Verbascum thapsus
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''Verbascum thapsus'', the great mullein, greater mullein or common mullein is a species of
mullein ''Verbascum'' is a genus of over 450 species of flowering plants, common name mullein (), in the figwort family Scrophulariaceae. They are native to Europe and Asia, with the highest species diversity in the Mediterranean. Mullein or "mullein le ...
native to Europe, northern Africa, and Asia, and introduced in the Americas and Australia. It is a hairy
biennial plant A biennial plant is a flowering plant that, generally in a temperate climate, takes two years to complete its biological life cycle. Life cycle In its first year, the biennal plant undergoes primary growth, during which its vegetative structures ...
that can grow to 2 m tall or more. Its small, yellow flowers are densely grouped on a tall stem, which grows from a large rosette of leaves. It grows in a wide variety of habitats, but prefers well-lit, disturbed soils, where it can appear soon after the ground receives light, from long-lived seeds that persist in the
soil seed bank The soil seed bank is the natural storage of seeds, often dormant, within the soil of most ecosystems. The study of soil seed banks started in 1859 when Charles Darwin observed the emergence of seedlings using soil samples from the bottom of a lak ...
. It is a common weedy plant that spreads by prolifically producing seeds, and has become invasive in temperate world regions. It is a minor problem for most agricultural crops, since it is not a competitive species, being intolerant of shade from other plants and unable to survive
tilling Tilling can mean: * Tillage, an agricultural preparation of the soil. * TILLING (molecular biology) * Tilling is a fictional town in the Mapp and Lucia novels of E. F. Benson. * Tilling Green, Ledshire, is a fictional village in Patricia Wentwo ...
. It also hosts many insects, some of which can be harmful to other plants. Although individuals are easy to remove by hand, populations are difficult to eliminate permanently. Although commonly used in
traditional medicine Traditional medicine (also known as indigenous medicine or folk medicine) comprises medical aspects of traditional knowledge that developed over generations within the folk beliefs of various societies, including indigenous peoples, before the ...
, no
approved drugs An approved drug is a medicinal preparation that has been validated for a therapeutic use by a ruling authority of a government. This process is usually specific by country, unless specified otherwise. Process by country United States In the ...
are made from this plant. It has been used to make
dye A dye is a colored substance that chemically bonds to the substrate to which it is being applied. This distinguishes dyes from pigments which do not chemically bind to the material they color. Dye is generally applied in an aqueous solution an ...
s and
torch A torch is a stick with combustible material at one end, which is ignited and used as a light source. Torches have been used throughout history, and are still used in processions, symbolic and religious events, and in juggling entertainment. In ...
es.


Description

''V. thapsus'' is a
dicotyledon The dicotyledons, also known as dicots (or, more rarely, dicotyls), are one of the two groups into which all the flowering plants (angiosperms) were formerly divided. The name refers to one of the typical characteristics of the group: namely, t ...
ous plant that produces a rosette of leaves in its first year of growth.Blamey, M., & Grey-Wilson, C. (1989). ''Flora of Britain and Northern Europe''. Hodder & Stoughton, . The leaves are large, up to 50 cm long. The second-year plants normally produce a single unbranched stem, usually 1–2 m tall. In the eastern part of its range in China, it is, however, only reported to grow up to 1.5 m tall. The tall, pole-like stems end in a dense
spike Spike, spikes, or spiking may refer to: Arts, entertainment, and media Books * ''The Spike'' (novel), a novel by Arnaud de Borchgrave * ''The Spike'' (book), a nonfiction book by Damien Broderick * ''The Spike'', a starship in Peter F. Hamilto ...
of flowers that can occupy up to half the stem length. All parts of the plants are covered with star-shaped
trichome Trichomes (); ) are fine outgrowths or appendages on plants, algae, lichens, and certain protists. They are of diverse structure and function. Examples are hairs, glandular hairs, scales, and papillae. A covering of any kind of hair on a pla ...
s. This cover is particularly thick on the leaves, giving them a silvery appearance. The species'
chromosome number Ploidy () is the number of complete sets of chromosomes in a cell, and hence the number of possible alleles for autosomal and pseudoautosomal genes. Sets of chromosomes refer to the number of maternal and paternal chromosome copies, respectively ...
is 2n = 36. On flowering plants, the leaves are alternately arranged up the stem. They are thick and
decurrent ''Decurrent'' (sometimes decurring) is a term used in botany and mycology to describe plant or fungal parts that extend downward. In botany, the term is most often applied to leaf blades that partly wrap or have wings around the stem or petio ...
, with much variation in leaf shape between the upper and lower leaves on the stem, ranging from oblong to oblanceolate, and reaching sizes up to 50 cm long and 14 cm across (19 inches long and 5 inches wide).Huxley, A., ed. (1992). ''New RHS Dictionary of Gardening''. Macmillan. They become smaller higher up the stem, and less strongly decurrent down the stem. The flowering stem is solid and 2–2.5 cm (nearly an inch) across, and occasionally branched just below the
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed o ...
, usually following damage. After flowering and seed release, the stem and fruits usually persist in winter, drying into dark brown, stiff structures of densely packed, ovoid-shaped, and dry seed capsules. The dried stems may persist into the following spring or even the next summer. The plant produces a shallow
taproot A taproot is a large, central, and dominant root from which other roots sprout laterally. Typically a taproot is somewhat straight and very thick, is tapering in shape, and grows directly downward. In some plants, such as the carrot, the taproo ...
. Flowers are pentamerous with (usually) five
stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...
, a five-lobed
calyx tube A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coined b ...
, and a five-petalled corolla, the latter bright yellow and an wide. The flowers are almost
sessile Sessility, or sessile, may refer to: * Sessility (motility), organisms which are not able to move about * Sessility (botany), flowers or leaves that grow directly from the stem or peduncle of a plant * Sessility (medicine), tumors and polyps that ...
, with very short
pedicels In botany, a pedicel is a stem that attaches a single flower to the inflorescence. Such inflorescences are described as ''pedicellate''. Description Pedicel refers to a structure connecting a single flower to its inflorescence. In the absenc ...
(2 mm, 0.08 in). The five stamens are of two types, with the three upper stamens being shorter, their filaments covered by yellow or whitish hairs, and having smaller
anther The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filam ...
s, while the lower two stamens have
glabrous Glabrousness (from the Latin '' glaber'' meaning "bald", "hairless", "shaved", "smooth") is the technical term for a lack of hair, down, setae, trichomes or other such covering. A glabrous surface may be a natural characteristic of all or part of ...
filaments and larger anthers.They are all hairy in subspecies ''V. crassifolium'' and ''V. giganteum''. The plant produces small, ovoid (6 mm, 0.24 in) capsules that split open by way of two valves, each capsule containing large numbers of minute, brown
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiospe ...
s less than 1 mm (0.04 in) in size, marked with longitudinal ridges. A white-flowered form, ''V. thapsus'' f. ''candicans'', is known to occur. Flowering lasts up to three months from early to late summer (June to August in northern Europe), with flowering starting at the bottom of the spike and progressing irregularly upward; each flower opens for part of a day and only a few open at the same time around the stem.


