Threefin Blenny
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Threefin or triplefin blennies are blenniiforms, small percomorph marine
fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of ...
of the family Tripterygiidae. Found in tropical and temperate waters of the
Atlantic The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the " Old World" of Africa, Europe an ...
,
Pacific The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the contine ...
and
Indian Ocean The Indian Ocean is the third-largest of the world's five oceanic divisions, covering or ~19.8% of the water on Earth's surface. It is bounded by Asia to the north, Africa to the west and Australia to the east. To the south it is bounded by t ...
s, the family contains about 150 species in 30 genera. The family name derives from the
Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...
''tripteros'' meaning "with three wings". With an elongated, typical blenny form, threefin blennies differ from their relatives by having a
dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through c ...
separated into three parts (hence the name); the first two are spinous. The small, slender pelvic fins are located underneath the throat and possess a single spine; the large anal fin may have one or two spines. The pectoral fins are greatly enlarged, and the tail fin is rounded. The
New Zealand topknot The New Zealand topknot, ''Notoclinus fenestratus'', is a triplefin of the genus '' Notoclinus'', found around the North Island of New Zealand in reef A reef is a ridge or shoal of rock, coral or similar relatively stable material, lyi ...
, ''Notoclinus fenestratus'', is the largest species at 20 cm in total length; most other species do not exceed 6 cm. Many threefin blennies are brightly coloured, often for reasons of
camouflage Camouflage is the use of any combination of materials, coloration, or illumination for concealment, either by making animals or objects hard to see, or by disguising them as something else. Examples include the leopard's spotted coat, the b ...
; these species are popular in the aquarium hobby. As demersal fish, threefin blennies spend most of their time on or near the bottom on
coral Corals are marine invertebrates within the class Anthozoa of the phylum Cnidaria. They typically form compact colonies of many identical individual polyps. Coral species include the important reef builders that inhabit tropical oceans and ...
and rocks. The fish are typically found in shallow, clear waters with sun exposure, such as
lagoon A lagoon is a shallow body of water separated from a larger body of water by a narrow landform, such as reefs, barrier islands, barrier peninsulas, or isthmuses. Lagoons are commonly divided into ''coastal lagoons'' (or ''barrier lagoons'') ...
s and seaward
reef A reef is a ridge or shoal of rock, coral or similar relatively stable material, lying beneath the surface of a natural body of water. Many reefs result from natural, abiotic processes— deposition of sand, wave erosion planing down rock o ...
s; nervous fish, they retreat to rock crevices at any perceived threat. Threefin blennies are diurnal and territorial; many species exhibit
sexual dichromatism Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most ani ...
, with the females drab compared to the males. The second dorsal fin is also extended in the males of some species. Small
invertebrate Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chordate ...
s comprise the bulk of the threefin blenny diet.


Genera

FishBase lists about 150 species in 30 genera: *Subfamily Notoclininae Fricke, 2009 **'' Brachynectes'' Scott, 1957 **''
Notoclinus ''Notoclinus'' is a genus of triplefins in the family Tripterygiidae. Species * Brown topknot, '' Notoclinus compressus'' ( Hutton, 1872) * New Zealand topknot The New Zealand topknot, ''Notoclinus fenestratus'', is a triplefin of the genus ...
''
Gill A gill () is a respiratory organ that many aquatic organisms use to extract dissolved oxygen from water and to excrete carbon dioxide. The gills of some species, such as hermit crabs, have adapted to allow respiration on land provided they are ...
, 1893
* Subfamily Tripterygiinae Whitley, 1931 * '' Acanthanectes'' Holleman &
Buxton Buxton is a spa town in the Borough of High Peak, Derbyshire, England. It is England's highest market town, sited at some above sea level.Apopterygion'' Kuiter, 1986 * '' Axoclinus'' Fowler, 1944 * '' Bellapiscis'' Hardy, 1987 * '' Blennodon'' Hardy, 1987 * '' Ceratobregma'' Holleman, 1987 Yellow-and-black triplefin
''Forsterygion flavonigrum'' * '' Cremnochorites'' Holleman, 1982 * '' Crocodilichthys''
Allen Allen, Allen's or Allens may refer to: Buildings * Allen Arena, an indoor arena at Lipscomb University in Nashville, Tennessee * Allen Center, a skyscraper complex in downtown Houston, Texas * Allen Fieldhouse, an indoor sports arena on the Unive ...
&
Robertson Robertson may refer to: People * Robertson (surname) (includes a list of people with this name) * Robertson (given name) * Clan Robertson, a Scottish clan * Robertson, stage name of Belgian magician Étienne-Gaspard Robert (1763–1837) Places ...
, 1991
* '' Cryptichthys'' Hardy, 1987 * ''
Enneanectes ''Enneanectes'' is a genus of triplefin fish in the family Tripterygiidae. Species There are currently 15 recognized species in this genus: * '' Enneanectes altivelis'' Rosenblatt, 1960 (Lofty triplefin) * '' Enneanectes atrorus'' Rosenblatt, 1 ...
'' D.S. Jordan & Evermann, 1895 * '' Enneapterygius'' Rüppell, 1835 * ''
Forsterygion ''Forsterygion'' is a genus of triplefins in the family Tripterygiidae native to coastal New Zealand, but also introduced to Tasmania, Australia. Species The following species are classified in this genus: * Spotted robust triplefin, '' Forst ...
'' Whitley &
Phillipps Phillipps is both a given name and an English surname. Notable people with the name include: "Phillipps" has also been a shortened version of Philippson, a German surname especially prevalent amongst German Jews and Dutch Jews. People with th ...
, 1939
* '' Gilloblennius'' Whitley & Phillipps, 1939 * '' Helcogramma''
McCulloch McCulloch is a Scottish surname. It's a variation of the Northern Irish surname McCullough. It's commonly found in Galloway. Notable people with the surname include: *Alan McCulloch (politician), New Zealand politician *Alan McLeod McCulloch ( ...
& Waite, 1918
* '' Helcogrammoides''
Rosenblatt Rosenblatt is a surname of German and Jewish origin, meaning "rose leaf". People with this surname include: * Albert Rosenblatt (born 1936), New York Court of Appeals judge * Dana Rosenblatt, known as "Dangerous" (born 1972), American boxer * Elie ...
, 1990
* '' Karalepis'' Hardy, 1984 * '' Lepidoblennius'' Steindachner, 1867 * '' Lepidonectes'' Bussing, 1991 * '' Matanui''
Jawad Jawad is a Tehsil and a Nagar Panchayat in Neemuch district in the Indian state of Madhya Pradesh. There are six towns and 105 villages in Jawad. Demographics As of the 2011 Census of India The 2011 Census of India or the 15th India ...
& Clements, 2004
* '' Norfolkia'' Fowler, 1953 * '' Notoclinops'' Whitley, 1930 Blue-eyed triplefin
''Notoclinops segmentatus'' * '' Ruanoho'' Hardy, 1986 * '' Springerichthys'' Shen, 1994 * '' Trianectes'' McCulloch & Waite, 1918 * '' Trinorfolkia'' Fricke, 1994 * '' Tripterygion'' Risso, 1827 * ''
Ucla The University of California, Los Angeles (UCLA) is a public land-grant research university in Los Angeles, California. UCLA's academic roots were established in 1881 as a teachers college then known as the southern branch of the California ...
'' Holleman, 1993


