Thagomiser
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A thagomizer () is the distinctive arrangement of four spikes on the tails of stegosaurine dinosaurs. These spikes are believed to have been a defensive measure against predators.Carpenter, K., Sanders, F., McWhinney, L., and Wood, L. 2005. Evidence for predator-prey relationships: Example for ''Allosaurus'' and ''Stegosaurus''. Pp. 325-350 in Carpenter, K. (ed.) The Carnivorous Dinosaurs. Indiana University Press, Bloomington. The arrangement of spikes originally had no distinct name. Cartoonist Gary Larson invented the name "thagomizer" in 1982 as a joke in his comic strip ''
Far Side ''The Far Side'' is a single-panel comic created by Gary Larson and syndicated by Chronicle Features and then Universal Press Syndicate, which ran from December 31, 1979, to January 1, 1995 (when Larson retired as a cartoonist). Its surrealis ...
'', and it was gradually adopted as an informal term sometimes used within scientific circles, research, and education.


Etymology

The term ''thagomizer'' was coined by Gary Larson in jest. In a 1982 ''The Far Side''
comic a Media (communication), medium used to express ideas with images, often combined with text or other visual information. It typically the form of a sequence of Panel (comics), panels of images. Textual devices such as speech balloons, Glo ...
, a group of cavemen are taught by a caveman lecturer that the spikes on a stegosaur's tail were named "after the late Thag Simmons". The term was picked up initially by Kenneth Carpenter, then a paleontologist at the Denver Museum of Nature and Science, who used the term when describing a fossil at the
Society of Vertebrate Paleontology The Society of Vertebrate Paleontology (SVP) is a professional organization that was founded in the United States in 1940 to advance the science of vertebrate paleontology around the world. Mission and Activities SVP has about 2,300 members inter ...
Annual Meeting in 1993. ''Thagomizer'' has since been adopted as an informal
anatomical Anatomy () is the branch of biology concerned with the study of the structure of organisms and their parts. Anatomy is a branch of natural science that deals with the structural organization of living things. It is an old science, having it ...
term, and is used by the
Smithsonian Institution The Smithsonian Institution ( ), or simply the Smithsonian, is a group of museums and education and research centers, the largest such complex in the world, created by the U.S. government "for the increase and diffusion of knowledge". Founded ...
, the
Dinosaur National Monument Dinosaur National Monument is an American national monument located on the southeast flank of the Uinta Mountains on the border between Colorado and Utah at the confluence of the Green and Yampa rivers. Although most of the monument area is in ...
, the book ''The Complete Dinosaur'' and the BBC documentary series ''
Planet Dinosaur ''Planet Dinosaur'', is a six-part documentary television series created by Nigel Paterson and Phil Dobree, produced by the BBC, and narrated by John Hurt. It first aired in the United Kingdom in 2011, with VFX studio Jellyfish Pictures a ...
''. The term has also appeared in some technical papers describing stegosaurs and related dinosaurs.


Paleobiology

There has been debate about whether the thagomizer was used simply for display, as posited by Gilmore in 1914, or used as a defensive weapon. Robert Bakker noted that it is likely that the length of the stegosaur tail was much more flexible than those of other
ornithischia Ornithischia () is an extinct order of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. The name ''Ornithischia'', or "bird-hipped", reflects this similarity and is derived from the Greek s ...
n dinosaurs because it lacked ossified tendons, thus lending credence to the idea of the thagomizer being a weapon. He also observed that '' Stegosaurus'' could have maneuvered its rear easily by keeping its large hindlimbs stationary and pushing off with its very powerfully muscled but short forelimbs, allowing it to swivel deftly to deal with attack. In 2010, analysis of a digitized model of '' Kentrosaurus aethiopicus'' showed that the tail could bring the thagomizer around to the sides of the dinosaur, possibly striking an attacker beside it. In 2001, a study of thagomizers by McWhinney et al. showed a high incidence of trauma-related damage. This too supports the theory that the principle function of the thagomizer was defense in combat. There is also evidence for ''Stegosaurus'' defending itself, in the form of an '' Allosaurus'' tail vertebra with a partially healed puncture wound that fits a ''Stegosaurus'' tail spike. The species of stegosaur known as ''Stegosaurus stenops'' had four dermal spikes, each about long. Discoveries of articulated stegosaur armor show that, at least in some species, these spikes protruded horizontally from the tail, not vertically as is often depicted. Initially, Marsh described ''S. armatus'' as having eight spikes in its tail, unlike ''S. stenops''. However, recent research re-examined this and concluded this species also had four.


Mathematics

In a 2017 paper, the term
thagomizer graph In graph theory, a book graph (often written B_p ) may be any of several kinds of graph formed by multiple cycles sharing an edge. Variations One kind, which may be called a quadrilateral book, consists of ''p'' quadrilaterals sharing a comm ...
(and also the associated "thagomizer matroid") was introduced for the complete tripartite graph .


See also

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Tail club In zoology, a club is a bony mass at the end of the tail of some dinosaurs and of some mammals, most notably the ankylosaurids and the glyptodonts, as well as meiolaniid turtles. It is thought that this was a form of defensive armour or weapon ...
*
Timeline of stegosaur research This timeline of stegosaur research is a chronological listing of events in the history of paleontology focused on the stegosaurs, the iconic plate-backed, spike-tailed herbivorous eurypod dinosaurs that predominated during the Jurassic period ...
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References

{{Thefarside Dinosaur anatomy The Far Side Stegosaurs 1980s neologisms Words originating in fiction