Taxonomy

For the purpose of
botanical nomenclature Botanical nomenclature is the formal, scientific naming of plants. It is related to, but distinct from Alpha taxonomy, taxonomy. Plant taxonomy is concerned with grouping and classifying plants; botanical nomenclature then provides names for the ...
, ''Verbascum thapsus'' was first described by
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
in his 1753 '. The
specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''thapsus'' had been first used by
Theophrastus Theophrastus (; grc-gre, Θεόφραστος ; c. 371c. 287 BC), a Greek philosopher and the successor to Aristotle in the Peripatetic school. He was a native of Eresos in Lesbos.Gavin Hardy and Laurence Totelin, ''Ancient Botany'', Routledge ...
(as , ') for an unspecified herb from the
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic peri ...
settlement of
Thapsos Thapsos ( el, Θάψος) was a prehistoric village in Sicily of the middle Bronze Age. It was found by the Italian archaeologist Paolo Orsi on the small peninsula of Magnisi, near Priolo Gargallo. In its vicinity was born the Thapsos culture ...
, near modern
Syracuse Syracuse may refer to: Places Italy *Syracuse, Sicily, or spelled as ''Siracusa'' *Province of Syracuse United States *Syracuse, New York **East Syracuse, New York **North Syracuse, New York *Syracuse, Indiana * Syracuse, Kansas *Syracuse, Miss ...
,
Sicily (man) it, Siciliana (woman) , population_note = , population_blank1_title = , population_blank1 = , demographics_type1 = Ethnicity , demographics1_footnotes = , demographi ...
, though it is often assimilated to the ancient
Tunisia ) , image_map = Tunisia location (orthographic projection).svg , map_caption = Location of Tunisia in northern Africa , image_map2 = , capital = Tunis , largest_city = capital , ...
n city of
Thapsus Thapsus, also known as Tampsus and as Thapsus Minor to distinguish it from Thapsus in Sicily, was a Carthaginian and Roman port near present-day Bekalta, Tunisia. Geography Thapsus was established on Ras ed-Dimas, an easily defended promontory ...
. At the time, no
type specimen In biology, a type is a particular wiktionary:en:specimen, specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to a ...
was specified, as the practice only arose later, in the 19th century. When a lectotype (type selected amongst original material) was designated, it was assigned to specimen 242.1 of Linnaeus' herbarium, the only ''V. thapsus'' specimen. The species had previously been designated as
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
for ''
Verbascum ''Verbascum'' is a genus of over 450 species of flowering plants, common name mullein (), in the figwort family Scrophulariaceae. They are native to Europe and Asia, with the highest species diversity in the Mediterranean. Mullein or "mullein le ...
''. European plants exhibit considerable
phenotypical In genetics, the phenotype () is the set of observable characteristics or traits of an organism. The term covers the organism's morphology or physical form and structure, its developmental processes, its biochemical and physiological proper ...
variation, which has led to the plant acquiring many
synonyms A synonym is a word, morpheme, or phrase that means exactly or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are all ...
over the years. Introduced American populations show much less variation. The taxonomy of ''Verbascum'' has not undergone any significant revision since Svanve Mürbeck's monographies in the 1930s, with the exception of the work of Arthur Huber-Morath, who used informal grouping in organizing the genus for the florae of Iran and Turkey to account for many intermediate species. Since Huber-Morath's groups are not taxonomical, Mürbeck's treatment is the most current one available, as no study has yet sought to apply genetic or molecular data extensively to the genus. In Mürbeck's classification, ''V. thapsus'' is placed in section Bothrospermae subsect. ''Fasciculata'' (or sect. ''Verbascum'' subsect. ''Verbascum'' depending on nomenclatural choices) alongside species such as ''
Verbascum nigrum ''Verbascum nigrum'', the black mullein or dark mullein, is a species of biennial or short-lived perennial herbaceous plant in the mullein genus ''Verbascum'', native to dry open sites in temperate Europe. It grows to . ''Verbascum nigrum'' form ...
'' (black or dark mullein), ''
Verbascum lychnitis ''Verbascum lychnitis'', the white mullein, is a flowering plant in the figwort family (Scrophulariaceae) native to Asia and Europe. It has naturalized in parts of North America. The species was first formally named by Carl Linnaeus in 1753. Desp ...
'' (white mullein), and ''
Verbascum sinuatum ''Verbascum sinuatum'', commonly known as the scallop-leaved mullein,Wild Flowers of Israel
Verbascum ...
'' (wavy-leaved mullein).


Subspecies and hybrids

The three usually recognized
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
are: * ''V. t. thapsus''; type, widespread. * ''V. t. crassifolium'' (Lam.) Murb.; Mediterranean region and to 2000 metres in southwestern Austria. (syn. subsp. ''montanum'' (Scrad.) Bonnier & Layens) * ''V. t. giganteum'' (Willk.) Nyman; Spain, endemic. In all subspecies but the type, the lower stamens are also hairy. In ''V. t. crassifolium'', the hairiness is less dense and often absent from the upper part of the anthers, while lower leaves are hardly decurrent and have longer petioles. In ''V. t. giganteum'', the hairs are densely white tomentose, and lower leaves are strongly decurrent. ''V. t. crassifolium'' also differs from the type in having slightly larger flowers, which measure 15–30 mm wide, whereas in the type, they are 12–20 mm in diameter. Both ''V. t. giganteum'' and ''V. t. crassifolium'' were originally described as species. Due to its morphological variation, ''V. thapsus'' has had a great many subspecies described. A recent revision led its author to maintain ''V. giganteum'' but sink ''V. crassifolium'' into synonymy. The plant is also parent to several hybrids (''see table''). Of these, the most common is ''V.'' × ''semialbum'' Chaub. (× ''V. nigrum''). All occur in Eurasia, and three, ''V.'' × ''kerneri'' Fritsch, ''V.'' × ''pterocaulon'' Franch. and ''V.'' × ''thapsi'' L. ( syn. ''V.'' × ''spurium'' W.D.J.Koch), have also been reported in North America.


