Taxonomic Waste-basket
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Wastebasket taxon (also called a wastebin taxon, dustbin taxon or catch-all taxon) is a term used by some
taxonomists In biology, taxonomy () is the scientific study of naming, defining ( circumscribing) and classifying groups of biological organisms based on shared characteristics. Organisms are grouped into taxa (singular: taxon) and these groups are given ...
to refer to a taxon that has the sole purpose of classifying organisms that do not fit anywhere else. They are typically defined by either their designated members' often superficial similarity to each other, or their ''lack'' of one or more distinct character states or by their ''not'' belonging to one or more other taxa. Wastebasket taxa are by definition either
paraphyletic In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...
or polyphyletic, and are therefore not considered valid taxa under strict cladistic rules of taxonomy. The name of a wastebasket taxon may in some cases be retained as the designation of an evolutionary grade, however. The term was coined in a 1985 essay by Steven Jay Gould.


Examples

There are many examples of paraphyletic groups, but true "wastebasket" taxa are those that are known not to, and perhaps not intended to, represent natural groups, but are nevertheless used as convenient groups of organisms. The acritarchs are perhaps the most famous example. Wastebasket taxa are often old (and perhaps not described with the systematic rigour and precision that is possible in the light of accumulated knowledge of diversity) and populous; further characteristics are reviewed by. * The Flacourtiaceae, a now-defunct family of flowering plants – the Angiosperm Phylogeny Group has placed its tribes and genera in various other families, especially the Achariaceae and Salicaceae. * The obsolete kingdom
Protista A protist () is any eukaryotic organism (that is, an organism whose cells contain a cell nucleus) that is not an animal, plant, or fungus. While it is likely that protists share a common ancestor (the last eukaryotic common ancestor), the excl ...
is composed of all
eukaryote Eukaryotes () are organisms whose cells have a nucleus. All animals, plants, fungi, and many unicellular organisms, are Eukaryotes. They belong to the group of organisms Eukaryota or Eukarya, which is one of the three domains of life. Bacte ...
s that are not animals, plants or fungi, leaving to the protists all single-celled eukaryotes. * The Tricholomataceae is a fungal group, at one point composed of the white-, yellow-, or pink-spored genera in the
Agaricales The fungal order Agaricales, also known as gilled mushrooms (for their distinctive gills) or euagarics, contains some of the most familiar types of mushrooms. The order has 33 extant families, 413 genera, and over 13,000 described species, alo ...
not already classified as belonging to the
Amanitaceae The Amanitaceae is a family of mushroom-forming fungi. ''Amanita'' Pers. is one of the most specious and best-known fungal genera. The family, also commonly called the amanita family, is in order Agaricales, the gilled mushrooms. The family co ...
,
Lepiotaceae The Agaricaceae are a family of basidiomycete fungi and include the genus ''Agaricus'', as well as basidiomycetes previously classified in the families Tulostomataceae, Lepiotaceae, and Lycoperdaceae. Taxonomy The family Agaricaceae was publishe ...
,
Hygrophoraceae The Hygrophoraceae are a family of fungi in the order Agaricales. Originally conceived as containing white-spored, thick-gilled agarics (gilled mushrooms), including ''Hygrophorus'' and ''Hygrocybe'' species (the waxcaps or waxy caps), DNA evide ...
, Pluteaceae, or
Entolomataceae The Entolomataceae, also known as Rhodophyllaceae, are a large family of pink-spored terrestrial gilled mushrooms which includes the genera ''Entoloma'', '' Rhodocybe'', and ''Clitopilus''. The family collectively contains over 1500 species, the ...
. * Carnosauria and Thecodontia are fossil groups, banded together back when the limited fossil record did not allow for a more detailed scheme. * Condylarthra is an artificial clade into which ungulate mammals not clearly within Perissodactyla or Cetartiodactyla were traditionally shoved. Many of these groups, like Meridiungulata or '' Protungulatum'', may not represent laurasitherian mammals, while others like phenacodontids have been clearly established as early odd-toed ungulates. * The order Insectivora has traditionally been used as a dumping ground for placental insectivorous mammals (and similar forms such as colugos), usually aligned with carnivorans, ungulates and bats. While the core components ( moles, shrews, hedgehogs and their close relations) do in fact form a consistent clade, Eulipotyphla, that is part of Laurasiatheria with the aforementioned clades, other mammals historically placed in the order have been found to belong to other branches of the placental tree: tree shrews and colugos are euarchontans related to
Primates Primates are a diverse order of mammals. They are divided into the strepsirrhines, which include the lemurs, galagos, and lorisids, and the haplorhines, which include the tarsiers and the simians (monkeys and apes, the latter including huma ...
and sometimes grouped in
Sundatheria Sundatheria is a proposed clade of placental mammals. It includes the orders Scandentia and Dermoptera. An alternative phylogeny is the Primatomorpha The Primatomorpha are a proposed mirorder of mammals containing the flying lemurs (order ...
, while tenrecs, golden moles and elephant shrews are all afrotheres, probably forming the clade
Afroinsectiphilia The Afroinsectiphilia (African insectivores) is a clade that has been proposed based on the results of recent molecular phylogenetic studies. Many of the taxa within it were once regarded as part of the order Insectivora, but Insectivora is now ...
. Both of these clades have at times been accused of being wastebasket taxa themselves, grouping superficially similar animals in Euarchonta and Afrotheria, respectively, but they have been more strongly supported by genetic studies. * Vermes is an obsolete taxon of worm-like animals. It was a catch-all term used by Carl Linnaeus and Jean-Baptiste Lamarck for non-arthropod invertebrate animals.