Timeline

ImageSize = width:1000px height:auto barincrement:15px PlotArea = left:10px bottom:50px top:10px right:10px Period = from:-65.5 till:10 TimeAxis = orientation:horizontal ScaleMajor = unit:year increment:5 start:-65.5 ScaleMinor = unit:year increment:1 start:-65.5 TimeAxis = orientation:hor AlignBars = justify Colors = #legends id:CAR value:claret id:ANK value:rgb(0.4,0.3,0.196) id:HER value:teal id:HAD value:green id:OMN value:blue id:black value:black id:white value:white id:cenozoic value:rgb(0.54,0.54,0.258) id:paleogene value:rgb(0.99,0.6,0.32) id:paleocene value:rgb(0.99,0.65,0.37) id:eocene value:rgb(0.99,0.71,0.42) id:oligocene value:rgb(0.99,0.75,0.48) id:neogene value:rgb(0.999999,0.9,0.1) id:miocene value:rgb(0.999999,0.999999,0) id:pliocene value:rgb(0.97,0.98,0.68) id:quaternary value:rgb(0.98,0.98,0.5) id:pleistocene value:rgb(0.999999,0.95,0.68) id:holocene value:rgb(0.999,0.95,0.88) BarData= bar:eratop bar:space bar:periodtop bar:space bar:NAM1 bar:space bar:period bar:space bar:era PlotData= align:center textcolor:black fontsize:M mark:(line,black) width:25 shift:(7,-4) bar:periodtop from: -65.5 till: -55.8 color:paleocene text:
Paleocene The Paleocene, ( ) or Palaeocene, is a geological epoch that lasted from about 66 to 56 million years ago (mya). It is the first epoch of the Paleogene Period in the modern Cenozoic Era. The name is a combination of the Ancient Greek ''pal ...
from: -55.8 till: -33.9 color:eocene text:
Eocene The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', " ...
from: -33.9 till: -23.03 color:oligocene text: Oligocene from: -23.03 till: -5.332 color:miocene text:
Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
from: -5.332 till: -2.588 color:pliocene text: Plio. from: -2.588 till: -0.0117 color:pleistocene text:
Pleist. The Pleistocene ( , often referred to as the ''Ice age'') is the geological epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was finally confirmed in ...
from: -0.0117 till: 0 color:holocene text: H. bar:eratop from: -65.5 till: -23.03 color:paleogene text: Paleogene from: -23.03 till: -2.588 color:neogene text: Neogene from: -2.588 till: 0 color:quaternary text: Q. PlotData= align:left fontsize:M mark:(line,white) width:5 anchor:till align:left color:miocene bar:NAM1 from: -11.608 till: 0 text: Tripterygion PlotData= align:center textcolor:black fontsize:M mark:(line,black) width:25 bar:period from: -65.5 till: -55.8 color:paleocene text:
Paleocene The Paleocene, ( ) or Palaeocene, is a geological epoch that lasted from about 66 to 56 million years ago (mya). It is the first epoch of the Paleogene Period in the modern Cenozoic Era. The name is a combination of the Ancient Greek ''pal ...
from: -55.8 till: -33.9 color:eocene text:
Eocene The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', " ...
from: -33.9 till: -23.03 color:oligocene text: Oligocene from: -23.03 till: -5.332 color:miocene text:
Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
from: -5.332 till: -2.588 color:pliocene text: Plio. from: -2.588 till: -0.0117 color:pleistocene text:
Pleist. The Pleistocene ( , often referred to as the ''Ice age'') is the geological epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was finally confirmed in ...
from: -0.0117 till: 0 color:holocene text: H. bar:era from: -65.5 till: -23.03 color:paleogene text: Paleogene from: -23.03 till: -2.588 color:neogene text: Neogene from: -2.588 till: 0 color:quaternary text: Q.


References

* {{Taxonbar, from=Q1257814 Blenniiformes