Common names

''V. thapsus'' is known by a variety of names. European reference books call it "great mullein". Watts (2000), pp. 633–634. In North America, "common mullein" is used while western United States residents commonly refer to mullein as "cowboy toilet paper". In the 19th century, it had well over 40 different common names in English alone. Some of the more whimsical ones included "hig candlewick", "Indian rag weed", "bullicks lungwort", "Adams-rod", "hare's-beard", and "ice-leaf". Vernacular names include innumerable references to the plant's hairiness: "woolly mullein", "velvet mullein", or "blanket mullein", "beggar's blanket", "Moses' blanket", "poor man's blanket", "Our Lady's blanket", or "old man's blanket", and "feltwort", and so on ("flannel" is another common generic name). "Mullein" itself derives from the French word for "soft". Some names refer to the plant's size and shape: "shepherd's club(s)" or "staff", "
Aaron's rod Aaron's rod refers to any of the walking sticks carried by Moses's brother, Aaron, in the Torah. The Bible tells how, along with Moses's rod, Aaron's rod was endowed with miraculous power during the Plagues of Egypt that preceded the Exodus. T ...
" (a name it shares with a number of other plants with tall, yellow inflorescences), and a plethora of other "X's staff" and "X's rod". The name "velvet dock" or "mullein dock" is also recorded, where "dock" is a British name applied to any broad-leaved plant.


Distribution and habitat

''V. thapsus'' has a wide native range including Europe, northern Africa, and Asia, from the
Azores ) , motto =( en, "Rather die free than subjected in peace") , anthem= ( en, "Anthem of the Azores") , image_map=Locator_map_of_Azores_in_EU.svg , map_alt=Location of the Azores within the European Union , map_caption=Location of the Azores wi ...
and
Canary Islands The Canary Islands (; es, Canarias, ), also known informally as the Canaries, are a Spanish autonomous community and archipelago in the Atlantic Ocean, in Macaronesia. At their closest point to the African mainland, they are west of Morocc ...
east to western
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
, north to the
British Isles The British Isles are a group of islands in the North Atlantic Ocean off the north-western coast of continental Europe, consisting of the islands of Great Britain, Ireland, the Isle of Man, the Inner and Outer Hebrides, the Northern Isles, ...
,
Scandinavia Scandinavia; Sámi languages: /. ( ) is a subregion#Europe, subregion in Northern Europe, with strong historical, cultural, and linguistic ties between its constituent peoples. In English usage, ''Scandinavia'' most commonly refers to Denmark, ...
, and
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part of ...
, and south to the
Himalayas The Himalayas, or Himalaya (; ; ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the planet's highest peaks, including the very highest, Mount Everest. Over 100 ...
.Flora Europaea: ''Verbascum thapsus''
retrieved on November 6, 2009.
In northern Europe, it grows from sea level up to 1,850 m altitude, while in China it grows at 1,400–3,200 m altitude. It has been introduced throughout the temperate world, and is established as a
weed A weed is a plant considered undesirable in a particular situation, "a plant in the wrong place", or a plant growing where it is not wanted.Harlan, J. R., & deWet, J. M. (1965). Some thoughts about weeds. ''Economic botany'', ''19''(1), 16-24. ...
in
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
,
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...
, tropical Asia,
La Réunion LA most frequently refers to Los Angeles, the second largest city in the United States. La, LA, or L.A. may also refer to: Arts and entertainment Music * La (musical note), or A, the sixth note * "L.A.", a song by Elliott Smith on ''Figure ...
, North America,
Hawaii Hawaii ( ; haw, Hawaii or ) is a state in the Western United States, located in the Pacific Ocean about from the U.S. mainland. It is the only U.S. state outside North America, the only state that is an archipelago, and the only stat ...
,
Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the east a ...
,
Hispaniola Hispaniola (, also ; es, La Española; Latin and french: Hispaniola; ht, Ispayola; tnq, Ayiti or Quisqueya) is an island in the Caribbean that is part of the Greater Antilles. Hispaniola is the most populous island in the West Indies, and th ...
, and
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
. It has also been reported in
Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...
. In the United States, it was imported very early in the 18th century and cultivated for its medicinal and
piscicide A piscicide is a chemical substance which is poisonous to fish. The primary use for piscicides is to eliminate a dominant species of fish in a body of water, as the first step in attempting to populate the body of water with a different fish. They ...
properties. By 1818, it had begun spreading so much that
Amos Eaton Amos Eaton (May 17, 1776 – May 10, 1842) was an American botanist, geologist, and educator who is considered the founder of the modern scientific prospectus in education, which was a radical departure from the American liberal arts tradition of ...
thought it was a native plant. In 1839, it was already reported in
Michigan Michigan () is a state in the Great Lakes region of the upper Midwestern United States. With a population of nearly 10.12 million and an area of nearly , Michigan is the 10th-largest state by population, the 11th-largest by area, and the ...
and in 1876, in
California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...
. It is now found commonly in all the states. In
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...
, it is most common in the
Maritime Provinces The Maritimes, also called the Maritime provinces, is a region of Eastern Canada consisting of three provinces: New Brunswick, Nova Scotia, and Prince Edward Island. The Maritimes had a population of 1,899,324 in 2021, which makes up 5.1% of Ca ...
and southern
Quebec Quebec ( ; )According to the Canadian government, ''Québec'' (with the acute accent) is the official name in Canadian French and ''Quebec'' (without the accent) is the province's official name in Canadian English is one of the thirtee ...
,
Ontario Ontario ( ; ) is one of the thirteen provinces and territories of Canada.Ontario is located in the geographic eastern half of Canada, but it has historically and politically been considered to be part of Central Canada. Located in Central Ca ...
, and
British Columbia British Columbia (commonly abbreviated as BC) is the westernmost province of Canada, situated between the Pacific Ocean and the Rocky Mountains. It has a diverse geography, with rugged landscapes that include rocky coastlines, sandy beaches, ...
, with scattered populations in between. Great mullein most frequently grows as a colonist of bare and disturbed soil, usually on sandy or chalky ones. It grows best in dry, sandy, or gravelly soils, although it can grow in a variety of habitats, including banksides, meadows, roadsides, forest clearings, and pastures. This ability to grow in a wide range of habitats has been linked to strong phenotype variation rather than
adaptation In biology, adaptation has three related meanings. Firstly, it is the dynamic evolutionary process of natural selection that fits organisms to their environment, enhancing their evolutionary fitness. Secondly, it is a state reached by the po ...
capacities.


