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The tiny hawk (''Microspizias superciliosus'') is a small diurnal
bird of prey Birds of prey or predatory birds, also known as raptors, are hypercarnivorous bird species that actively hunt and feed on other vertebrates (mainly mammals, reptiles and other smaller birds). In addition to speed and strength, these predat ...
found in or near forests, primarily humid, throughout much of the
Neotropics The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the Americas and the entire South American temperate zone. Definition In bioge ...
. It is primarily a bird-eater, and is known to prey on hummingbirds.


Taxonomy

The tiny hawk was formally described by the Swedish naturalist
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his Nobility#Ennoblement, ennoblement in 1761 as Carl von Linné#Blunt, Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalise ...
in 1766 in the twelfth edition of his '' Systema Naturae'' under the binomial name ''Falco superciliosus''. Linnaeus specified the
locality Locality may refer to: * Locality (association), an association of community regeneration organizations in England * Locality (linguistics) * Locality (settlement) * Suburbs and localities (Australia), in which a locality is a geographic subdivis ...
as Suriname. The specific epithet ''superciliosus'' is Latin meaning "eye-browed". There are two subspecies separated by the northern
Andes The Andes, Andes Mountains or Andean Mountains (; ) are the longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range is long, wide (widest between 18°S – 20°S ...
. The two differ most significantly in relative tail length; differences in size and color are small, and can be hard to distinguish. *''A. s. superciliosus'' is found east of the Andes in South America. Slightly the larger of the two subspecies, it is paler and has diffuse, grayish barring on the underparts. *''A. s. fontanieri'' is found from
Nicaragua Nicaragua (; ), officially the Republic of Nicaragua (), is the largest country in Central America, bordered by Honduras to the north, the Caribbean to the east, Costa Rica to the south, and the Pacific Ocean to the west. Managua is the countr ...
down to western
Ecuador Ecuador ( ; ; Quechua: ''Ikwayur''; Shuar: ''Ecuador'' or ''Ekuatur''), officially the Republic of Ecuador ( es, República del Ecuador, which literally translates as "Republic of the Equator"; Quechua: ''Ikwadur Ripuwlika''; Shuar: ' ...
. It is somewhat smaller and darker than the nominate ''A. s. superciliosus'', with a shorter tail and sharper, blacker barring below. The tiny hawk and its
sister species In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...
, the upland-dwelling semicollared hawk (''"A." collaris''), form a superspecies. Tiny hawk is the only
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of ''
Accipiter ''Accipiter'' is a genus of birds of prey in the family Accipitridae. With 51 recognized species it is the most diverse genus in its family. Most species are called goshawks or sparrowhawks, although almost all New World species (excepting th ...
'' yet studied which has a large procoracoid
foramen In anatomy and osteology, a foramen (;Entry "foramen"
in
. It also differs from the typical sparrowhawks in other respects of its
anatomy Anatomy () is the branch of biology concerned with the study of the structure of organisms and their parts. Anatomy is a branch of natural science that deals with the structural organization of living things. It is an old science, having it ...
and in its DNA sequence. Consequently, its old name ''Hieraspiza superciliosa'' may be more appropriate. In 2021, a phylogenetic study found it and the semicollared hawk to form a distinct group from the rest of ''Accipiter'', that is sister to '' Kaupifalco''. For this reason, both were reclassified into the new genus '' Microspizias''.


