Todus
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''Todus'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of birds in the family Todidae, the todies, found in the
Caribbean The Caribbean (, ) ( es, El Caribe; french: la Caraïbe; ht, Karayib; nl, De Caraïben) is a region of the Americas that consists of the Caribbean Sea, its islands (some surrounded by the Caribbean Sea and some bordering both the Caribbean Se ...
. It is the only extant genus within the family Todidae. The five
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
are small,
near passerine Near passerines and higher land-bird assemblage are terms of traditional, pre-cladistic taxonomy that have often been given to tree-dwelling birds or those most often believed to be related to the true passerines (order Passeriformes) owing to mor ...
birds of the forests of the
Greater Antilles The Greater Antilles ( es, Grandes Antillas or Antillas Mayores; french: Grandes Antilles; ht, Gwo Zantiy; jam, Grieta hAntiliiz) is a grouping of the larger islands in the Caribbean Sea, including Cuba, Hispaniola, Puerto Rico, Jamaica, and ...
: Puerto Rico, Jamaica, and Cuba, with adjacent islands, have one species each, and
Hispaniola Hispaniola (, also ; es, La Española; Latin and french: Hispaniola; ht, Ispayola; tnq, Ayiti or Quisqueya) is an island in the Caribbean that is part of the Greater Antilles. Hispaniola is the most populous island in the West Indies, and th ...
has two, the broad-billed tody in the lowlands (including
Gonâve Island Gonâve Island or Zile Lagonav (french: Île de la Gonâve, ; also ''La Gonâve'') is an island of Haiti located west-northwest of Port-au-Prince in the Gulf of Gonâve. It is the largest of the Hispaniolan satellite islands. The island is an a ...
) and the
narrow-billed tody The narrow-billed tody (''Todus angustirostris'') is a species of bird in the family Todidae. It is one of two Todus species endemic to Hispaniola, a Caribbean island shared by the Dominican Republic and Haiti. Taxonomy and systematics ...
in the highlands.


Taxonomy and systematics

The
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''Todus'' was introduced by the French zoologist
Mathurin Jacques Brisson Mathurin Jacques Brisson (; 30 April 1723 – 23 June 1806) was a French zoologist and natural philosopher. Brisson was born at Fontenay-le-Comte. The earlier part of his life was spent in the pursuit of natural history; his published works ...
in 1760 with the
Jamaican tody The Jamaican tody (''Todus todus'') is a species of bird in the genus ''Todus'' endemic to Jamaica. Local names for the Jamaican tody include ''rasta bird'', ''robin'' and ''robin redbreast''. Taxonomy It was thought that the genus ''Todus'' w ...
(''Todus todus'') as the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
. ''Todus'' is a Latin word for a small bird mentioned by the Roman playwright
Plautus Titus Maccius Plautus (; c. 254 – 184 BC), commonly known as Plautus, was a Roman playwright of the Old Latin period. His comedies are the earliest Latin literary works to have survived in their entirety. He wrote Palliata comoedia, the gen ...
and the grammarian
Sextus Pompeius Festus Sextus Pompeius Festus, usually known simply as Festus, was a Roman grammarian who probably flourished in the later 2nd century AD, perhaps at Narbo (Narbonne) in Gaul. Work He made a 20-volume epitome of Verrius Flaccus's voluminous and encyclo ...
. This name had earlier been used for the Jamaican tody by the Irish physician Patrick Browne in his book ''The Civil and Natural History of Jamaica'' which was published in 1756.


Extant species

Five
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
are recognized:


Former species

*
Emperor fairywren The emperor fairywren (''Malurus cyanocephalus'') is a species of bird in the Australasian wren family, Maluridae. It is found in New Guinea in its natural habitat of subtropical or tropical moist lowland forests. It is the largest species of fa ...
(as ''Todus cyanocephalus'') *
Leaden flycatcher The leaden flycatcher (''Myiagra rubecula'') is a species of passerine bird in the family Monarchidae. Around 15 cm (6 in) in length, the male is lustrous azure with white underparts, while the female possesses leaden head, mantle and ...
(as ''Todus rubecula'')


Description

Todies range in weight from 5 to 7 g and in length from 10 to 11.5 cm. They have colourful
plumage Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...
and resemble kingfishers in their general shape. They have green heads, backs and wings, red throats (absent in immature Puerto Rican, broad-billed, and narrow-billed Todies) with a white and blue-grey stripe on each side, and yellow undertail coverts; the colour of the rest of the undersides is pale and varies according to species. The irises are pale grey. They have long, flattened bills (as do many flycatching birds) with serrated edges; the upper mandible is black and the lower is red with a little black. The legs, and especially the feet, are small. Todies are highly
vocal The human voice consists of sound made by a human being using the vocal tract, including talking, singing, laughing, crying, screaming, shouting, humming or yelling. The human voice frequency is specifically a part of human sound production i ...
, except that the Jamaican tody seldom calls in the non-breeding season (August to November); they give simple, unmusical buzzing notes, beeps, and guttural rattles, puffing their throats out with every call. Their wings produce a "strange, whirring rattle", though mostly when courting or defending territory in the Puerto Rican tody.


Behaviour and ecology

Todies are generally sedentary; the longest single flight known for the broad-billed tody is 40 m. Their activity is greatest in the morning when sunny weather follows rain, and in March and September.


Breeding

Like most of the
Coraciiformes The Coraciiformes are a group of usually colourful birds including the kingfishers, the bee-eaters, the rollers, the motmots, and the todies. They generally have syndactyly, with three forward-pointing toes (and toes 3 & 4 fused at their base) ...
, todies nest in tunnels, which they dig with their beaks and feet in steep banks or rotten tree trunks. The tunnel is 30 cm long in the Cuban and narrow-billed Todies, 30 to 60 cm in the broad-billed tody, and ends in a nest chamber, generally not reused. They lay about four round white eggs in the chamber. Both parents incubate but are surprisingly inattentive to the eggs. The young are
altricial In biology, altricial species are those in which the young are underdeveloped at the time of birth, but with the aid of their parents mature after birth. Precocial species are those in which the young are relatively mature and mobile from the mome ...
and stay in the nest until they can fly. Both parents also care for the nestlings, much more attentively; they may feed each chick up to 140 times per day, the highest rate known among birds.


Food and feeding

Todies eat small prey such as insects and lizards. Insects, particularly grasshoppers, crickets, beetles, bugs, butterflies, bees, wasps, and ants, form the greater part of the diet. Spiders and millipedes may also be taken, as is a small amount of fruit (2% of the diet). Todies typically sit on a low, small branch, singly or in pairs, keeping still or stepping or hopping sideways. When they see prey moving on the lower surface of a leaf, they fly a short distance (averaging 2.2 m in the broad-billed tody and 1.0 m in the
Puerto Rican tody The Puerto Rican tody (''Todus mexicanus'') is a bird endemic to Puerto Rico. It is locally known in Spanish as "San Pedrito" ("little Saint Peter") and "medio peso" ("half-dollar bird"). Taxonomy Todies are the closest relative to the motmots ...
), diagonally upward to
glean Gleaning is the act of collecting leftover crops from farmers' fields after they have been commercially harvested or on fields where it is not economically profitable to harvest. It is a practice described in the Hebrew Bible that became a legall ...
it. They may also take prey from the ground, occasionally chasing it with a few hops.


References


External links


Tody videos
on the Internet Bird Collection {{Taxonbar, from=Q726375 Higher-level bird taxa restricted to the West Indies * Bird genera