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''Lama'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
containing four South American
camelid Camelids are members of the biological family Camelidae, the only currently living family in the suborder Tylopoda. The seven extant members of this group are: dromedary camels, Bactrian camels, wild Bactrian camels, llamas, alpacas, vicuñas, ...
s, the wild
guanaco The guanaco (; ''Lama guanicoe'') is a camelid native to South America, closely related to the llama. Guanacos are one of two wild South American camelids, the other being the vicuña, which lives at higher elevations. Etymology The guanaco g ...
and
vicuña The vicuña (''Lama vicugna'') or vicuna (both , very rarely spelled ''vicugna'', its former genus name) is one of the two wild South American camelids, which live in the high alpine areas of the Andes, the other being the guanaco, which live ...
, and the domesticated
llama The llama (; ) (''Lama glama'') is a domesticated South American camelid, widely used as a List of meat animals, meat and pack animal by Inca empire, Andean cultures since the Pre-Columbian era. Llamas are social animals and live with othe ...
and
alpaca The alpaca (''Lama pacos'') is a species of South American camelid mammal. It is similar to, and often confused with, the llama. However, alpacas are often noticeably smaller than llamas. The two animals are closely related and can successfu ...
. Before the
Spanish Spanish might refer to: * Items from or related to Spain: **Spaniards are a nation and ethnic group indigenous to Spain **Spanish language, spoken in Spain and many Latin American countries **Spanish cuisine Other places * Spanish, Ontario, Cana ...
conquest of the
Americas The Americas, which are sometimes collectively called America, are a landmass comprising the totality of North and South America. The Americas make up most of the land in Earth's Western Hemisphere and comprise the New World. Along with th ...
, llamas and alpacas were the only domesticated
ungulate Ungulates ( ) are members of the diverse clade Ungulata which primarily consists of large mammals with hooves. These include odd-toed ungulates such as horses, rhinoceroses, and tapirs; and even-toed ungulates such as cattle, pigs, giraffes, cam ...
s of the
continent A continent is any of several large landmasses. Generally identified by convention rather than any strict criteria, up to seven geographical regions are commonly regarded as continents. Ordered from largest in area to smallest, these seven ...
. They were kept not only for their value as beasts of burden, but also for their flesh, hides, and
wool Wool is the textile fibre obtained from sheep and other mammals, especially goats, rabbits, and camelids. The term may also refer to inorganic materials, such as mineral wool and glass wool, that have properties similar to animal wool. As ...
.


