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''Thuringothyris'' is an
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
Early Permian 01 or '01 may refer to: * The year 2001, or any year ending with 01 * The month of January * 1 (number) Music * '01 (Richard Müller album), 01'' (Richard Müller album), 2001 * 01 (Son of Dave album), ''01'' (Son of Dave album), 2000 * 01 (Urban ...
eureptiles Eureptilia ("true reptiles") is one of the two major subgroups of the clade Sauropsida, the other one being Parareptilia. Eureptilia includes Diapsida (the clade containing all modern reptiles and birds), as well as a number of primitive Perm ...
known from the
Thuringian Forest The Thuringian Forest (''Thüringer Wald'' in German), is a mountain range in the southern parts of the German state of Thuringia, running northwest to southeast. Skirting from its southerly source in foothills to a gorge on its north-west side i ...
in central
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...
.


Description

''Thuringothyris'' is known from the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
MNG Multiple-image Network Graphics (MNG) is a graphics file format, published in 2001, for animated images. Its specification is publicly documented and there are free software Free software or libre software is computer software distribute ...
 7729, articulated well-preserved
skull The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, the ...
and partial
postcrania Postcrania (postcranium, adjective: postcranial) in zoology and vertebrate paleontology is all or part of the skeleton apart from the skull. Frequently, fossil remains, e.g. of dinosaurs or other extinct tetrapods, consist of partial or isolated sk ...
l
skeleton A skeleton is the structural frame that supports the body of an animal. There are several types of skeletons, including the exoskeleton, which is the stable outer shell of an organism, the endoskeleton, which forms the support structure inside ...
, and from the referred specimens MNG 10652, poorly preserved skull and partial vertebral column, MNG 10647, disarticulated
cranial Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position pro ...
and postcranial remains of at least four individuals, MNG 10183, slightly crushed skull and partial postcranial skeleton and MNG 11191, poorly preserved skull and partial limbs. All specimens were collected from the Tambach-Sandstein Member, the uppermost part of the
Tambach Formation The Tambach Formation is an Early Permian-age geologic formation in central Germany. It consists of red to brown-colored sedimentary rocks (red beds) such as conglomerate, sandstone, and mudstone, and is the oldest portion of the Upper Rotliegend ...
, dating to the
Artinskian In the geologic timescale, the Artinskian is an age or stage of the Permian. It is a subdivision of the Cisuralian Epoch or Series. The Artinskian likely lasted between and million years ago (Ma) according to the most recent revision of the ...
stage of the Late
Cisuralian The Cisuralian is the first Series (stratigraphy), series/Epoch (geology), epoch of the Permian. The Cisuralian was preceded by the Pennsylvanian (geology), Pennsylvanian and followed by the Guadalupian. The Cisuralian Epoch is named after the w ...
Series Series may refer to: People with the name * Caroline Series (born 1951), English mathematician, daughter of George Series * George Series (1920–1995), English physicist Arts, entertainment, and media Music * Series, the ordered sets used in ...
(or alternatively upper
Rotliegend The Rotliegend, Rotliegend Group or Rotliegendes (german: the underlying red) is a lithostratigraphy, lithostratigraphic unit (a sequence of rock strata) of latest Carboniferous to Guadalupian (middle Permian) age that is found in the subsurface of ...
), about 284–279.5
million years ago The abbreviation Myr, "million years", is a unit of a quantity of (i.e. ) years, or 31.556926 teraseconds. Usage Myr (million years) is in common use in fields such as Earth science and cosmology. Myr is also used with Mya (million years ago). ...
. They were found in the Bromacker Quarry, the middle part of the Thuringian Forest, near the small town of
Tambach-Dietharz Tambach-Dietharz is a town in the district of Gotha, in Thuringia, Germany. It is situated in the Thuringian Forest, 19 km south of Gotha. Mayor Since 2012, Marco Schütz (independent) is the mayor. His predecessor was the former lieutenan ...
. ''Thuringothyris'' was originally thought to be protorothyridid. A redescription of all known ''Thuringothyris'' specimens by Johannes Müller, David S. Berman, Amy C. Henrici, Thomas Martens and Stuart S. Sumida in 2006 suggested that it is a sister taxon of Captorhinidae. A noval phylogenic study of primitive reptile relationships by Müller & Reisz in 2006 recovered ''Thuringothyris'' as a sister taxon of the Captorhinidae.Muller, J. and Reisz, R.R. (2006). "The phylogeny of early eureptiles: Comparing parsimony and Bayesian approaches in the investigation of a basal fossil clade." ''Systematic Biology'', 55(3):503-511. The same results were obtained in later phylogenic analyses.


Etymology

''Thuringothyris'' was first named by Jürgen A. Boy and Thomas Martens in
1991 File:1991 Events Collage.png, From left, clockwise: Boris Yeltsin, elected as Russia's first president, waves the new flag of Russia after the 1991 Soviet coup d'état attempt, orchestrated by Soviet hardliners; Mount Pinatubo erupts in the Phil ...
and the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
is ''Thuringothyris mahlendorffae''. The generic name is named after its finding place
Thuringia Thuringia (; german: Thüringen ), officially the Free State of Thuringia ( ), is a state of central Germany, covering , the sixth smallest of the sixteen German states. It has a population of about 2.1 million. Erfurt is the capital and larg ...
. The specific name honors Ursula R. Mahlendorf.


References

Prehistoric life of Europe Permian reptiles of Europe Fossil taxa described in 1991 Captorhinids Prehistoric reptile genera {{permian-reptile-stub