:
* Anaerobic adhesive, a bonding a ...
and
aerobic respiration
Cellular respiration is the process by which biological fuels are oxidised in the presence of an inorganic electron acceptor such as oxygen to produce large amounts of energy, to drive the bulk production of ATP. Cellular respiration may be des ...
in the soil influences the nutrient cycling of carbon, hydrogen, oxygen, and nitrogen, and the solubility of phosphorus thus contributing to the chemical variations in its water. Wetlands with low pH and saline conductivity may reflect the presence of acid
sulfate
The sulfate or sulphate ion is a polyatomic anion with the empirical formula . Salts, acid derivatives, and peroxides of sulfate are widely used in industry. Sulfates occur widely in everyday life. Sulfates are salts of sulfuric acid and many ar ...
s and wetlands with average salinity levels can be heavily influenced by calcium or magnesium. Biogeochemical processes in wetlands are determined by soils with low
redox
Redox (reduction–oxidation, , ) is a type of chemical reaction in which the oxidation states of substrate (chemistry), substrate change. Oxidation is the loss of Electron, electrons or an increase in the oxidation state, while reduction ...
potential. Wetland soils are identified by redoxymorphic mottles (often from iron oxide rust) or low chroma intensity, as determined by the
Munsell Color System
In colorimetry, the Munsell color system is a color space that specifies colors based on three properties of color: hue (basic color), chroma (color intensity), and value ( lightness). It was created by Professor Albert H. Munsell in the first ...
.
Water chemistry
Due to the low dissolved oxygen (DO) content, and relatively low nutrient balance of wetland environments, most wetlands are very susceptible to alterations in water chemistry. Key factors that are assessed to determine water quality include:
* Major
anion
An ion () is an atom or molecule with a net electrical charge.
The charge of an electron is considered to be negative by convention and this charge is equal and opposite to the charge of a proton, which is considered to be positive by convent ...
analysis: (HCO3−,Cl−,NO3−,SO42-)
* Major
cation
An ion () is an atom or molecule with a net electrical charge.
The charge of an electron is considered to be negative by convention and this charge is equal and opposite to the charge of a proton, which is considered to be positive by convent ...
Conductivity
Conductivity may refer to:
*Electrical conductivity, a measure of a material's ability to conduct an electric current
**Conductivity (electrolytic), the electrical conductivity of an electrolyte in solution
**Ionic conductivity (solid state), elec ...
- conductivity increases with more dissolved ions in the water
*
Turbidity
Turbidity is the cloudiness or haziness of a fluid caused by large numbers of individual particles that are generally invisible to the naked eye, similar to smoke in air. The measurement of turbidity is a key test of water quality.
Fluids can ...
*
Dissolved oxygen
Oxygen saturation (symbol SO2) is a relative measure of the concentration of oxygen that is dissolved or carried in a given medium as a proportion of the maximal concentration that can be dissolved in that medium at the given temperature. It ca ...
* Temperature
*
Total dissolved solids
Total dissolved solids (TDS) is a measure of the dissolved combined content of all inorganic and organic substances present in a liquid in molecular, ionized, or micro-granular ( colloidal sol) suspended form. TDS concentrations are often repor ...
* Gas emissions (
carbon dioxide
Carbon dioxide (chemical formula ) is a chemical compound made up of molecules that each have one carbon atom covalently double bonded to two oxygen atoms. It is found in the gas state at room temperature. In the air, carbon dioxide is transpar ...
and
methane
Methane ( , ) is a chemical compound with the chemical formula (one carbon atom bonded to four hydrogen atoms). It is a group-14 hydride, the simplest alkane, and the main constituent of natural gas. The relative abundance of methane on Eart ...
; CO2 and CH4)
These chemical factors can be used to quantify wetland disturbances, and often provide information as to whether a wetland is fed by precipitation, surface water or groundwater, due to the different ion characteristics of the different water sources. Wetlands are adept at impacting the water chemistry of streams or water bodies that interact with them, and can process ions that result from water pollution such as
acid mine drainage
Acid mine drainage, acid and metalliferous drainage (AMD), or acid rock drainage (ARD) is the outflow of acidic water from metal mines or coal mines.
Acid rock drainage occurs naturally within some environments as part of the rock weathering ...
or
urban runoff
Urban runoff is surface runoff of rainwater, landscape irrigation, and car washing created by urbanization. Impervious surfaces (roads, parking lots and sidewalks) are constructed during land development. During rain , storms and other precipit ...
.,
Biota
The biota of a wetland system includes its plants (
flora
Flora is all the plant life present in a particular region or time, generally the naturally occurring (indigenous) native plants. Sometimes bacteria and fungi are also referred to as flora, as in the terms '' gut flora'' or '' skin flora''.
E ...
) and animals (
fauna
Fauna is all of the animal life present in a particular region or time. The corresponding term for plants is ''flora'', and for fungi, it is '' funga''. Flora, fauna, funga and other forms of life are collectively referred to as '' biota''. Zoo ...
) and
microbes
A microorganism, or microbe,, ''mikros'', "small") and ''organism'' from the el, ὀργανισμός, ''organismós'', "organism"). It is usually written as a single word but is sometimes hyphenated (''micro-organism''), especially in olde ...
(bacteria, fungi). The most important factor affecting the biota is the hydroperiod, or the duration of flooding. Other important factors include fertility and salinity of the water or soils. The chemistry of water flowing into wetlands depends on the source of water, the geological material that it flows through and the nutrients discharged from organic matter in the soils and plants at higher elevations. Biota may vary within a wetland seasonally or in response to flood regimes.
Flora
There are four main groups of
hydrophyte
Aquatic plants are plants that have adapted to living in aquatic environments (saltwater or freshwater). They are also referred to as hydrophytes or macrophytes to distinguish them from algae and other microphytes. A macrophyte is a plant that g ...
s that are found in wetland systems throughout the world.
Submerged wetland vegetation can grow in saline and fresh-water conditions. Some species have underwater flowers, while others have long stems to allow the flowers to reach the surface. Submerged species provide a food source for native fauna, habitat for invertebrates, and also possess filtration capabilities. Examples include
seagrasses
Seagrasses are the only flowering plants which grow in marine environments. There are about 60 species of fully marine seagrasses which belong to four families (Posidoniaceae, Zosteraceae, Hydrocharitaceae and Cymodoceaceae), all in the orde ...