Classification

Although they were often compared to sheep by early writers, their affinity to the camel was soon perceived. They were included in the genus ''
Camelus A camel (from: la, camelus and grc-gre, κάμηλος (''kamēlos'') from Hebrew or Phoenician: גָמָל ''gāmāl''.) is an even-toed ungulate in the genus ''Camelus'' that bears distinctive fatty deposits known as "humps" on its back. C ...
'' in the ''Systema Naturae'' of
Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
. In 1800,
Cuvier Jean Léopold Nicolas Frédéric, Baron Cuvier (; 23 August 1769 – 13 May 1832), known as Georges Cuvier, was a French naturalist and zoologist, sometimes referred to as the "founding father of paleontology". Cuvier was a major figure in nat ...
moved the llama, alpaca, and guanaco to the genus ''Lama'', and the vicuña to the genus ''Vicugna''. Later, the alpaca was transferred to ''Vicugna''; both were eventually returned to ''Lama'' by the
American Society of Mammalogists The American Society of Mammalogists (ASM) was founded in 1919. Its primary purpose is to encourage the study of mammals, and professions studying them. There are over 4,500 members of this society, and they are primarily professional scientists ...
in 2021. These camelids are, with the two species of true
camel A camel (from: la, camelus and grc-gre, κάμηλος (''kamēlos'') from Hebrew or Phoenician: גָמָל ''gāmāl''.) is an even-toed ungulate in the genus ''Camelus'' that bears distinctive fatty deposits known as "humps" on its back. C ...
s, the sole
extant Extant is the opposite of the word extinct. It may refer to: * Extant hereditary titles * Extant literature, surviving literature, such as ''Beowulf'', the oldest extant manuscript written in English * Extant taxon, a taxon which is not extinct, ...
representatives of a distinct section of
Artiodactyla The even-toed ungulates (Artiodactyla , ) are ungulates—hoofed animals—which bear weight equally on two (an even number) of their five toes: the third and fourth. The other three toes are either present, absent, vestigial, or pointing poster ...
(even-toed ungulates) called
Tylopoda Tylopoda (meaning "calloused foot") is a suborder of terrestrial herbivorous even-toed ungulates belonging to the order Artiodactyla. They are found in the wild in their native ranges of South America and Asia, while Australian feral camels are ...
, or "hump-footed", from the peculiar bumps on the soles of their feet. This section consists of a single family, the
Camelidae Camelids are members of the biological family Camelidae, the only currently living family in the suborder Tylopoda. The seven extant members of this group are: dromedary camels, Bactrian camels, wild Bactrian camels, llamas, alpacas, vicuñas, ...
, the other sections of the same great division being the
Suina Suina (also known as Suiformes) is a suborder of omnivorous, non-ruminant artiodactyl mammals that includes the domestic pig and peccaries. A member of this clade is known as a suine. Suina includes the family Suidae, termed suids, known in Eng ...
or
pig The pig (''Sus domesticus''), often called swine, hog, or domestic pig when distinguishing from other members of the genus '' Sus'', is an omnivorous, domesticated, even-toed, hoofed mammal. It is variously considered a subspecies of ''Sus s ...
s, the Tragulina or
chevrotain Chevrotains, or mouse-deer, are small even-toed ungulates that make up the family Tragulidae, the only extant members of the infraorder Tragulina. The 10  extant species are placed in three genera, but several species also are known only ...
s, and the Pecora or true
ruminants Ruminants ( suborder Ruminantia) are hoofed herbivorous grazing or browsing mammals that are able to acquire nutrients from plant-based food by fermenting it in a specialized stomach prior to digestion, principally through microbial actions. The ...
, to each of which the Tylopoda have some affinity, standing in some respects in a central position between them, sharing some characters from each, but showing special modifications not found in any of the others. Discovery of the
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
fauna of the American continent of the Paleogene and Neogene periods, starting with the 19th-century paleontologists Leidy,
Cope The cope (known in Latin as ''pluviale'' 'rain coat' or ''cappa'' 'cape') is a liturgical vestment, more precisely a long mantle or cloak, open in front and fastened at the breast with a band or clasp. It may be of any liturgical colours, litu ...
, and
Marsh A marsh is a wetland that is dominated by herbaceous rather than woody plant species.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p Marshes can often be found at ...
, has revealed the early history of this family. Llamas were not always confined to South America; their remains are abundant in the
Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fina ...
deposits of the
Rocky Mountains The Rocky Mountains, also known as the Rockies, are a major mountain range and the largest mountain system in North America. The Rocky Mountains stretch in straight-line distance from the northernmost part of western Canada, to New Mexico in ...
region, and in