Wastebasket taxa in science

Fossil groups that are poorly known due to fragmentary remains are sometimes grouped together on gross morphology or
stratigraphy Stratigraphy is a branch of geology concerned with the study of rock (geology), rock layers (Stratum, strata) and layering (stratification). It is primarily used in the study of sedimentary rock, sedimentary and layered volcanic rocks. Stratigrap ...
, only later to be found to be wastebasket taxa, such as the crocodile-like Triassic group Rauisuchia. One of the roles of taxonomists is to identify wastebasket taxa and reclassify the content into more natural units. Sometimes, during taxonomic revisions, a wastebasket taxon can be salvaged after doing thorough research on its members, and then imposing tighter restrictions on what continues to be included. Such techniques "saved" Carnosauria and '' Megalosaurus''. Other times, the
taxonomic Taxonomy is the practice and science of categorization or classification. A taxonomy (or taxonomical classification) is a scheme of classification, especially a hierarchical classification, in which things are organized into groups or types. ...
name contains too much unrelated "baggage" to be successfully salvaged. As such, it is usually dumped in favour of a new, more restrictive name (for example,
Rhynchocephalia Rhynchocephalia (; ) is an order of lizard-like reptiles that includes only one living species, the tuatara (''Sphenodon punctatus'') of New Zealand. Despite its current lack of diversity, during the Mesozoic rhynchocephalians were a diverse g ...
), or abandoned altogether (for example, ''
Simia In his '' Systema Naturae'' of 1758, Carl Linnaeus divided the Order Primates within ''Mammalia'' into four genera: ''Homo'', ''Simia'', ''Lemur'', and ''Vespertilio''. His ''Vespertilio'' included all bats, and has since been moved from Primates ...
'').


Related concepts

A related concept is that of form taxon, "wastebasket" groupings that are united by gross morphology. This is often result of a common mode of life, often one that is
generalist A generalist is a person with a wide array of knowledge on a variety of subjects, useful or not. It may also refer to: Occupations * a physician who provides general health care, as opposed to a medical specialist; see also: ** General pract ...
, leading to generally similar body shapes by convergent evolution. The term wastebasket taxon is sometimes employed in a derogatory fashion to refer to an evolutionary grade taxon.


See also

* Lazarus taxon * Elvis taxon * '' Incertae sedis'' * Glossary of scientific naming


References

{{Reflist, 2 Obsolete taxa Taxonomy (biology) Paraphyletic groups Polyphyletic groups