''Tendaguria'' ( ; meaning "the
Tendaguru
The Tendaguru Formation, or Tendaguru Beds are a highly fossiliferous formation and Lagerstätte located in the Lindi Region of southeastern Tanzania. The formation represents the oldest sedimentary unit of the Mandawa Basin, overlying Neoprotero ...
one") is a
genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of herbivorous
sauropod
Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their bo ...
dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
from the Late
Jurassic
The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The J ...
of
Lindi Region,
Tanzania
Tanzania (; ), officially the United Republic of Tanzania ( sw, Jamhuri ya Muungano wa Tanzania), is a country in East Africa within the African Great Lakes region. It borders Uganda to the north; Kenya to the northeast; Comoro Islands and ...
.
Discovery and naming
In 1911, German geologist
Wilhelm Bornhardt at
Nambango in
German East Africa
German East Africa (GEA; german: Deutsch-Ostafrika) was a German colony in the African Great Lakes region, which included present-day Burundi, Rwanda, the Tanzania mainland, and the Kionga Triangle, a small region later incorporated into Mozam ...
discovered two sauropod vertebrae, fifteen kilometers (nine miles) southeast of Tendaguru Hill. These were described by
Werner Janensch
Werner Ernst Martin Janensch (11 November 1878 – 20 October 1969) was a German paleontologist and geologist.
Biography
Janensch was born at Herzberg (Elster).
In addition to Friedrich von Huene, Janensch was probably Germany's most impo ...
in 1929 but not named.
The finds were formally named by
José Fernando Bonaparte,
Wolf-Dieter Heinrich and
Rupert Wild
Rupert may refer to:
People
* Rupert (name), various people known by the given name or surname "Rupert"
Places Canada
*Rupert, Quebec, a village
* Rupert Bay, a large bay located on the south-east shore of James Bay
*Rupert River, Quebec
* Ruper ...
in 2000. The
type species
In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
is ''Tendaguria tanzaniensis'' () Bonaparte, Heinrich & Wild 2000. The generic name refers to the
Tendaguru
The Tendaguru Formation, or Tendaguru Beds are a highly fossiliferous formation and Lagerstätte located in the Lindi Region of southeastern Tanzania. The formation represents the oldest sedimentary unit of the Mandawa Basin, overlying Neoprotero ...
, the area of the great German palaeontological expeditions between 1909 and 1912. The
specific name was "after Tanzania, the country where the holotype was collected".
[J.F. Bonaparte, W.-D. Heinrich, and R. Wild, 2000, "Review of ''Janenschia'' Wild, with the description of a new sauropod from the Tendaguru beds of Tanzania and a discussion on the systematic value of procoelous caudal vertebrae in the Sauropoda", ''Palaeontographica Abteilung A'' 256(1-3): 25-76] The territory of present Tanzania largely coincides with that of the former German East Africa.
The type specimen consists of two
syntype
In biological nomenclature, a syntype is any one of two or more biological types that is listed in a description of a taxon where no holotype was designated. Precise definitions of this and related terms for types have been established as part of ...
s, MB.R.2092.1 (NB4) and MB.R.2092.2 (NB5), probably uncovered in the
Upper Saurian Bed
Upper may refer to:
* Shoe upper or ''vamp'', the part of a shoe on the top of the foot
* Stimulant, drugs which induce temporary improvements in either mental or physical function or both
* ''Upper'', the original film title for the 2013 found f ...
(''Obere Dinosauriermergel''),
Tendaguru Series, dating from the
Late Jurassic
The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987.
In European lithostratigraphy, the name ...
Tithonian
In the geological timescale, the Tithonian is the latest age of the Late Jurassic Epoch and the uppermost stage of the Upper Jurassic Series. It spans the time between 152.1 ± 4 Ma and 145.0 ± 4 Ma (million years ago). It is preceded by the K ...
. The specimens are two anterior dorsal vertebrae, part of the collection of the
Berlin's Natural History Museum. They probably belong to the same individual, having been found at a short distance from each other.