Ecology

Great mullein is a biennial and generally requires winter dormancy before it can flower. This dormancy is linked to
starch Starch or amylum is a polymeric carbohydrate consisting of numerous glucose units joined by glycosidic bonds. This polysaccharide is produced by most green plants for energy storage. Worldwide, it is the most common carbohydrate in human diets ...
degradation activated by low temperatures in the root, and
gibberellin Gibberellins (GAs) are plant hormones that regulate various developmental processes, including stem elongation, germination, dormancy, flowering, flower development, and leaf and fruit senescence. GAs are one of the longest-known classes of p ...
application bypasses this requirement. Seeds germinate almost solely in bare soil, at temperatures between 10 and 40 °C. While they can germinate in total darkness if proper conditions are present (tests give a 35% germination rate under ideal conditions), in the wild, they in practice only do so when exposed to light, or very close to the soil surface, which explains the plant's habitat preferences. While it can also grow in areas where some vegetation already exists, growth of the rosettes on bare soil is four to seven times more rapid. Seeds germinate in spring and summer. Those that germinate in autumn produce plants that overwinter if they are large enough, while rosettes less than across die in winter. After flowering, the entire plant usually dies at the end of its second year, but some individuals, especially in the northern parts of the range, require a longer growth period and flower in their third year. Under better growing conditions, some individuals flower in the first year. Triennial individuals have been found to produce fewer seeds than biennial and annual ones. While year of flowering and size are linked to the environment, most other characteristics appear to be genetic. A given flower is open only for a single day, opening before dawn and closing in the afternoon. Flowers are self-fecundating and
protogynous Sequential hermaphroditism (called dichogamy in botany) is a type of hermaphroditism In reproductive biology, a hermaphrodite () is an organism that has both kinds of reproductive organs and can produce both gametes associated with male and ...
(with female parts maturing first), and will self-pollinate if they have not been
pollinated Pollination is the transfer of pollen from an anther of a plant to the stigma of a plant, later enabling fertilisation and the production of seeds, most often by an animal or by wind. Pollinating agents can be animals such as insects, birds, ...
by insects during the day. While many insects visit the flowers, only some bees actually accomplish pollination. The flowering period of ''V. thapsus'' lasts from June to August in most of its range, extending to September or October in warmer climates. Visitors include halictid bees and
hoverflies Hover flies, also called flower flies or syrphid flies, make up the insect family Syrphidae. As their common name suggests, they are often seen hovering or nectaring at flowers; the adults of many species feed mainly on nectar and pollen, while ...
. The hair on lower stamens may serve to provide footholds for visitors. The seeds maintain their germinative powers for decades, up to 100 years, according to some studies. Because of this, and because the plant is an extremely prolific seed bearer (each plant produces hundreds of capsules, each containing up to 700 seeds, with a total up to 180,000 or 240,000 seeds), it remains in the
soil seed bank The soil seed bank is the natural storage of seeds, often dormant, within the soil of most ecosystems. The study of soil seed banks started in 1859 when Charles Darwin observed the emergence of seedlings using soil samples from the bottom of a lak ...
for extended periods of time, and can sprout from apparently bare ground, or shortly after
forest fire A wildfire, forest fire, bushfire, wildland fire or rural fire is an unplanned, uncontrolled and unpredictable fire in an area of Combustibility and flammability, combustible vegetation. Depending on the type of vegetation present, a wildfire ...
s long after previous plants have died. Its population pattern typically consists of an ephemeral adult population followed by a long period of dormancy as seeds. Great mullein rarely establishes on new grounds without human intervention because its seeds do not disperse very far. Seed dispersion requires the stem to be moved by wind or animal movement; 75% of the seeds fall within 1 m of the parent plant, and 93% fall within 5 m. Megachilid bees of the genus ''
Anthidium ''Anthidium'' is a genus of bees often called carder or potter bees, who use conifer resin, plant hairs, mud, or a mix of them to build nests. They are in the family Megachilidae which is cosmopolitan in distribution and made up of species that ...
'' use the hair (amongst that of various woolly plants) in making their
nest A nest is a structure built for certain animals to hold eggs or young. Although nests are most closely associated with birds, members of all classes of vertebrates and some invertebrates construct nests. They may be composed of organic materia ...
s. The seeds are generally too small for birds to feed on, although the
American goldfinch The American goldfinch (''Spinus tristis'') is a small North American bird in the finch family. It is migratory, ranging from mid-Alberta to North Carolina during the breeding season, and from just south of the Canada–United States border to ...
has been reported to consume them. Other bird species have been reported to consume the leaves (
Hawaiian goose The nene (''Branta sandvicensis''), also known as the nēnē or the Hawaiian goose, is a species of bird Endemism in birds, endemic to the Hawaiian Islands. The nene is exclusively found in the wild on the islands of Oahu, Maui, Kauai, Kauai, Mo ...
) or flowers (
palila The palila (''Loxioides bailleui'') is a critically endangered finch-billed species of Hawaiian honeycreeper. It has a golden-yellow head and breast, with a light belly, gray back, and greenish wings and tail. The bird has a close ecological rel ...
), or to use the plant as a source when
foraging Foraging is searching for wild food resources. It affects an animal's Fitness (biology), fitness because it plays an important role in an animal's ability to survive and reproduce. Optimal foraging theory, Foraging theory is a branch of behaviora ...
for insects (
white-headed woodpecker The white-headed woodpecker (''Leuconotopicus albolarvatus'') is a non-migratory woodpecker that resides in pine forests of the mountains of western North America. Description It has a black body (approximately long) and white head. It has whit ...
). Additionally, deer and elk eat the leaves.


