Venenosaurus
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''Venenosaurus'' ( ) is a genus of
sauropod Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their bo ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
that lived in what is now Utah during the Early Cretaceous. Its type and only species is ''Venenosaurus dicrocei''. Fossils of ''Venenosaurus'' were first discovered in 1998, by Denver Museum of Natural History volunteer Anthony DiCroce, and described as a new genus and species in 2001 by Virginia Tidwell and colleagues, who named the species for DiCroce. ''Venenosaurus'' was a relatively small sauropod, and was similar to ''
Cedarosaurus ''Cedarosaurus'' (meaning "Cedar lizard" - named after the Cedar Mountain Formation, in which it was discovered) was a nasal-crested macronarian dinosaur genus from the Early Cretaceous Period (Valanginian). It was a sauropod which lived in what i ...
'', another sauropod from the Early Cretaceous of Utah.


Discovery and naming

In the spring of 1998, a team of volunteers for the
Denver Museum of Natural History The Denver Museum of Nature & Science is a municipal natural history and science museum in Denver, Colorado. It is a resource for informal science education in the Rocky Mountain region. A variety of exhibitions, programs, and activities help mus ...
, including Anthony "Tony" DiCroce, found a fossil site in Grand County, Utah, which they named Tony's Bone Bed. The bone bed was in the Poison Strip Member of the Cedar Mountain Formation, and contained the remains of the iguanodont ''
Planicoxa ''Planicoxa'' is a genus of advanced iguanodontian dinosaur from the Early Cretaceous of North America. It is known from the partial skeletons of several individual specimens. Its fossils were discovered in Utah, United States. The type species ...
'', a theropod, and an adult and juvenile of a sauropod. In 2001, Virginia Tidwell, Kenneth Carpenter, and Susanne Meyer described the sauropod remains as a new genus and species, ''Venenosaurus dicrocei''. The genus name comes from the Latin ''veneno'', "poison", in reference to the Poison Strip Member, and the species name honors DiCroce for his discovery of the fossils.


Fossil record

''Venenosaurus'' fossils are known from the Poison Strip Member of the Cedar Mountain Formation, including the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
, DMNS 40932, a partial skeleton of an adult individual, which consists of nine tail
vertebra The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic ...
e, the left
scapula The scapula (plural scapulae or scapulas), also known as the shoulder blade, is the bone that connects the humerus (upper arm bone) with the clavicle (collar bone). Like their connected bones, the scapulae are paired, with each scapula on eithe ...
, right
radius In classical geometry, a radius ( : radii) of a circle or sphere is any of the line segments from its center to its perimeter, and in more modern usage, it is also their length. The name comes from the latin ''radius'', meaning ray but also the ...
, left
ulna The ulna (''pl''. ulnae or ulnas) is a long bone found in the forearm that stretches from the elbow to the smallest finger, and when in anatomical position, is found on the medial side of the forearm. That is, the ulna is on the same side of t ...
, metacarpals, forefoot phalanges, right pubis, left and right
ischia Ischia ( , , ) is a volcanic island in the Tyrrhenian Sea. It lies at the northern end of the Gulf of Naples, about from Naples. It is the largest of the Phlegrean Islands. Roughly trapezoidal in shape, it measures approximately east to west ...
, metatarsals,
chevrons Chevron (often relating to V-shaped patterns) may refer to: Science and technology * Chevron (aerospace), sawtooth patterns on some jet engines * Chevron (anatomy), a bone * ''Eulithis testata'', a moth * Chevron (geology), a fold in rock lay ...
, and
rib In vertebrate anatomy, ribs ( la, costae) are the long curved bones which form the rib cage, part of the axial skeleton. In most tetrapods, ribs surround the chest, enabling the lungs to expand and thus facilitate breathing by expanding the ches ...
s. A few bones of a juvenile individual are also known from the same site. These specimens are housed at the Denver Museum of Nature and Science. Additional specimens of ''Venenosaurus'' have been reported from Dalton Wells Quarry, in the Yellow Cat Member of the Cedar Mountain Formation, comprising the fragmentary remains of least three individuals. However, it is not certain whether these specimens belong to ''Venenosaurus'' or its close relative ''Cedarosaurus''.


Anatomy

The scientists who first described ''V. dicrocei'' observed that the new species most closely resembles ''
Cedarosaurus ''Cedarosaurus'' (meaning "Cedar lizard" - named after the Cedar Mountain Formation, in which it was discovered) was a nasal-crested macronarian dinosaur genus from the Early Cretaceous Period (Valanginian). It was a sauropod which lived in what i ...
'', while still being distinct. ''Venenosaurus'' is a relatively small titanosauriform sauropod, measuring long and weighing .


