''Titanotylopus'' is an

extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

terrestrial Terrestrial refers to things related to land or the planet Earth. Terrestrial may also refer to: * Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on or near the ground, as opposed to ...

herbivore A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpart ...

in the

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

Camelidae Camelids are members of the biological family Camelidae, the only currently living family in the suborder Tylopoda. The seven extant members of this group are: dromedary camels, Bactrian camels, wild Bactrian camels, llamas, alpacas, vicuñas, ...

, endemic to

North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...

from the late Hemphillian stage of the

Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...

through the Irvingtonian stage of the

Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fina ...

. It was one of the last surviving North American camels, after its extinction, only ''

Camelops ''Camelops''Being occasionally called ''Western Camel'' or ''Yesterday's Camel''. is an extinct genus of camels that lived in North and Central America, ranging from Alaska to Guatemala, from the middle Pliocene to the end of the Pleistocene. It ...

'' remained. Its closest living relative is the

Bactrian camel The Bactrian camel (''Camelus bactrianus''), also known as the Mongolian camel or domestic Bactrian camel, is a large even-toed ungulate native to the steppes of Central Asia. It has two humps on its back, in contrast to the single-humped drom ...

. Its name is derived from the

Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...

words Τιτάν, τύλος and πούς — " Titan", "knob" and "foot"; thus, "giant knobby-foot."

Paleobiology

''Titanotylopus'' is distinguished from other early large camelids by its large upper

canines Canine may refer to: Zoology and anatomy * a dog-like Canid animal in the subfamily Caninae ** ''Canis'', a genus including dogs, wolves, coyotes, and jackals ** Dog, the domestic dog * Canine tooth, in mammalian oral anatomy People with the surn ...

and other distinguishing dental characteristics, and absence of

lacrimal vacuities The term Lacrimal or lachrymal, may refer to: Anatomy * Lacrimal apparatus * Lacrimal artery * Lacrimal bone * Lacrimal canaliculi (singular: canaliculus), also known as ''Lacrimal ducts'' * Lacrimal fossa (disambiguation) * Lacrimal fluid, see ...

in the skull. Unlike the smaller, contemporaneous ''

'', ''Titanotylopus'' had relatively broad second

phalanges The phalanges (singular: ''phalanx'' ) are digital bones in the hands and feet of most vertebrates. In primates, the thumbs and big toes have two phalanges while the other digits have three phalanges. The phalanges are classed as long bones. ...

, suggesting that it had true padded "cameltoes," like modern camels.Björn Kurtén and Elaine Anderson ''Pleistoceone Mammals of North America'' (New York : Columbia University Press, 1980), pp. 301–302 The species ''Titanotylopus spatulus'' was characterized by broad, spatula-like

incisor Incisors (from Latin ''incidere'', "to cut") are the front teeth present in most mammals. They are located in the premaxilla above and on the mandible below. Humans have a total of eight (two on each side, top and bottom). Opossums have 18, whe ...

s. It has been found at Grand View, Red Light fauna of the

Love Formation The Love Formation is a geologic formation in Texas. It preserves fossils. See also * List of fossiliferous stratigraphic units in Texas * Paleontology in Texas Paleontology in Texas refers to paleontological research occurring within or cond ...

Hudspeth County, Texas Hudspeth County is a county located in the U.S. state of Texas. As of the 2020 census, the population was 3,202. Its county seat is Sierra Blanca, and the largest community is Fort Hancock. The county is named for Claude Benton Hudspeth, a ...

Donnelly Ranch Donnelly is a surname of Irish origin. It is the anglicized form of the Gaelic ''Ó Donnghaile'' meaning ‘descendant of Donnghal’ a given name composed of the elements "donn" (dark or brown), plus "gal" (valour). O'Donnelly is derived from ...

White Rock, Kansas White Rock is a ghost town in Republic County, Kansas, United States. It is located 6 miles north of Courtland. History Several attempts were made to settle the area, but it wasn't until 1866 when Thomas Lovewell settled the town permanently. W ...

Mullen II Mullen is a surname of Irish and Scottish origin. Notable people with the surname include: * Alex Mullen (born 1992), Australian professional footballer *Andrew Mullen (born 1996), British Paralympian swimmer * Ann A. Mullen (1935–1994), Ame ...

(

Kansas Kansas () is a state in the Midwestern United States. Its capital is Topeka, and its largest city is Wichita. Kansas is a landlocked state bordered by Nebraska to the north; Missouri to the east; Oklahoma to the south; and Colorado to the ...

), Sandahl Local Fauna (

Nebraska Nebraska () is a state in the Midwestern region of the United States. It is bordered by South Dakota to the north; Iowa to the east and Missouri to the southeast, both across the Missouri River; Kansas to the south; Colorado to the southwe ...

), Vallecito Creek,

Colorado Colorado (, other variants) is a state in the Mountain West subregion of the Western United States. It encompasses most of the Southern Rocky Mountains, as well as the northeastern portion of the Colorado Plateau and the western edge of t ...

and 111 Ranch, Arizona in North America.

Appearance

''Titanotylopus'' possessed long and massive limbs, a comparatively small braincase, and a convex slope between the eyes. It reached a shoulder height of over and a weight of . Like modern camels, it possessed a hump for fat storage; evidence for this is provided by the long neural spines on its

thoracic The thorax or chest is a part of the anatomy of humans, mammals, and other tetrapod animals located between the neck and the abdomen. In insects, crustaceans, and the extinct trilobites, the thorax is one of the three main divisions of the crea ...

vertebra The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic ...

Alternate classification

While some authors have considered ''Gigantocamelus'' and ''Titanotylopus'' to be congeneric, others have maintained them separately. Voorhies and Corner, based on previously unreported material, documented that the two are indeed worthy of separate generic status. Harrison (1985) followed Voorhies and Corner in advocating the use of ''Titanotylopus'' for only ''T. nebraskensis'', based on a lower

jaw The jaw is any opposable articulated structure at the entrance of the mouth, typically used for grasping and manipulating food. The term ''jaws'' is also broadly applied to the whole of the structures constituting the vault of the mouth and serv ...

, and ''Gigantocamelus'' for ''G. spatula'', which includes ''G. fricki''. There is a clear difference between the proximal phalanx of specimens assigned to ''Gigantocamelus'' and to ''Titanotylopus'', based on

skeleton A skeleton is the structural frame that supports the body of an animal. There are several types of skeletons, including the exoskeleton, which is the stable outer shell of an organism, the endoskeleton, which forms the support structure inside ...

s associated with

skull The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, the ...

material.

Paleobiology

Appearance

Alternate classification

See also

References

Further reading