Fossil record

Seeds of ''V. thapsus'' have been recorded from part of the
Cromer Forest Bed The Cromer Forest Bed is a geological formation in Norfolk, England. It consists of river gravels, estuary and floodplain sediments predominantly clays and muds as well as sands along the coast of northern Norfolk. It is the type locality for the ...
series and at
West Wittering West Wittering is a village and civil parish situated on the Manhood Peninsula in the Chichester district of West Sussex, England. It lies near the mouth of Chichester Harbour on the B2179 road southwest of Chichester close to the border with Ha ...
in
Sussex Sussex (), from the Old English (), is a historic county in South East England that was formerly an independent medieval Anglo-Saxon kingdom. It is bounded to the west by Hampshire, north by Surrey, northeast by Kent, south by the English ...
from some parts of the
Ipswichian interglacial The Eemian (also called the last interglacial, Sangamonian Stage, Ipswichian, Mikulin, Kaydaky, penultimate,NOAA - Penultimate Interglacial Period http://www.ncdc.noaa.gov/global-warming/penultimate-interglacial-period Valdivia or Riss-Würm) wa ...
layers.


Agricultural impacts and control

Because it cannot compete with established plants, great mullein is no longer considered a serious agricultural weed and is easily crowded out in cultivation, Reprinted in Mulligan, G.A. (1979), ''The Biology of Canadian Weeds'' I, , pp. 320–331. except in areas where vegetation is sparse to begin with, such as
California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...
n semidesertic areas of the eastern
Sierra Nevada The Sierra Nevada () is a mountain range in the Western United States, between the Central Valley of California and the Great Basin. The vast majority of the range lies in the state of California, although the Carson Range spur lies primarily ...
. In such ecological contexts, it crowds out native herbs and grasses; its tendency to appear after forest fires also disturbs the normal
ecological succession Ecological succession is the process of change in the species structure of an ecological community over time. The time scale can be decades (for example, after a wildfire) or more or less. Bacteria allows for the cycling of nutrients such as ca ...
. Although not an agricultural threat, its presence can be very difficult to eradicate and is especially problematic in
overgrazed Overgrazing occurs when plants are exposed to intensive grazing for extended periods of time, or without sufficient recovery periods. It can be caused by either livestock in poorly managed agricultural applications, game reserves, or nature rese ...
pastures. The species is legally listed as a
noxious weed A noxious weed, harmful weed or injurious weed is a weed that has been designated by an agricultural or other governing authority as a plant that is injurious to agricultural or horticultural crops, natural habitats or ecosystems, or humans or liv ...
in the US state of
Colorado Colorado (, other variants) is a state in the Mountain West subregion of the Western United States. It encompasses most of the Southern Rocky Mountains, as well as the northeastern portion of the Colorado Plateau and the western edge of t ...
(class C) and Hawaii, and the Australian state of
Victoria Victoria most commonly refers to: * Victoria (Australia), a state of the Commonwealth of Australia * Victoria, British Columbia, provincial capital of British Columbia, Canada * Victoria (mythology), Roman goddess of Victory * Victoria, Seychelle ...
(regionally prohibited in the
West Gippsland West Gippsland, a region of Gippsland in Victoria, Australia, extends from the southeastern limits of metropolitan Melbourne and Western Port Bay in the west to the Latrobe Valley in the east, and is bounded by the Strzelecki Ranges to the south ...
region, and regionally controlled in several others). Despite not being an agricultural weed in itself, it
hosts A host is a person responsible for guests at an event or for providing hospitality during it. Host may also refer to: Places *Host, Pennsylvania, a village in Berks County People *Jim Host (born 1937), American businessman *Michel Host ( ...
a number of insects and
disease A disease is a particular abnormal condition that negatively affects the structure or function of all or part of an organism, and that is not immediately due to any external injury. Diseases are often known to be medical conditions that a ...
s, including both
pests PESTS was an anonymous American activist group formed in 1986 to critique racism, tokenism, and exclusion in the art world. PESTS produced newsletters, posters, and other print material highlighting examples of discrimination in gallery represent ...
and
beneficial insects Beneficial insects (sometimes called beneficial bugs) are any of a number of species of insects that perform valued services like pollination and pest control. The concept of ''beneficial'' is subjective and only arises in light of desired outcome ...
. It is also a potential reservoir of the
cucumber mosaic virus ''Cucumber mosaic virus'' (CMV) is a plant pathogenic virus in the family ''Bromoviridae''. This virus has a worldwide distribution and a very wide host range, having the reputation of the widest host range of any known plant virus. It can be tra ...
, '' Erysiphum cichoraceum'' (the cucurbit
powdery mildew Powdery mildew is a fungal disease that affects a wide range of plants. Powdery mildew diseases are caused by many different species of ascomycete fungi in the order Erysiphales. Powdery mildew is one of the easier plant diseases to identify, as ...
) and
Texas root rot Texas root rot (also known as Phymatotrichopsis root rot, Phymatotrichum root rot, cotton root rot, or, in the older literature, Ozonium root rot) is a disease that is fairly common in Mexico and the southwestern United States resulting in sudden ...
. A study found ''V. thapsus'' hosts insects from 29 different
families Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Ideall ...
. Most of the pests found were
western flower thrips The western flower thrips 'Frankliniella occidentalis'' (Pergande)is an invasive pest insect in agriculture. This species of thrips is native to the Southwestern United States but has spread to other continents, including Europe, Australia (wher ...
(''Frankliniella occidentalis''), ''