Limbs

The
radius In classical geometry, a radius ( : radii) of a circle or sphere is any of the line segments from its center to its perimeter, and in more modern usage, it is also their length. The name comes from the latin ''radius'', meaning ray but also the ...
of ''Venenosaurus'' is more slender than the radii of ''
Alamosaurus ''Alamosaurus'' (; meaning "Ojo Alamo lizard") is a genus of opisthocoelicaudiine titanosaurian sauropod dinosaurs, containing a single known species, ''Alamosaurus sanjuanensis'', from the late Cretaceous Period of what is now southern North Am ...
'', '' Chubutisaurus'', '' Opisthocoelicaudia'', and ''Saltasaurus''. The ratio of the radius' least circumference to length produces a ratio of .33, more gracile than the radius of ''
Camarasaurus lewisi ''Camarasaurus lewisi'' is a species of sauropod dinosaur from the Upper Jurassic of the United States. It was named by James A. Jensen in 1988. ''C. lewisi'' was originally placed in its own genus, ''Cathetosaurus'', but in 1996 it was reclassif ...
'' and '' C. grandis''. ''Cedarosaurus'', however, has a slightly more gracile ratio of .31. The team found that '' Brachiosaurus brancai'''s radius is the closest anatomical match to that of ''Venenosaurus''. The metacarpals of ''Venenosaurus'' are long and slender. With the exception of the incomplete first metacarpal, all of the right metacarpals are known. Metatarsal I is the shortest and most robust of the three recovered metatarsals. ''Cedarosaurus'' had a more gracile
ulna The ulna (''pl''. ulnae or ulnas) is a long bone found in the forearm that stretches from the elbow to the smallest finger, and when in anatomical position, is found on the medial side of the forearm. That is, the ulna is on the same side of t ...
and radius than ''Venenosaurus''.


Pelvis

''Venenosaurus'' shows a mixture of titanosaur and non-titanosaur
ischium The ischium () form ...
-to- pubis proportions. Its hips most closely resemble those of ''Brachiosaurus''.


Vertebrae

The vertebrae in the middle and toward the end of the tail were short, distinguishing it from
titanosaurs Titanosaurs (or titanosaurians; members of the group Titanosauria) were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still thr ...
like '' Andesaurus'', '' Malawisaurus'', ''
Aeolosaurus ''Aeolosaurus'' (; "Aeolus' lizard") is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous Period of what is now South America. Like most sauropods, it would have been a quadrupedal herbivore with a long neck and tail. ''Aeolosau ...
'', ''
Alamosaurus ''Alamosaurus'' (; meaning "Ojo Alamo lizard") is a genus of opisthocoelicaudiine titanosaurian sauropod dinosaurs, containing a single known species, ''Alamosaurus sanjuanensis'', from the late Cretaceous Period of what is now southern North Am ...
'', and '' Saltasaurus''. The neural spines in the middle tail vertebrae are angled toward the front of the animal. These vertebrae resemble those of ''
Cedarosaurus ''Cedarosaurus'' (meaning "Cedar lizard" - named after the Cedar Mountain Formation, in which it was discovered) was a nasal-crested macronarian dinosaur genus from the Early Cretaceous Period (Valanginian). It was a sauropod which lived in what i ...
'', ''
Aeolosaurus ''Aeolosaurus'' (; "Aeolus' lizard") is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous Period of what is now South America. Like most sauropods, it would have been a quadrupedal herbivore with a long neck and tail. ''Aeolosau ...
'', and ''
Gondwanatitan ''Gondwanatitan'' (meaning "giant from Gondwana") was a titanosaurian sauropod dinosaur. ''Gondwanatitan'' was found in Brazil, at the time part of the southern supercontinent Gondwana, in the late Cretaceous Period (70 mya). Like some other ...
''. The vertebrae are located at a transitional position from anterior to posterior caudal vertebrae. ''Venenosaurus'' had unusual lateral fossae, which looked like deep depressions in the outside walls of the vertebral centra. Some fossae are divided into two chambers by a ridge inside the depression. In most sauropods the fossae would form pneumatic openings leading to the interior of the centrum, rather than just being a depression. Less well-developed, but similar fossae are known from ''Cedarosaurus''. Fossae that similarly resemble shallow depressions are known from ''Saltasaurus'', ''Alamosaurus'', ''Aeolosaurus'', ''Gondwanatitan'', and ''Malawisaurus''. These taxa differ, however, in that their fossae are even shallower, lack the division into chambers, and do not extend as far into the vertebral columns as those of ''Venenosaurus''. The vertebrae near the base of the tail are extremely useful for classifying sauropods. Derived titanosaurs had vertebrae that were convex on the front and back. Primitive sauropods had vertebrae that were either flat on both ends ( amphiplaty) or concave on both ( amphicoely). ''Venenosaurus'' may have had a condition intermediate between the two. The possession of amphiplatyan caudal centra with anteriorly facing neural spines is a unique identifier of this species. Sometimes the form of central articulations change within a single individual's vertebral column.


Footnotes


References


Works cited

* * * * * * {{Taxonbar, from=Q144253 Early Cretaceous dinosaurs of North America Macronarians Fossil taxa described in 2001 Taxa named by Kenneth Carpenter Paleontology in Utah Cretaceous Utah Aptian genus first appearances Albian genus extinctions Early Cretaceous sauropods Sauropods of North America