Description

The tiny hawk is aptly named; males measure a mere , about the size of a
starling Starlings are small to medium-sized passerine birds in the family Sturnidae. The Sturnidae are named for the genus '' Sturnus'', which in turn comes from the Latin word for starling, ''sturnus''. Many Asian species, particularly the larger ones, ...
, though females are slightly larger at 26.5 cm (10.5 in). It is one of the smallest true raptors in the world and is one of the smallest ''
Accipiter ''Accipiter'' is a genus of birds of prey in the family Accipitridae. With 51 recognized species it is the most diverse genus in its family. Most species are called goshawks or sparrowhawks, although almost all New World species (excepting th ...
'' species, though the
little sparrowhawk The little sparrowhawk (''Accipiter minullus'') is a species of Afrotropical bird of prey in the family Accipitridae. It is the smallest member of the genus ''Accipiter'' and forms a superspecies with the red-thighed sparrowhawk (''Accipiter er ...
of Africa is of similar or even smaller size. The birds range in weight from As with most raptors, there is considerable
sexual dimorphism Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most an ...
in size, with females measuring up to 25% longer and as much as 60% heavier than males. The adult male tiny hawk is dark slaty gray above, with a mottled gray face, a paler gray supercilium, and a blackish crown and nape. The white underparts are covered with fine gray barring, except on the unmarked throat. The longish tail is dark with three paler gray bands. Adult females are similar, but browner above with buffier underparts and browner barring below. The
cere The beak, bill, or rostrum is an external anatomical structure found mostly in birds, but also in turtles, non-avian dinosaurs and a few mammals. A beak is used for eating, preening, manipulating objects, killing prey, fighting, probing for food ...
and legs of all ages are yellow, and the bill, which is relatively heavy, is gray. Adults have red or red-brown irises, while those of immatures often are yellowish. Unlike most
accipiter ''Accipiter'' is a genus of birds of prey in the family Accipitridae. With 51 recognized species it is the most diverse genus in its family. Most species are called goshawks or sparrowhawks, although almost all New World species (excepting th ...
s, the tiny hawk's tail is quite short; it is squared or notched at the tip. Its wings are medium-length, with pointed tips, and its legs and toes are long. Immature birds come in two color morphs: the brown and the
rufous Rufous () is a color that may be described as reddish-brown or brownish-red, as of rust or oxidised iron. The first recorded use of ''rufous'' as a color name in English was in 1782. However, the color is also recorded earlier in 1527 as a dia ...
. Brown morph birds are dark sooty-brown above, finely barred with black, and white below, heavily barred with cinnamon. Rufous morph immatures are more chestnut above, and barred with rufous (rather than cinnamon) on the underparts. They also have duskier heads and brighter rufous tails than do brown morph birds. The
call Call or Calls may refer to: Arts, entertainment, and media Games * Call, a type of betting in poker * Call, in the game of contract bridge, a bid, pass, double, or redouble in the bidding stage Music and dance * Call (band), from Lahore, Paki ...
of the tiny hawk is a shrill, high-pitched, somewhat quavery series of 20–30 notes of uneven pitch. After an initial few accelerating notes, the call settles into a steady rhythm, variously transcribed as ''caucau-ca-ca-ca'', ''keer-keer-keer'' or ''kree-ree-ree-ree''.


Distribution and habitat

Tiny hawks are patchily distributed from eastern
Nicaragua Nicaragua (; ), officially the Republic of Nicaragua (), is the largest country in Central America, bordered by Honduras to the north, the Caribbean to the east, Costa Rica to the south, and the Pacific Ocean to the west. Managua is the countr ...
down to western
Ecuador Ecuador ( ; ; Quechua: ''Ikwayur''; Shuar: ''Ecuador'' or ''Ekuatur''), officially the Republic of Ecuador ( es, República del Ecuador, which literally translates as "Republic of the Equator"; Quechua: ''Ikwadur Ripuwlika''; Shuar: ' ...
, northern Bolivia, northern
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
,
Paraguay Paraguay (; ), officially the Republic of Paraguay ( es, República del Paraguay, links=no; gn, Tavakuairetã Paraguái, links=si), is a landlocked country in South America. It is bordered by Argentina to the south and southwest, Brazil to th ...
and southern
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
. They frequent humid and wet
lowland Upland and lowland are conditional descriptions of a plain based on elevation above sea level. In studies of the ecology of freshwater rivers, habitats are classified as upland or lowland. Definitions Upland and lowland are portions of p ...
forest A forest is an area of land dominated by trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological function. The United Nations' ...
s (including second growth) and forested
foothills Foothills or piedmont are geographically defined as gradual increases in elevation at the base of a mountain range, higher hill range or an upland area. They are a transition zone between plains and low relief hills and the adjacent topogr ...
from sea level to about 1800 meters (5900 ft), though most records come from elevations below 1000 m (3300 ft). The species is believed to be fairly sedentary.


