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''Themeda quadrivalvis'' is a species of
grass Poaceae () or Gramineae () is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos and the grasses of natural grassland and species cultivated in lawns an ...
known by the common names grader grass, habana grass, and kangaroo grass, not to be confused with ''
Themeda triandra } ''Themeda triandra'' is a species of perennial tussock-forming grass widespread in Africa, Australia, Asia and the Pacific. In Australia it is commonly known as kangaroo grass and in East Africa and South Africa it is known as red grass and re ...
'', which is also known as kangaroo grass. It is native to
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
,''Themeda quadrivalvis''.
Weeds Australia Weed Identification. Retrieved August 9, 2012.
Nepal Nepal (; ne, नेपाल ), formerly the Federal Democratic Republic of Nepal ( ne, सङ्घीय लोकतान्त्रिक गणतन्त्र नेपाल ), is a landlocked country in South Asia. It is mai ...
, and
Malaysia Malaysia ( ; ) is a country in Southeast Asia. The federation, federal constitutional monarchy consists of States and federal territories of Malaysia, thirteen states and three federal territories, separated by the South China Sea into two r ...
.''Themeda quadrivalvis''.
Grass Manual Treatment. Retrieved August 9, 2012.
It can also be found in many other places as an
introduced species An introduced species, alien species, exotic species, adventive species, immigrant species, foreign species, non-indigenous species, or non-native species is a species living outside its native distributional range, but which has arrived there ...
and often a
noxious weed A noxious weed, harmful weed or injurious weed is a weed that has been designated by an agricultural or other governing authority as a plant that is injurious to agricultural or horticultural crops, natural habitats or ecosystems, or humans or liv ...
. It occurs in the
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
,
New Caledonia ) , anthem = "" , image_map = New Caledonia on the globe (small islands magnified) (Polynesia centered).svg , map_alt = Location of New Caledonia , map_caption = Location of New Caledonia , mapsize = 290px , subdivision_type = Sovereign st ...
,
Fiji Fiji ( , ,; fj, Viti, ; Fiji Hindi: फ़िजी, ''Fijī''), officially the Republic of Fiji, is an island country in Melanesia, part of Oceania in the South Pacific Ocean. It lies about north-northeast of New Zealand. Fiji consists ...
,
Mauritius Mauritius ( ; french: Maurice, link=no ; mfe, label=Mauritian Creole, Moris ), officially the Republic of Mauritius, is an island nation in the Indian Ocean about off the southeast coast of the African continent, east of Madagascar. It incl ...
,
Thailand Thailand ( ), historically known as Siam () and officially the Kingdom of Thailand, is a country in Southeast Asia, located at the centre of the Indochinese Peninsula, spanning , with a population of almost 70 million. The country is bo ...
,
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guine ...
,
Papua New Guinea Papua New Guinea (abbreviated PNG; , ; tpi, Papua Niugini; ho, Papua Niu Gini), officially the Independent State of Papua New Guinea ( tpi, Independen Stet bilong Papua Niugini; ho, Independen Stet bilong Papua Niu Gini), is a country i ...
,
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
, the
Middle East The Middle East ( ar, الشرق الأوسط, ISO 233: ) is a geopolitical region commonly encompassing Arabian Peninsula, Arabia (including the Arabian Peninsula and Bahrain), Anatolia, Asia Minor (Asian part of Turkey except Hatay Pro ...
and tropical America. It is a troublesome exotic weed in
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
, especially in northern regions.''Themeda quadrivalvis''.
Northern Territory Government NRETAS. Retrieved 1 May 2018.
It is also an agricultural weed in crops such as
sugar cane Sugarcane or sugar cane is a species of (often hybrid) tall, perennial grass (in the genus ''Saccharum'', tribe Andropogoneae) that is used for sugar production. The plants are 2–6 m (6–20 ft) tall with stout, jointed, fibrous stalks t ...
and
lucerne Lucerne ( , ; High Alemannic German, High Alemannic: ''Lozärn'') or Luzern ()Other languages: gsw, Lozärn, label=Lucerne German; it, Lucerna ; rm, Lucerna . is a city in central Switzerland, in the Languages of Switzerland, German-speaking po ...
.''Themeda quadrivalvis''.
Pacific Island Ecosystems at Risk. Retrieved May 1, 2018.


Description

This species is an annual grass growing up to tall. The usually folded leaves are up to long. The seed heads are rounded or fan-shaped and are accompanied by leaflike
bract In botany, a bract is a modified or specialized leaf, especially one associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves. They may be smaller, larger, or of ...
s. The grass becomes orange, red, or golden brown at maturity. The clustered spikelets in the seed heads have some tubercle-based hairs and twisted awns, which can be up to 5 centimeters long. The awn is hygroscopic, twisting when moist and drilling the seed into the soil. This species is similar to ''Themeda triandra'', a native species which tends to be smaller and more brown in color. Grader grass is a prolific producer of seed; there can be up to 1000 seeds in a single seed head.Grader Grass management guide.
NQ Dry Tropics, Northern Gulf Resource Management Group and Southern Gulf Catchments. Retrieved August 9, 2012.
The seed is dispersed by
grader A grader, also commonly referred to as a road grader, motor grader, or simply a blade, is a form of heavy equipment with a long blade used to create a flat surface during grading. Although the earliest models were towed behind horses, and lat ...
s, on fur and clothing, and as a contaminant of pasture seed supplies. The seed is sometimes found as a contaminant of
bird seed Bird food or bird seed is food (often varieties of seeds, nuts, and/or dried fruits) intended for consumption by wild and domestic birds. While most bird food is fed to commercial fowl (such as chicken or turkey), bird food is also used to fee ...
, as well.


Invasiveness

This grass is a major noxious weed in parts of Australia, where it was introduced in the 1930s. It may have arrived in supplies of straw packing. The plant was first recorded in Australia in September 1935 near Habana in the
Mackay Region The Mackay Region is a local government area located in North Queensland, Queensland, Australia. Established in 2008, it was preceded by three previous local government areas with modern histories extending back as far as 1869. It has an estima ...
of
Queensland ) , nickname = Sunshine State , image_map = Queensland in Australia.svg , map_caption = Location of Queensland in Australia , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , established_ ...
. It then spread as a contaminant of seed. The weed easily invades
ecosystem An ecosystem (or ecological system) consists of all the organisms and the physical environment with which they interact. These biotic and abiotic components are linked together through nutrient cycles and energy flows. Energy enters the syste ...
s, displacing native vegetation. It grows quickly, producing flowers within 5 to 6 weeks after
germination Germination is the process by which an organism grows from a seed or spore. The term is applied to the sprouting of a seedling from a seed of an angiosperm or gymnosperm, the growth of a sporeling from a spore, such as the spores of fungi, fer ...
, and seed within 10 weeks. Flowering generally occurs in February to June in Australia. Germination can occur throughout the year, however, given adequate moisture. The grass easily colonizes disturbed areas such as roadsides. While the grass is cultivated for animal fodder in India, it is unpalatable throughout most of the year in Australia, forming wide monotypic stands that displace palatable grasses. It also forms a heavy fuel load, increasing the likelihood of fire. The species is also invasive in
New Caledonia ) , anthem = "" , image_map = New Caledonia on the globe (small islands magnified) (Polynesia centered).svg , map_alt = Location of New Caledonia , map_caption = Location of New Caledonia , mapsize = 290px , subdivision_type = Sovereign st ...
.


References

{{Taxonbar, from=Q7781242 Andropogoneae Grasses of Africa Grasses of Asia Grasses of India Grasses of Pakistan Grasses of South Africa Taxa named by Carl Linnaeus