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
; some of these extinct forms were much larger than any now living. None of these transitional forms has been found in Old World strata;
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
was the original home of the Tylopoda. Camelids invaded the Old World via
Beringia Beringia is defined today as the land and maritime area bounded on the west by the Lena River in Russia; on the east by the Mackenzie River in Canada; on the north by 72 degrees north latitude in the Chukchi Sea; and on the south by the tip ...
, and South America via the
Isthmus of Panama The Isthmus of Panama ( es, Istmo de Panamá), also historically known as the Isthmus of Darien (), is the narrow strip of land that lies between the Caribbean Sea and the Pacific Ocean, linking North and South America. It contains the country ...
, as part of the
Great American Interchange The Great American Biotic Interchange (commonly abbreviated as GABI), also known as the Great American Interchange and the Great American Faunal Interchange, was an important late Cenozoic paleozoogeographic biotic interchange event in which lan ...
. The Old World forms were gradually driven southward, perhaps by changes of
climate Climate is the long-term weather pattern in an area, typically averaged over 30 years. More rigorously, it is the mean and variability of meteorological variables over a time spanning from months to millions of years. Some of the meteorologic ...
, and having become isolated, they have undergone further special modifications. Meanwhile, the New World members of the family became restricted to
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
following the peopling of the Americas by Paleo-Indians and the accompanying extinction of the
megafauna In terrestrial zoology, the megafauna (from Greek μέγας ''megas'' "large" and New Latin ''fauna'' "animal life") comprises the large or giant animals of an area, habitat, or geological period, extinct and/or extant. The most common threshold ...
. A possible variety is the hueque or
chilihueque The chilihueque or hueque was a South American camelid variety or species that existed in central and south-central Chile in Pre-Hispanic and colonial times. There are two main hypotheses on their status among South American camelids: the firs ...
that existed in
central Central is an adjective usually referring to being in the center of some place or (mathematical) object. Central may also refer to: Directions and generalised locations * Central Africa, a region in the centre of Africa continent, also known as ...
and south-central Chile in
pre-Hispanic In the history of the Americas, the pre-Columbian era spans from the original settlement of North and South America in the Upper Paleolithic period through European colonization, which began with Christopher Columbus's voyage of 1492. Usually, th ...
and early colonial times. Two main hypotheses on their status among South American camelids are given: the first one suggests they are locally domesticated
guanacos The guanaco (; ''Lama guanicoe'') is a camelid native to South America, closely related to the llama. Guanacos are one of two wild South American camelids, the other being the vicuña, which lives at higher elevations. Etymology The guanaco g ...
and the second suggests they are a variety of
llama The llama (; ) (''Lama glama'') is a domesticated South American camelid, widely used as a List of meat animals, meat and pack animal by Inca empire, Andean cultures since the Pre-Columbian era. Llamas are social animals and live with othe ...
s brought from the north into south-central Chile.Bonacic, Cristián (1991)
Características biológicas y productivas de los camélidos sudamericanos
. ''Avances en ciencias veterinarias''. Vol. 6, No. 2.
Chilihueques became extinct in the 16th or 17th century, being replaced by European livestock. The causes of its extinction are unknown. According to
Juan Ignacio Molina Fr. Juan Ignacio Molina (; (June 24, 1740 – September 12, 1829) was a Chilean Jesuit priest, naturalist, historian, translator, geographer, botanist, ornithologist, and linguist. He is usually referred to as Abate Molina (a form of Abbot Moli ...
, the Dutch captain
Joris van Spilbergen Joris van Spilbergen (1568 in Antwerp – January 31, 1620 in Bergen op Zoom) was a Dutch naval officer. Joris van Spilbergen was born in Antwerp in 1568. His first major expedition was in 1596, when he sailed to Africa. He then left for As ...
observed the use of chilihueques by native
Mapuche The Mapuche ( (Mapuche & Spanish: )) are a group of indigenous inhabitants of south-central Chile and southwestern Argentina, including parts of Patagonia. The collective term refers to a wide-ranging ethnicity composed of various groups who sha ...
s of
Mocha Island Mocha Island ( es, link=no, Isla Mocha ) is a small Chilean island located west of the coast of Arauco Province in the Pacific Ocean. The island is approximately in area, with a small chain of mountains running roughly in north-south direction. ...
as plough animals in 1614.''The Geographical, Natural and Civil History of Chili''
Pages 15 and 16, Volume II
/ref>