Behavior

On clear mornings (and occasionally in late afternoons), the tiny hawk will sometimes sun itself on a high open branch. Occasionally, pairs will sun together. Otherwise, this is a secretive species and easily overlooked. It generally hunts from a perch, located anywhere from the undergrowth to the
canopy Canopy may refer to: Plants * Canopy (biology), aboveground portion of plant community or crop (including forests) * Canopy (grape), aboveground portion of grapes Religion and ceremonies * Baldachin or canopy of state, typically placed over an ...
, though most often in the middle story. When hunting in low growth, it often flashes from one perch to another in rapid succession. It is a fast flier; instead of the typical flap-flap-glide cadence of the accipiters, it makes a few quick flaps and then briefly closes its wings before flapping again. Tiny hawks occasionally soar above the forest canopy.


Food and feeding

Like all
accipiter ''Accipiter'' is a genus of birds of prey in the family Accipitridae. With 51 recognized species it is the most diverse genus in its family. Most species are called goshawks or sparrowhawks, although almost all New World species (excepting th ...
s, the tiny hawk feeds primarily on birds. It hunts hummingbirds and small
passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped'), which includes more than half of all bird species. Sometimes known as perching birds, passerines are distinguished from other orders of birds by th ...
s, typically darting out from a place of concealment to snatch them as they pass by, but also ambushing them when the smaller birds are perched. There is some evidence that it learns the regular perches of some hummingbirds and hunts for them there. However, the species cannot be considered a specialist predator of hummingbirds. Some individuals also hunt
rodent Rodents (from Latin , 'to gnaw') are mammals of the order Rodentia (), which are characterized by a single pair of continuously growing incisors in each of the upper and lower jaws. About 40% of all mammal species are rodents. They are n ...
s and bats.


Breeding

Little is known about the tiny hawk's
breeding Breeding is sexual reproduction that produces offspring, usually animals or plants. It can only occur between a male and a female animal or plant. Breeding may refer to: * Animal husbandry, through selected specimens such as dogs, horses, and ra ...
biology. Their breeding season may vary depending on their location; from
Panama Panama ( , ; es, link=no, Panamá ), officially the Republic of Panama ( es, República de Panamá), is a transcontinental country spanning the southern part of North America and the northern part of South America. It is bordered by Co ...
to Colombia, it apparently runs between February and June, while in the southern part of the range, it may range from October to January. They are known to build stick
nest A nest is a structure built for certain animals to hold eggs or young. Although nests are most closely associated with birds, members of all classes of vertebrates and some invertebrates construct nests. They may be composed of organic materi ...
s, at least sometimes in the canopy of tall
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
s. In
Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in th ...
, there is a record of a pair nesting in an old black-collared hawk nest. Females lay one to three bluish-white
eggs Humans and human ancestors have scavenged and eaten animal eggs for millions of years. Humans in Southeast Asia had domesticated chickens and harvested their eggs for food by 1,500 BCE. The most widely consumed eggs are those of fowl, especial ...
, faintly streaked and spotted with brown. Incubation and
fledging Fledging is the stage in a flying animal's life between hatching or birth and becoming capable of flight. This term is most frequently applied to birds, but is also used for bats. For altricial birds, those that spend more time in vulnerable c ...
periods are not known.


References


Sources

* * * For further information on the procoracoid foramen in the birds of prey, see also Olson (1987) "Variation in the procoracoid foramen in the Accipitridae" ''Riv. Ital. Orn.'' 57(3-4):161-16

!-- argh, JavaScript cite journal toolbar not working, will save and re-edit: not in a mood to hand-type the whole thing ~~~~ --> "The typical and presumably primitive condition in the Accipitridae is to have the procoracoid process with a distinct foramen for the supracoracoideus nerve piercing the bone."


External links


Tiny Hawk videos
on the Internet Bird Collection website

ttp://vireo.acnatsci.org/species_image.php?species=Accipiter+superciliosus Photo-High Res--(Close-up)on Visual Resources for Ornithology (VIREO) website {{Taxonbar, from=Q127828 tiny hawk Birds of prey Birds of the Amazon Basin Birds of Brazil Birds of Colombia Birds of Venezuela Birds of Ecuador Birds of the Guianas Birds of Costa Rica Birds of Panama tiny hawk tiny hawk