Lygus The genus ''Lygus'' includes over 40 species of plant-feeding insects in the family Miridae. The term lygus bug is used for any member of genus ''Lygus''. Species At one time, nearly 200 species were classified as genus ''Lygus'', but most of th ...
'' species such as the
tarnished plant bug The tarnished plant bug (TPB), ''Lygus lineolaris'', is a species of plant-feeding insect in the family Miridae. It has piercing-sucking mouthparts and has become a serious pest on small fruits and vegetables in North America. It is considered a ...
(''L. lineolaris''), and various spider mites from the family
Tetranychidae Spider mites are members of the Tetranychidae family, which includes about 1,200 species. They are part of the subclass Mite, Acari (mites). Spider mites generally live on the undersides of leaf, leaves of plants, where they may spin protective s ...
. These make the plant a potential reservoir for
overwintering Overwintering is the process by which some organisms pass through or wait out the winter season, or pass through that period of the year when "winter" conditions (cold or sub-zero temperatures, ice, snow, limited food supplies) make normal activi ...
pests. Other insects commonly found on great mullein feed exclusively on ''Verbascum'' species in general or ''V. thapsus'' in particular. They include mullein thrips ('' Haplothrips verbasci''), '' Gymnaetron tetrum'' (whose larva consume the seeds), and the
mullein moth The mullein moth (''Cucullia verbasci'') is a noctuid moth with a Palearctic distribution. The species was first described by Carl Linnaeus in his landmark 1758 10th edition of ''Systema Naturae''. Description The forewing is broad (for the ...
(''Cucullia verbasci''). Useful insects are also hosted by great mullein, including predatory mites of the genera '' Galendromus'', '' Typhlodromus'', and '' Amblyseius'', the
minute pirate bug Anthocoridae is a family of bugs, commonly called minute pirate bugs or flower bugs. Worldwide there are 500-600 species. Description Anthocoridae are 1.5–5 mm long and have soft, elongated oval, flat bodies, often patterned in black an ...
''Orius tristicolor'', and the mullein plant bug (''Campylomma verbasci''). The plant's ability to host both pests and beneficials makes it potentially useful to maintain stable populations of insects used for biological control in other cultures, like ''Campylomma verbasci'' and ''
Dicyphus hesperus ''Dicyphus hesperus'' is a species of true bug in the family Miridae. It is a generalist predator of other insects and also feeds on plant tissues. It is native to North America and has been used there in biological control of agricultural pest ...
'' (
Miridae The Miridae are a large and diverse insect family at one time known by the taxonomic synonym Capsidae. Species in the family may be referred to as capsid bugs or "mirid bugs". Common names include plant bugs, leaf bugs, and grass bugs. It is the ...
), a predator of
whiteflies Whiteflies are Hemipterans that typically feed on the undersides of plant leaves. They comprise the family Aleyrodidae, the only family in the superfamily Aleyrodoidea. More than 1550 species have been described. Description and taxonomy The ...
. A number of pest
Lepidoptera Lepidoptera ( ) is an order (biology), order of insects that includes butterfly, butterflies and moths (both are called lepidopterans). About 180,000 species of the Lepidoptera are described, in 126 Family (biology), families and 46 Taxonomic r ...
species, including the stalk borer (''Papaipema nebris'') and
gray hairstreak The gray hairstreak (''Strymon melinus'') is also called the bean lycaenid or cotton square borer. It is a member of the Lycaenidae family, known as the gossamer-winged butterflies and the second-largest family of butterflies. It is one of the mos ...
(''Strymon melinus''), also use ''V. thapsus'' as a host plant. Control of the plant, when desired, is best managed via mechanical means, such as hand pulling and hoeing, preferably followed by sowing of native plants. Animals rarely graze it because of its irritating hairs, and liquid
herbicide Herbicides (, ), also commonly known as weedkillers, are substances used to control undesired plants, also known as weeds.EPA. February 201Pesticides Industry. Sales and Usage 2006 and 2007: Market Estimates. Summary in press releasMain page fo ...
s require
surfactant Surfactants are chemical compounds that decrease the surface tension between two liquids, between a gas and a liquid, or interfacial tension between a liquid and a solid. Surfactants may act as detergents, wetting agents, emulsifiers, foaming ...
s to be effective, as the hair causes water to roll off the plant, much like the
lotus effect The lotus effect refers to self-cleaning properties that are a result of ultrahydrophobicity as exhibited by the leaves of ''Nelumbo'', the lotus flower. Dirt particles are picked up by water droplets due to the micro- and nanoscopic architect ...
. Burning is ineffective, as it only creates new bare areas for
seedling A seedling is a young sporophyte developing out of a plant embryo from a seed. Seedling development starts with germination of the seed. A typical young seedling consists of three main parts: the radicle (embryonic root), the hypocotyl (embry ...
s to occupy. ''G. tetrum'' and ''Cucullia verbasci'' usually have little effect on ''V. thapsus'' populations as a whole. Goats and chickens have also been proposed to control mullein. Effective (when used with a surfactant) contact herbicides include
glyphosate Glyphosate (IUPAC name: ''N''-(phosphonomethyl)glycine) is a broad-spectrum Herbicide, systemic herbicide and Crop desiccation, crop desiccant. It is an organophosphorus compound, specifically a phosphonate, which acts by inhibiting the plan ...
,
triclopyr Triclopyr (3,5,6-trichloro-2-pyridinyloxyacetic acid) is an organic compound in the pyridine group that is used as a systemic foliar herbicide and fungicide. Uses Triclopyr is used to control broadleaf weeds while leaving grasses and conifers ...
and sulfurometuron-methyl. Ground herbicides, like
tebuthiuron Tebuthiuron is a nonselective broad spectrum herbicide of the urea class. It is used in a number of herbicides manufactured by Dow AgroSciences, and is sold under several trade names, depending on the formulation. It is used to control weeds, w ...
, are also effective, but recreate bare ground and require repeated application to prevent regrowth.