Extant species


Characteristics

These characters apply especially to llamas.
Dentition Dentition pertains to the development of teeth and their arrangement in the mouth. In particular, it is the characteristic arrangement, kind, and number of teeth in a given species at a given age. That is, the number, type, and morpho-physiolo ...
of adults:-incisors 1/3 canines 1/1, premolars 2/2, molars 3/2; total 32. In the upper jaw is a compressed, sharp, pointed laniariform
incisor Incisors (from Latin ''incidere'', "to cut") are the front teeth present in most mammals. They are located in the premaxilla above and on the mandible below. Humans have a total of eight (two on each side, top and bottom). Opossums have 18, whe ...
near the hinder edge of the
premaxilla The premaxilla (or praemaxilla) is one of a pair of small cranial bones at the very tip of the upper jaw of many animals, usually, but not always, bearing teeth. In humans, they are fused with the maxilla. The "premaxilla" of therian mammal has b ...
, followed in the male at least by a moderate-sized, pointed, curved true canine in the anterior part of the maxilla. The isolated canine-like
premolar The premolars, also called premolar teeth, or bicuspids, are transitional teeth located between the canine and molar teeth. In humans, there are two premolars per quadrant in the permanent set of teeth, making eight premolars total in the mouth ...
which follows in the camels is not present. The teeth of the molar series which are in contact with each other consist of two very small premolars (the first almost rudimentary) and three broad
molars The molars or molar teeth are large, flat teeth at the back of the mouth. They are more developed in mammals. They are used primarily to grind food during chewing. The name ''molar'' derives from Latin, ''molaris dens'', meaning "millstone to ...
, constructed generally like those of ''Camelus''. In the lower jaw, the three incisors are long, spatulate, and procumbent; the outer ones are the smallest. Next to these is a curved, suberect canine, followed after an interval by an isolated minute and often deciduous simple conical premolar; then a contiguous series of one premolar and three molars, which differ from those of ''Camelus'' in having a small accessory column at the anterior outer edge. The skull generally resembles that of ''Camelus'', the relatively larger brain cavity and orbits and less developed
cranial ridges The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, th ...
being due to its smaller size. The nasal bones are shorter and broader, and are joined by the
premaxilla The premaxilla (or praemaxilla) is one of a pair of small cranial bones at the very tip of the upper jaw of many animals, usually, but not always, bearing teeth. In humans, they are fused with the maxilla. The "premaxilla" of therian mammal has b ...
.
Vertebrae The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic i ...
: * cervical 7, * dorsal 12, * lumbar 7, * sacral 4, * caudal 15 to 20. Ears are rather long and pointed. No dorsal hump is present. Feet are narrow, the toes being more separated than in the camels, each having a distinct plantar pad. The tail is short, and the fur is long and woolly. The llama and alpaca are only known in the domestic state, and are variable in size and color, being often white, black, or piebald. The wild guanaco and vicuña are of a nearly uniform light-brown colour, passing into white below. The vicuña and guanaco share an obvious family resemblance and may be difficult to tell apart at a distance. The vicuña is smaller and slenderer in its proportions, and has a shorter head than the guanaco. The guanaco has an extensive geographical range, from the high lands of the Andean region of Ecuador and Peru to the open plains of
Patagonia Patagonia () refers to a geographical region that encompasses the southern end of South America, governed by Argentina and Chile. The region comprises the southern section of the Andes Mountains with lakes, fjords, temperate rainforests, and gl ...
, and even the wooded islands of
Tierra del Fuego Tierra del Fuego (, ; Spanish for "Land of the Fire", rarely also Fireland in English) is an archipelago off the southernmost tip of the South American mainland, across the Strait of Magellan. The archipelago consists of the main island, Isla G ...
. It constituted the principal food of the Patagonian Indians, and they use its skin for the material from which their long robes are made. It is about the size of a
European red deer The red deer (''Cervus elaphus'') is one of the largest deer species. A male red deer is called a stag or hart, and a female is called a hind. The red deer inhabits most of Europe, the Caucasus Mountains region, Anatolia, Iran, and parts of wes ...
, and is an elegant animal with a long, slender, gracefully curved neck and slim legs. The vicuña ranges throughout the Western Andes.


References

{{Authority control Camelids Mammal genera Taxa named by Georges Cuvier