Description
''Tendaguria'' was a large sauropod from the Tendaguru fossil locality. Its length is estimated at about twenty meters (sixty-six feet). The vertebrae are
opisthocoelous (convex at the front, hollow behind) and differ from other known sauropods in their very low, almost non-existent neural spines, which are not distinct bodies of bone, do not rise above the surrounding area of the neural arch and are continuous at their rear with the transverse processes, including the
epipophyses. Viewed from above, the spines have the form of a low and broad transverse ridge. ''Tendaguria'' is thus a highly derived or
apomorphic sauropod; the axial musculature probably shifted from the neural spines to the dorsal surface of the transverse processes. Other
autapomorphies
In phylogenetics, an autapomorphy is a distinctive feature, known as a derived trait, that is unique to a given taxon. That is, it is found only in one taxon, but not found in any others or outgroup taxa, not even those most closely related to t ...
are: a depression in the upper part of the outer side of each
prezygapophysis
The articular processes or zygapophyses (Greek ζυγον = "yoke" (because it links two vertebrae) + απο = "away" + φυσις = "process") of a vertebra are projections of the vertebra that serve the purpose of fitting with an adjacent vertebr ...
; deep depressions in the front side of the transverse processes combined with shallow depressions in the rear side; the possession of robust epipophyses.
Classification
Due to its unique
morphology
Morphology, from the Greek and meaning "study of shape", may refer to:
Disciplines
* Morphology (archaeology), study of the shapes or forms of artifacts
* Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies ...
, ''Tendaguria'' defied classification in the original description, where the authors placed it as
Sauropoda
Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their bo ...
''
incertae sedis
' () or ''problematica'' is a term used for a taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertainty ...
'', though also naming a separate Tendaguriidae. It shows a mix of basal and derived traits, indicating a position outside, respectively inside,
Neosauropoda
Neosauropoda is a clade within Dinosauria, coined in 1986 by Argentine paleontologist José Bonaparte and currently described as ''Saltasaurus loricatus'', ''Diplodocus longus'', and all animals directly descended from their most recent common ...
. A cervical vertebra referred to ''Tendaguria'' because of a similar low spine shows some similarities to ''
Camarasaurus''.
In 2014 it was included in a
cladogram
A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to d ...
generated in the redescription of ''
Diamantinasaurus
''Diamantinasaurus'' is a genus of titanosaurian sauropod from Australia that lived during the early Late Cretaceous, about 94 million years ago. The type species of the genus is ''D. matildae'', first described and named in 2009 in paleontology, ...
'', where it was placed as the sister to ''
Wintonotitan
''Wintonotitan'' (meaning " Winton titan") is a genus of titanosauriform dinosaur from late Albian (Early Cretaceous)-age Winton Formation of Australia. It is known from partial postcranial remains.
Description and history
Fossils that are ...
'' within
Somphospondyli
Somphospondyli is an extinct clade of titanosauriform sauropods that lived from the Late Jurassic until the end of the Late Cretaceous, comprising all titanosauriforms more closely related to Titanosauria proper than Brachiosauridae. The remains ...
.
Mannion (2019) and Mannion ''et al.'' (2019) assign ''Tendaguria'' to Turiasauria based on comparative anatomy and a phylogenetic analysis where it is recovered as the sister taxon of ''Moabosaurus'' within the Turiasauria.
[Philip D Mannion, Paul Upchurch, Daniela Schwarz, Oliver Wings; Taxonomic affinities of the putative titanosaurs from the Late Jurassic Tendaguru Formation of Tanzania: phylogenetic and biogeographic implications for eusauropod dinosaur evolution, Zoological Journal of the Linnean Society, , zly068, https://doi.org/10.1093/zoolinnean/zly068]
References
External links
Dinosauria Translation and Pronunciation Guide
{{Taxonbar, from=Q278093
Turiasauria
Late Jurassic dinosaurs of Africa
Macronarians
Fossil taxa described in 2000
Taxa named by José Bonaparte