Uses


Phytochemicals

Phytochemical Phytochemicals are chemical compounds produced by plants, generally to help them resist fungi, bacteria and plant virus infections, and also consumption by insects and other animals. The name comes . Some phytochemicals have been used as poisons ...
s in ''V. thapsus'' flowers and leaves include
saponin Saponins (Latin "sapon", soap + "-in", one of), also selectively referred to as triterpene glycosides, are bitter-tasting usually toxic plant-derived organic chemicals that have a foamy quality when agitated in water. They are widely distributed ...
s,
polysaccharide Polysaccharides (), or polycarbohydrates, are the most abundant carbohydrates found in food. They are long chain polymeric carbohydrates composed of monosaccharide units bound together by glycosidic linkages. This carbohydrate can react with wa ...
s,
mucilage Mucilage is a thick, gluey substance produced by nearly all plants and some microorganisms. These microorganisms include protists which use it for their locomotion. The direction of their movement is always opposite to that of the secretion of m ...
,
flavonoid Flavonoids (or bioflavonoids; from the Latin word ''flavus'', meaning yellow, their color in nature) are a class of polyphenolic secondary metabolites found in plants, and thus commonly consumed in the diets of humans. Chemically, flavonoids ...
s,
tannin Tannins (or tannoids) are a class of astringent, polyphenolic biomolecules that bind to and precipitate proteins and various other organic compounds including amino acids and alkaloids. The term ''tannin'' (from Anglo-Norman ''tanner'', ...
s,
iridoid Iridoids are a type of monoterpenoids in the general form of cyclopentanopyran, found in a wide variety of plants and some animals. They are biosynthetically derived from 8-oxogeranial. Iridoids are typically found in plants as glycosides, mos ...
and
lignin Lignin is a class of complex organic polymers that form key structural materials in the support tissues of most plants. Lignins are particularly important in the formation of cell walls, especially in wood and bark, because they lend rigidity ...
glycoside In chemistry, a glycoside is a molecule in which a sugar is bound to another functional group via a glycosidic bond. Glycosides play numerous important roles in living organisms. Many plants store chemicals in the form of inactive glycosides. ...
s, and
essential oil An essential oil is a concentrated hydrophobic liquid containing volatile (easily evaporated at normal temperatures) chemical compounds from plants. Essential oils are also known as volatile oils, ethereal oils, aetheroleum, or simply as the o ...
s. The plant's leaves, in addition to the seeds, have been reported to contain
rotenone Rotenone is an odorless, colorless, crystalline isoflavone used as a broad-spectrum insecticide, piscicide, and pesticide. It occurs naturally in the seeds and stems of several plants, such as the jicama vine plant, and the roots of several member ...
, although quantities are unknown.


Traditional medicine

Although long used in
herbal medicine Herbal medicine (also herbalism) is the study of pharmacognosy and the use of medicinal plants, which are a basis of traditional medicine. With worldwide research into pharmacology, some herbal medicines have been translated into modern remed ...
, no
drugs A drug is any chemical substance that causes a change in an organism's physiology or psychology when consumed. Drugs are typically distinguished from food and substances that provide nutritional support. Consumption of drugs can be via inhalat ...
are manufactured from its components.
Dioscorides Pedanius Dioscorides ( grc-gre, Πεδάνιος Διοσκουρίδης, ; 40–90 AD), “the father of pharmacognosy”, was a Greek physician, pharmacologist, botanist, and author of ''De materia medica'' (, On Medical Material) —a 5-vol ...
first recommended the plant 2000 years ago, considering it useful as a
folk medicine Traditional medicine (also known as indigenous medicine or folk medicine) comprises medical aspects of traditional knowledge that developed over generations within the folk beliefs of various societies, including indigenous peoples, before the ...
for pulmonary diseases. Leaves were smoked to attempt to treat lung ailments, a tradition that in America was rapidly transmitted to Native American peoples. The
Zuni people The Zuni ( zun, A:shiwi; formerly spelled ''Zuñi'') are Native American Pueblo peoples native to the Zuni River valley. The Zuni are a Federally recognized tribe and most live in the Pueblo of Zuni on the Zuni River, a tributary of the Lit ...
, however, use the plant in
poultice A poultice, also called a cataplasm, is a soft moist mass, often heated and medicated, that is spread on cloth and placed over the skin to treat an aching, inflamed, or painful part of the body. It can be used on wounds, such as cuts. 'Poultice' ...
s of powdered root applied to
sore Sore may refer to: * Ulcer (dermatology), a sore on the skin or a mucous membrane * Sore, a mild pain or ache * ''Sore'' (album), by Buzzov*en * Sore (band), an Indonesian rock band * Sore, Landes, a village in the Landes département of France * ...
s,
rash A rash is a change of the human skin which affects its color, appearance, or texture. A rash may be localized in one part of the body, or affect all the skin. Rashes may cause the skin to change color, itch, become warm, bumpy, chapped, dry, cr ...
es, and skin infections. An infusion of the root is also used to treat
athlete's foot Athlete's foot, known medically as ''tinea pedis'', is a common skin infection of the feet caused by a fungus. Signs and symptoms often include itching, scaling, cracking and redness. In rare cases the skin may blister. Athlete's foot fungus m ...
. All preparations meant to be drunk have to be finely filtered to eliminate the irritating hairs. Oil from the flowers was used against
catarrh Catarrh is an exudate of inflamed mucous membranes in one of the airways or cavities of the body, usually with reference to the throat and paranasal sinuses. It can result in a thick exudate of mucus and white blood cells caused by the swelling o ...
s,
colic Colic or cholic () is a form of pain that starts and stops abruptly. It occurs due to muscular contractions of a hollow tube ( small and large intestine, gall bladder, ureter, etc.) in an attempt to relieve an obstruction by forcing content out ...
s,
earache Ear pain, also known as earache or otalgia, is pain in the ear. Primary ear pain is pain that originates from the ear. Secondary ear pain is a type of referred pain, meaning that the source of the pain differs from the location where the pain i ...
s,
frostbite Frostbite is a skin injury that occurs when exposed to extreme low temperatures, causing the freezing of the skin or other tissues, commonly affecting the fingers, toes, nose, ears, cheeks and chin areas. Most often, frostbite occurs in the han ...
,
eczema Dermatitis is inflammation of the Human skin, skin, typically characterized by itchiness, erythema, redness and a rash. In cases of short duration, there may be small blisters, while in long-term cases the skin may become lichenification, thick ...
, and other external conditions. Topical application of various ''V. thapsus''-based preparations was recommended for the treatment of
wart Warts are typically small, rough, hard growths that are similar in color to the rest of the skin. They typically do not result in other symptoms, except when on the bottom of the feet, where they may be painful. While they usually occur on the ...
s,
boil A boil, also called a furuncle, is a deep folliculitis, which is an infection of the hair follicle. It is most commonly caused by infection by the bacterium ''Staphylococcus aureus'', resulting in a painful swollen area on the skin caused by an ...
s,
carbuncle A carbuncle is a cluster of boils caused by bacterial infection, most commonly with ''Staphylococcus aureus'' or ''Streptococcus pyogenes''. The presence of a carbuncle is a sign that the immune system is active and fighting the infection. The ...
s, hemorrhoids, and
chilblain Chilblains, also known as pernio, is a medical condition in which damage occurs to capillary beds in the skin, most often in the hands or feet, when blood perfuses into the nearby tissue resulting in redness, itching, inflammation, and possibly ...
s, amongst others.
Glycyrrhizin Glycyrrhizin (or glycyrrhizic acid or glycyrrhizinic acid) is the chief sweet-tasting constituent of ''Glycyrrhiza glabra'' ( liquorice) root. Structurally, it is a saponin used as an emulsifier and gel-forming agent in foodstuffs and cosmet ...
compounds with
bactericide A bactericide or bacteriocide, sometimes abbreviated Bcidal, is a substance which kills bacteria. Bactericides are disinfectants, antiseptics, or antibiotics. However, material surfaces can also have bactericidal properties based solely on thei ...
effects in vitro were isolated from flowers. The German
Commission E The German ''Commission E'' is a scientific advisory board of the Federal Institute for Drugs and Medical Devices formed in 1978. The commission gives scientific expertise for the approval of substances and products previously used in traditional, ...
describes uses of the plant for respiratory infections. It was also part of the
National Formulary A formulary is a list of pharmaceutical drugs, often decided upon by a group of people, for various reasons such as insurance coverage or use at a medical facility. Traditionally, a formulary contained a collection of formulas for the compounding ...
in the United States and United Kingdom. The plant has been used in an attempt to treat colds,
croup Croup, also known as laryngotracheobronchitis, is a type of respiratory infection that is usually caused by a virus. The infection leads to swelling inside the trachea, which interferes with normal breathing and produces the classic symptoms o ...
, sunburn, and other skin irritations.


Other uses

Roman soldiers are said to have dipped the plant stalks in grease for use as torches. Other cultures use the leaves as wicks. Native Americans and American colonists lined their shoes with leaves from the plant to keep out the cold. Mullein may be cultivated as an
ornamental plant Ornamental plants or garden plants are plants that are primarily grown for their beauty but also for qualities such as scent or how they shape physical space. Many flowering plants and garden varieties tend to be specially bred cultivars that i ...
. As for many plants, (
Pliny the Elder Gaius Plinius Secundus (AD 23/2479), called Pliny the Elder (), was a Roman author, naturalist and natural philosopher, and naval and army commander of the early Roman Empire, and a friend of the emperor Vespasian. He wrote the encyclopedic '' ...
described it in his ''
Naturalis Historia The ''Natural History'' ( la, Naturalis historia) is a work by Pliny the Elder. The largest single work to have survived from the Roman Empire to the modern day, the ''Natural History'' compiles information gleaned from other ancient authors. ...
''), great mullein was linked to
witch Witchcraft traditionally means the use of Magic (supernatural), magic or supernatural powers to harm others. A practitioner is a witch. In Middle Ages, medieval and early modern Europe, where the term originated, accused witches were usually ...
es, although the relationship remained generally ambiguous, and the plant was also widely held to ward off
curse A curse (also called an imprecation, malediction, execration, malison, anathema, or commination) is any expressed wish that some form of adversity or misfortune will befall or attach to one or more persons, a place, or an object. In particular, ...
s and
evil spirit Evil, in a general sense, is defined as the opposite or absence of good. It can be an extremely broad concept, although in everyday usage it is often more narrowly used to talk about profound wickedness and against common good. It is generally ...
s. The seeds contain several compounds (
saponin Saponins (Latin "sapon", soap + "-in", one of), also selectively referred to as triterpene glycosides, are bitter-tasting usually toxic plant-derived organic chemicals that have a foamy quality when agitated in water. They are widely distributed ...
s,
glycoside In chemistry, a glycoside is a molecule in which a sugar is bound to another functional group via a glycosidic bond. Glycosides play numerous important roles in living organisms. Many plants store chemicals in the form of inactive glycosides. ...
s,
coumarin Coumarin () or 2''H''-chromen-2-one is an aromatic organic chemical compound with formula . Its molecule can be described as a benzene molecule with two adjacent hydrogen atoms replaced by a lactone-like chain , forming a second six-membered h ...
,
rotenone Rotenone is an odorless, colorless, crystalline isoflavone used as a broad-spectrum insecticide, piscicide, and pesticide. It occurs naturally in the seeds and stems of several plants, such as the jicama vine plant, and the roots of several member ...
) that are toxic to fish, and have been widely used as
piscicide A piscicide is a chemical substance which is poisonous to fish. The primary use for piscicides is to eliminate a dominant species of fish in a body of water, as the first step in attempting to populate the body of water with a different fish. They ...
for
fishing Fishing is the activity of trying to catch fish. Fish are often caught as wildlife from the natural environment, but may also be caught from stocked bodies of water such as ponds, canals, park wetlands and reservoirs. Fishing techniques inclu ...
. Due to its weedy capacities, the plant, unlike other species of the genus (such as '' V. phoeniceum''), is not often cultivated.


Notes


References


Further reading

*


External links


Verbascum.org

The type specimen of ''Verbascum thapsus''




from the
UBC The University of British Columbia (UBC) is a public research university with campuses near Vancouver and in Kelowna, British Columbia. Established in 1908, it is British Columbia's oldest university. The university ranks among the top three ...
collection
JLindquist.com: webpage with pictures of tall specimens
{{Taxonbar, from=Q160095
thapsus Thapsus, also known as Tampsus and as Thapsus Minor to distinguish it from Thapsus in Sicily, was a Carthaginian and Roman port near present-day Bekalta, Tunisia. Geography Thapsus was established on Ras ed-Dimas, an easily defended promontory ...
Flora of Europe Flora of North Africa Flora of temperate Asia Flora of Spain Demulcents Medicinal plants Plants used in traditional Native American medicine Plants described in 1753 Taxa named by Carl Linnaeus Introduced plants of South America Flora naturalised in Australia